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padding:0 .2em;\"\u003EDas Aussehen der deutschsprachigen Wikipedia hat sich geändert: Das Hauptmenü ist nun links neben dem Logo erreichbar, die eigene Diskussionsseite und Beitragsliste rechts neben dem Link zum Anmelden und die Werkzeugleiste mit Verweisen zu Schwesterprojekten wie Wikimedia Commons rechts neben dem Link zur Versionsgeschichte. \u003Ca href=\"/wiki/Hilfe:Skin/Vector_2022\" title=\"Hilfe:Skin/Vector 2022\"\u003EWeitere Informationen\u003C/a\u003E\u003C/div\u003E\u003C/div\u003E\u003C/div\u003E\u003C/div\u003E\u003C/div\u003E";}}());</script></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="Website"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="Inhaltsverzeichnis" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Inhaltsverzeichnis</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">In die Seitenleiste verschieben</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">Verbergen</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">(Anfang)</div> </a> </li> <li id="toc-Eigenschaften" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Eigenschaften"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Eigenschaften</span> </div> </a> <ul id="toc-Eigenschaften-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Einzelnachweise" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Einzelnachweise"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Einzelnachweise</span> </div> </a> <ul id="toc-Einzelnachweise-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Inhaltsverzeichnis" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" title="Inhaltsverzeichnis" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Inhaltsverzeichnis umschalten" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Inhaltsverzeichnis umschalten</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">CCL18</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Zu einem Artikel in einer anderen Sprache gehen. Verfügbar in 6 Sprachen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-6" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">6 Sprachen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/CCL18" title="CCL18 – Bosnisch" lang="bs" hreflang="bs" data-title="CCL18" data-language-autonym="Bosanski" data-language-local-name="Bosnisch" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/CCL18" title="CCL18 – Englisch" lang="en" hreflang="en" data-title="CCL18" data-language-autonym="English" data-language-local-name="Englisch" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/CCL18" title="CCL18 – Serbokroatisch" lang="sh" hreflang="sh" data-title="CCL18" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbokroatisch" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/CCL18" title="CCL18 – Serbisch" lang="sr" hreflang="sr" data-title="CCL18" data-language-autonym="Српски / srpski" data-language-local-name="Serbisch" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/CCL18" title="CCL18 – Tatarisch" lang="tt" hreflang="tt" data-title="CCL18" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatarisch" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/CCL18" title="CCL18 – Ukrainisch" lang="uk" hreflang="uk" data-title="CCL18" data-language-autonym="Українська" data-language-local-name="Ukrainisch" class="interlanguage-link-target"><span>Українська</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q21113934#sitelinks-wikipedia" title="Links auf Artikel in anderen Sprachen bearbeiten" class="wbc-editpage">Links bearbeiten</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namensräume"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/CCL18" title="Seiteninhalt anzeigen [c]" accesskey="c"><span>Artikel</span></a></li><li id="ca-talk" class="new vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Diskussion:CCL18&action=edit&redlink=1" rel="discussion" class="new" title="Diskussion zum Seiteninhalt (Seite nicht vorhanden) [t]" accesskey="t"><span>Diskussion</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="Sprachvariante ändern" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Deutsch</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="Ansichten"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/CCL18"><span>Lesen</span></a></li><li id="ca-ve-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=CCL18&veaction=edit" title="Diese Seite mit dem VisualEditor bearbeiten [v]" accesskey="v"><span>Bearbeiten</span></a></li><li id="ca-edit" class="collapsible vector-tab-noicon mw-list-item"><a href="/w/index.php?title=CCL18&action=edit" title="Den Quelltext dieser Seite bearbeiten [e]" accesskey="e"><span>Quelltext bearbeiten</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=CCL18&action=history" title="Frühere Versionen dieser Seite [h]" accesskey="h"><span>Versionsgeschichte</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="Seitenwerkzeuge"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="Werkzeuge" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Werkzeuge</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">Werkzeuge</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">In die Seitenleiste verschieben</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">Verbergen</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="Weitere Optionen" > <div class="vector-menu-heading"> Aktionen </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/CCL18"><span>Lesen</span></a></li><li id="ca-more-ve-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=CCL18&veaction=edit" title="Diese Seite mit dem VisualEditor bearbeiten [v]" accesskey="v"><span>Bearbeiten</span></a></li><li id="ca-more-edit" class="collapsible vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=CCL18&action=edit" title="Den Quelltext dieser Seite bearbeiten [e]" accesskey="e"><span>Quelltext bearbeiten</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=CCL18&action=history"><span>Versionsgeschichte</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> Allgemein </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/Spezial:Linkliste/CCL18" title="Liste aller Seiten, die hierher verlinken [j]" accesskey="j"><span>Links auf diese Seite</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Spezial:%C3%84nderungen_an_verlinkten_Seiten/CCL18" rel="nofollow" title="Letzte Änderungen an Seiten, die von hier verlinkt sind [k]" accesskey="k"><span>Änderungen an verlinkten Seiten</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=CCL18&oldid=223702235" title="Dauerhafter Link zu dieser Seitenversion"><span>Permanenter Link</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=CCL18&action=info" title="Weitere Informationen über diese Seite"><span>Seiteninformationen</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=Spezial:Zitierhilfe&page=CCL18&id=223702235&wpFormIdentifier=titleform" title="Hinweise, wie diese Seite zitiert werden kann"><span>Artikel zitieren</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=Spezial:URL-K%C3%BCrzung&url=https%3A%2F%2Fde.wikipedia.org%2Fwiki%2FCCL18"><span>Kurzlink</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=Spezial:QrCode&url=https%3A%2F%2Fde.wikipedia.org%2Fwiki%2FCCL18"><span>QR-Code herunterladen</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> Drucken/exportieren </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=Spezial:DownloadAsPdf&page=CCL18&action=show-download-screen"><span>Als PDF herunterladen</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=CCL18&printable=yes" title="Druckansicht dieser Seite [p]" accesskey="p"><span>Druckversion</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> In anderen Projekten </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q21113934" title="Link zum verbundenen Objekt im Datenrepositorium [g]" accesskey="g"><span>Wikidata-Datenobjekt</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Seitenwerkzeuge"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Erscheinungsbild"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Erscheinungsbild</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">In die Seitenleiste verschieben</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">Verbergen</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">aus Wikipedia, der freien Enzyklopädie</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="de" dir="ltr"><table class="wikitable hintergrundfarbe-basis infobox float-right" id="Vorlage_Infobox_Protein" style="font-size:90%; margin-top:0; width:350px;" summary="Infobox Protein"> <tbody><tr> <th colspan="3" style="background:#90EE90; color:#202122;">CCL18 </th></tr> <tr> <td>Andere Namen </td> <td colspan="2"> <ul><li>Chemokine (C-C Motif) Ligand 18 (Pulmonary And Activation-Regulated)</li> <li>SCYA18</li> <li>AMAC1</li> <li>DCCK1</li> <li>PARC</li> <li>Small Inducible Cytokine Subfamily A (Cys-Cys), Member 18, Pulmonary And Activation-Regulated</li> <li>Alternative Macrophage Activation-Associated CC Chemokine 1</li> <li>Macrophage Inflammatory Protein 4</li> <li>Dendritic Cell Chemokine 1</li> <li>CKb7</li> <li>MIP4</li></ul> </td></tr> <tr> <td colspan="3" class="hintergrundfarbe1" style="font-size:smaller;"> <p>Vorhandene Strukturdaten: <span class="plainlinks"><a rel="nofollow" class="external text" href="https://www.rcsb.org/structure/4MHE">4MHE</a></span> </p> </td></tr> <tr> <th colspan="3" style="background:#90EE90; color:#202122;;">Eigenschaften des menschlichen Proteins </th></tr> <tr> <td><a href="/wiki/Molare_Masse" title="Molare Masse">Masse</a>/Länge <a href="/wiki/Prim%C3%A4rstruktur" title="Primärstruktur">Primärstruktur</a> </td> <td colspan="2" style="text-align:center;">89 Aminosäuren, 9849 Da </td></tr> <tr> <th colspan="3" style="background:#90EE90; color:#202122;">Bezeichner </th></tr> <tr> <td>Externe IDs </td> <td colspan="2" class="plainlinks"> <ul><li><a href="/wiki/GeneCards" title="GeneCards">GeneCards</a>: <a rel="nofollow" class="external text" href="http://www.genecards.org/cgi-bin/carddisp.pl?gene=CCL18">CCL18</a></li> <li><a href="/wiki/Online_Mendelian_Inheritance_in_Man" title="Online Mendelian Inheritance in Man">OMIM</a>: <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/omim/603757">603757</a></li> <li><a href="/wiki/UniProt" title="UniProt">UniProt</a> <a rel="nofollow" class="external text" href="https://www.uniprot.org/uniprotkb/P55774">P55774</a></li></ul> </td></tr> <tr> <th colspan="3" style="background:#90EE90; color:#202122;">Vorkommen </th></tr> <tr> <td style="background:#C3FDB8; color:#202122;">Homologie-Familie </td> <td colspan="2" style="text-align:center;"><a rel="nofollow" class="external text" href="http://hogenom.univ-lyon1.fr/query_sequence?seq=P55774">Hovergen</a> </td></tr> </tbody></table><p><span class="editoronly" style="display:none;"></span> </p><p><b>CCL18</b> (<span style="font-style:normal;font-weight:normal"><a href="/wiki/Englische_Sprache" title="Englische Sprache">englisch</a></span> <span lang="en-Latn" style="font-style:italic">CC-chemokine ligand 18</span>) ist ein <a href="/wiki/Zytokin" title="Zytokin">Zytokin</a> aus der Familie der CC-<a href="/wiki/Chemokin" title="Chemokin">Chemokine</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Eigenschaften">Eigenschaften</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=CCL18&veaction=edit&section=1" title="Abschnitt bearbeiten: Eigenschaften" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=CCL18&action=edit&section=1" title="Quellcode des Abschnitts bearbeiten: Eigenschaften"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>CCL18 ist an <a href="/wiki/Entz%C3%BCndung" title="Entzündung">Entzündungsprozessen</a> beteiligt. Es wird als <a href="/wiki/Pr%C3%A4protein" class="mw-redirect" title="Präprotein">Präprotein</a> im <a href="/wiki/Endoplasmatisches_Retikulum" title="Endoplasmatisches Retikulum">endoplasmatischen Retikulum</a> auf seine aktive Form von 69 Aminosäuren gekürzt.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Es wird von <a href="/wiki/Dendritische_Zelle" title="Dendritische Zelle">dendritischen Zellen</a>, <a href="/wiki/Monozyt" title="Monozyt">Monozyten</a> und <a href="/wiki/Makrophage" title="Makrophage">Makrophagen</a> gebildet,<sup id="cite_ref-PMID23102918_2-0" class="reference"><a href="#cite_note-PMID23102918-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-PMID18959625_3-0" class="reference"><a href="#cite_note-PMID18959625-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-PMID22117697_4-0" class="reference"><a href="#cite_note-PMID22117697-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> nicht aber von <a href="/wiki/T-Zelle" class="mw-redirect" title="T-Zelle">T-Zellen</a> oder <a href="/wiki/B-Zelle" class="mw-redirect" title="B-Zelle">B-Zellen</a>.<sup id="cite_ref-PMID23102918_2-1" class="reference"><a href="#cite_note-PMID23102918-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> CCL18 wird durch <a href="/wiki/Interleukin_10" class="mw-redirect" title="Interleukin 10">IL-10</a>, <a href="/wiki/Interleukin_4" class="mw-redirect" title="Interleukin 4">IL-4</a> und <a href="/wiki/Interleukin_13" class="mw-redirect" title="Interleukin 13">IL-13</a> induziert, also eher durch Zytokine der Aktivierung von <a href="/wiki/TH2" class="mw-redirect" title="TH2">TH2</a>-Zellen bzw. eine <a href="/wiki/Humorale_Immunantwort" title="Humorale Immunantwort">humorale Immunantwort</a> oder der <a href="/wiki/Immunsuppression" title="Immunsuppression">Immunsuppression</a>. <a href="/wiki/Interferon-gamma" class="mw-redirect" title="Interferon-gamma">Interferon-gamma</a> hemmt die Bildung von CCL18.<sup id="cite_ref-PMID12646652_11_5-0" class="reference"><a href="#cite_note-PMID12646652_11-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>CCL18 bindet an die Rezeptoren <a href="/w/index.php?title=PITPNM3&action=edit&redlink=1" class="new" title="PITPNM3 (Seite nicht vorhanden)">PITPNM3</a>, <a href="/w/index.php?title=GPR30&action=edit&redlink=1" class="new" title="GPR30 (Seite nicht vorhanden)">GPR30</a> und <a href="/w/index.php?title=CCR8&action=edit&redlink=1" class="new" title="CCR8 (Seite nicht vorhanden)">CCR8</a> auf anderen Zellen. CCL18 ist <a href="/wiki/Chemotaxis" title="Chemotaxis">chemotaktisch</a> für verschiedene <a href="/wiki/Immunzelle" class="mw-redirect" title="Immunzelle">Immunzellen</a>, darunter naive T-Zellen,<sup id="cite_ref-PMID_9192897_6-0" class="reference"><a href="#cite_note-PMID_9192897-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Regulatorische_T-Zelle" title="Regulatorische T-Zelle">regulatorische T-Zellen</a>,<sup id="cite_ref-PMID23102918_2-2" class="reference"><a href="#cite_note-PMID23102918-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-PMID22649201_7-0" class="reference"><a href="#cite_note-PMID22649201-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> TH2-Zellen,<sup id="cite_ref-PMID16670340_6_8-0" class="reference"><a href="#cite_note-PMID16670340_6-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> immunosuppressive und unreife dendritische Zellen,<sup id="cite_ref-PMID23102918_2-3" class="reference"><a href="#cite_note-PMID23102918-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-PMID12646652_11_5-1" class="reference"><a href="#cite_note-PMID12646652_11-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Basophile" class="mw-redirect" title="Basophile">Basophile</a><sup id="cite_ref-PMID16670340_6_8-1" class="reference"><a href="#cite_note-PMID16670340_6-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> und <a href="/wiki/B-Zelle" class="mw-redirect" title="B-Zelle">B-Zellen</a> (naive und effektor B-Zellen).<sup id="cite_ref-PMID15784687_9-0" class="reference"><a href="#cite_note-PMID15784687-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> Die von CCL18 gebundenen regulatorischen T-Zellen sind <a href="/wiki/FOXP3" class="mw-redirect" title="FOXP3">Foxp3</a>-negativ und sezernieren IL-10.<sup id="cite_ref-PMID22117697_4-1" class="reference"><a href="#cite_note-PMID22117697-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Einzelnachweise">Einzelnachweise</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=CCL18&veaction=edit&section=2" title="Abschnitt bearbeiten: Einzelnachweise" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=CCL18&action=edit&section=2" title="Quellcode des Abschnitts bearbeiten: Einzelnachweise"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text">K. Hieshima, T. Imai, M. Baba, K. Shoudai, K. Ishizuka, T. Nakagawa, J. Tsuruta, M. Takeya, Y. Sakaki, K. Takatsuki, R. Miura, G. Opdenakker, J. Van Damme, O. Yoshie, H. Nomiyama: <i>A novel human CC chemokine PARC that is most homologous to macrophage-inflammatory protein-1 alpha/LD78 alpha and chemotactic for T lymphocytes, but not for monocytes.</i> In: <i>Journal of Immunology (Baltimore, Md. : 1950).</i> Band 159, Nummer 3, August 1997, S. 1140–1149, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/9233607?dopt=Abstract">PMID 9233607</a>.</span> </li> <li id="cite_note-PMID23102918-2"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-PMID23102918_2-0">a</a></sup> <sup><a href="#cite_ref-PMID23102918_2-1">b</a></sup> <sup><a href="#cite_ref-PMID23102918_2-2">c</a></sup> <sup><a href="#cite_ref-PMID23102918_2-3">d</a></sup></span> <span class="reference-text">Bellinghausen I, Reuter S, Martin H, Maxeiner J, Luxemburger U, Türeci Ö, Grabbe S, Taube C, Saloga J: <cite style="font-style:italic">Enhanced production of CCL18 by tolerogenic dendritic cells is associated with inhibition of allergic airway reactivity</cite>. In: <cite style="font-style:italic">J. Allergy Clin. Immunol.</cite> 130. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>6</span>, Dezember 2012, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>1384–93</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1016/j.jaci.2012.08.039">10.1016/j.jaci.2012.08.039</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/23102918?dopt=Abstract">PMID 23102918</a>.<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:CCL18&rft.atitle=Enhanced+production+of+CCL18+by+tolerogenic+dendritic+cells+is+associated+with+inhibition+of+allergic+airway+reactivity&rft.au=Bellinghausen+I%2C+Reuter+S%2C+Martin+H%2C+...&rft.date=2012-12&rft.doi=10.1016%2Fj.jaci.2012.08.039&rft.genre=journal&rft.issue=6&rft.jtitle=J.+Allergy+Clin.+Immunol.&rft.pages=1384-93&rft.pmid=23102918&rft.volume=130.+Jahrgang" style="display:none"> </span></span> </li> <li id="cite_note-PMID18959625-3"><span class="mw-cite-backlink"><a href="#cite_ref-PMID18959625_3-0">↑</a></span> <span class="reference-text">Ferrara G, Bleck B, Richeldi L, Reibman J, Fabbri LM, Rom WN, Condos R: <cite style="font-style:italic">Mycobacterium tuberculosis induces CCL18 expression in human macrophages</cite>. In: <cite style="font-style:italic">Scand. J. Immunol.</cite> 68. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>6</span>, Dezember 2008, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>668–74</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1111/j.1365-3083.2008.02182.x">10.1111/j.1365-3083.2008.02182.x</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/18959625?dopt=Abstract">PMID 18959625</a>.<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:CCL18&rft.atitle=Mycobacterium+tuberculosis+induces+CCL18+expression+in+human+macrophages&rft.au=Ferrara+G%2C+Bleck+B%2C+Richeldi+L%2C+...&rft.date=2008-12&rft.doi=10.1111%2Fj.1365-3083.2008.02182.x&rft.genre=journal&rft.issue=6&rft.jtitle=Scand.+J.+Immunol.&rft.pages=668-74&rft.pmid=18959625&rft.volume=68.+Jahrgang" style="display:none"> </span></span> </li> <li id="cite_note-PMID22117697-4"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-PMID22117697_4-0">a</a></sup> <sup><a href="#cite_ref-PMID22117697_4-1">b</a></sup></span> <span class="reference-text">Schraufstatter IU, Zhao M, Khaldoyanidi SK, Discipio RG: <cite style="font-style:italic">The chemokine CCL18 causes maturation of cultured monocytes to macrophages in the M2 spectrum</cite>. In: <cite style="font-style:italic">Immunology</cite>. 135. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>4</span>, April 2012, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>287–98</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1111/j.1365-2567.2011.03541.x">10.1111/j.1365-2567.2011.03541.x</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/22117697?dopt=Abstract">PMID 22117697</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3372745/">PMC 3372745</a> (freier Volltext).<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:CCL18&rft.atitle=The+chemokine+CCL18+causes+maturation+of+cultured+monocytes+to+macrophages+in+the+M2+spectrum&rft.au=Schraufstatter+IU%2C+Zhao+M%2C+Khaldoyanidi+SK%2C+...&rft.date=2012-04&rft.doi=10.1111%2Fj.1365-2567.2011.03541.x&rft.genre=journal&rft.issue=4&rft.jtitle=Immunology&rft.pages=287-98&rft.pmc=3372745&rft.pmid=22117697&rft.volume=135.+Jahrgang" style="display:none"> </span></span> </li> <li id="cite_note-PMID12646652_11-5"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-PMID12646652_11_5-0">a</a></sup> <sup><a href="#cite_ref-PMID12646652_11_5-1">b</a></sup></span> <span class="reference-text">Vulcano M, Struyf S, Scapini P, Cassatella M, Bernasconi S, Bonecchi R, Calleri A, Penna G, Adorini L, Luini W, Mantovani A, Van Damme J, Sozzani S: <cite style="font-style:italic">Unique regulation of CCL18 production by maturing dendritic cells</cite>. In: <cite style="font-style:italic">J. Immunol.</cite> 170. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>7</span>, April 2003, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>3843–9</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.4049/jimmunol.170.7.3843">10.4049/jimmunol.170.7.3843</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/12646652?dopt=Abstract">PMID 12646652</a>.<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:CCL18&rft.atitle=Unique+regulation+of+CCL18+production+by+maturing+dendritic+cells&rft.au=Vulcano+M%2C+Struyf+S%2C+Scapini+P%2C+...&rft.date=2003-04&rft.doi=10.4049%2Fjimmunol.170.7.3843&rft.genre=journal&rft.issue=7&rft.jtitle=J.+Immunol.&rft.pages=3843-9&rft.pmid=12646652&rft.volume=170.+Jahrgang" style="display:none"> </span></span> </li> <li id="cite_note-PMID_9192897-6"><span class="mw-cite-backlink"><a href="#cite_ref-PMID_9192897_6-0">↑</a></span> <span class="reference-text">Adema GJ, Hartgers F, Verstraten R, de Vries E, Marland G, Menon S, Foster J, Xu Y, Nooyen P, McClanahan T, Bacon KB, Figdor CG: <cite style="font-style:italic">A dendritic-cell-derived C-C chemokine that preferentially attracts naive T cells</cite>. In: <cite style="font-style:italic">Nature</cite>. 387. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>6634</span>, Juni 1997, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>713–7</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1038/42716">10.1038/42716</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/9192897?dopt=Abstract">PMID 9192897</a>.<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:CCL18&rft.atitle=A+dendritic-cell-derived+C-C+chemokine+that+preferentially+attracts+naive+T+cells&rft.au=Adema+GJ%2C+Hartgers+F%2C+Verstraten+R%2C+...&rft.date=1997-06&rft.doi=10.1038%2F42716&rft.genre=journal&rft.issue=6634&rft.jtitle=Nature&rft.pages=713-7&rft.pmid=9192897&rft.volume=387.+Jahrgang" style="display:none"> </span></span> </li> <li id="cite_note-PMID22649201-7"><span class="mw-cite-backlink"><a href="#cite_ref-PMID22649201_7-0">↑</a></span> <span class="reference-text">Chenivesse C, Chang Y, Azzaoui I, Ait Yahia S, Morales O, Plé C, Foussat A, Tonnel AB, Delhem N, Yssel H, Vorng H, Wallaert B, Tsicopoulos A: <cite style="font-style:italic">Pulmonary CCL18 recruits human regulatory T cells</cite>. In: <cite style="font-style:italic">J. Immunol.</cite> 189. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>1</span>, Juli 2012, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>128–37</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.4049/jimmunol.1003616">10.4049/jimmunol.1003616</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/22649201?dopt=Abstract">PMID 22649201</a>.<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:CCL18&rft.atitle=Pulmonary+CCL18+recruits+human+regulatory+T+cells&rft.au=Chenivesse+C%2C+Chang+Y%2C+Azzaoui+I%2C+...&rft.date=2012-07&rft.doi=10.4049%2Fjimmunol.1003616&rft.genre=journal&rft.issue=1&rft.jtitle=J.+Immunol.&rft.pages=128-37&rft.pmid=22649201&rft.volume=189.+Jahrgang" style="display:none"> </span></span> </li> <li id="cite_note-PMID16670340_6-8"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-PMID16670340_6_8-0">a</a></sup> <sup><a href="#cite_ref-PMID16670340_6_8-1">b</a></sup></span> <span class="reference-text">de Nadaï P, Charbonnier AS, Chenivesse C, Sénéchal S, Fournier C, Gilet J, Vorng H, Chang Y, Gosset P, Wallaert B, Tonnel AB, Lassalle P, Tsicopoulos A: <cite style="font-style:italic">Involvement of CCL18 in allergic asthma</cite>. In: <cite style="font-style:italic">J. Immunol.</cite> 176. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em"> </span>10</span>, Mai 2006, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em"> </span>6286–93</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.4049/jimmunol.176.10.6286">10.4049/jimmunol.176.10.6286</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/16670340?dopt=Abstract">PMID 16670340</a>.<span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rfr_id=info:sid/de.wikipedia.org:CCL18&rft.atitle=Involvement+of+CCL18+in+allergic+asthma&rft.au=de+Nada%C3%AF+P%2C+Charbonnier+AS%2C+Chenivesse+C%2C+...&rft.date=2006-05&rft.doi=10.4049%2Fjimmunol.176.10.6286&rft.genre=journal&rft.issue=10&rft.jtitle=J.+Immunol.&rft.pages=6286-93&rft.pmid=16670340&rft.volume=176.+Jahrgang" style="display:none"> </span></span> </li> <li id="cite_note-PMID15784687-9"><span class="mw-cite-backlink"><a href="#cite_ref-PMID15784687_9-0">↑</a></span> <span class="reference-text">E. Schutyser, A. Richmond, J. 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