Patch use, apprehension, and vigilance behavior of Nubian Ibex under perceived risk of predation | Behavioral Ecology

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Foraging theory predicts that animals will sacrifice feeding effort in order to reduce predation risk. 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A test with fox squirrels, Sciurus niger; Ann Zool Fenn; citation_year=1992; citation_volume=29; citation_pages=311-318; " /><meta name="citation_reference" content="citation_title=Antipredator defenses in birds and mammals; citation_publisher=University of Chicago Press, Chicago (IL); citation_year=2005; " /><meta name="citation_reference" content="citation_title=The estimation and analysis of preference and its relationship to foraging models; Ecology; citation_year=1983; citation_volume=64; citation_issue=5; citation_pages=1297-1304; " /><meta name="citation_reference" content="citation_title=Attention, ‘apprehension’ and gerbils searching in patches; Ann Zool Fenn; citation_year=2001; citation_volume=38; citation_pages=15-23; " /><meta name="citation_reference" content="citation_title=Predator vigilance and group size in mammals and birds: a critical review of the empirical evidence; Biol Rev Camb Philos Soc; citation_year=1989; citation_volume=64; citation_issue=1; citation_pages=13-33; " /><meta name="citation_reference" content="citation_title=Vigilance by female Dall's sheep; interactions between predation risk factor; Anim Behav; citation_year=1997; citation_volume=53; citation_pages=799-808; " /><meta name="citation_reference" content="citation_title=Feeding and chewing behaviors of Nubian ibex: compensation for sex-related differences in body size; Func Ecol; citation_year=1995; citation_volume=9; citation_pages=385-393; " /><meta name="citation_reference" content="citation_title=Grouping patterns and spatial segregation by Nubian Ibex; J Arid Environ; citation_year=1995; citation_volume=30; citation_pages=423-439; " /><meta name="citation_reference" content="citation_title=Vigilance behaviour in African ungulates: the role of predation pressure; Behaviour; citation_year=1998; citation_volume=135; citation_pages=195-211; " /><meta name="citation_reference" content="citation_title=Cost of vigilance in foraging ungulates; Anim Behav; citation_year=1994; citation_volume=47; citation_pages=481-484; " /><meta name="citation_reference" content="citation_title=Antipredator constraints on lacting Nubian Ibex; J Mammal; citation_year=1996; citation_volume=77; citation_issue=4; citation_pages=1122-1131; " /><meta name="citation_reference" content="citation_title=Environmental heterogeneity and the coexistence of desert rodents; Ann Rev Ecol Syst; citation_year=1988; citation_volume=19; citation_pages=281-307; " /><meta name="citation_reference" content="citation_title=Apprehension and time allocation in gerbils: the effects of predatory risk and energetic state; Ecology; citation_year=2004; citation_volume=85; citation_pages=917-922; " /><meta name="citation_reference" content="citation_title=Foraging games between gerbils and their predator: temporal dynamics of resource depletion and apprehension in gerbils; Evol Ecol Res; citation_year=2002; citation_volume=4; citation_pages=495-518; " /><meta name="citation_reference" content="citation_title=Applying patch use to assess aspects of foraging behavior in Nubian Ibex; J Wildl Manag; citation_year=1994; citation_volume=58; citation_issue=2; citation_pages=299-307; " /><meta name="citation_reference" content="citation_title=Predation and competition: the interaction of two types of species interactions; Oikos; citation_year=1989; citation_volume=54; citation_issue=2; citation_pages=256-260; " /><meta name="citation_reference" content="citation_title=Wolves, elk, and bison: reestablishing the “landscape of fear” in Yellowstone National Park, USA; Can J Zool; citation_year=2001; citation_volume=79; citation_issue=8; citation_pages=1401-1409; " /><meta name="citation_reference" content="citation_title=Crèche behavior of Nubian Ibex Capra ibex nubiana in the Negev Desert Highlands, Israel; Isr J Zool; citation_year=1991; citation_volume=37; citation_pages=125-137; " /><meta name="citation_reference" content="citation_title=The influence of models of interpretation of vigilance; citation_publisher=Westview Press, Explanation, Evolution and Adaptation. Boulder (CO); Interpretation and explotation in the study of animal behavior; citation_year=1995; citation_volume=Vol 2; citation_pages=246-267; " /><meta name="citation_reference" content="citation_title=Back to the basics of anti-predatory vigilance: the group size effect; Anim Behav; citation_year=1995; citation_volume=49; citation_pages=11-20; " /><meta name="citation_reference" content="citation_title=Behavioral decisions made under the risk of predation: a review and prospectus; Can J Zool; citation_year=1990; citation_volume=68; citation_pages=619-640; " /><meta name="citation_reference" content="citation_title=Foraging-efficiency–predation-risk trade-off in the grey squirrel; Anim Behav; citation_year=1985; citation_volume=33; citation_pages=155-165; " /><meta name="citation_reference" content="citation_title=A Nubian Ibex nursery: crèche or natural trap?; Isr J Zool; citation_year=1995; citation_volume=41; citation_pages=163-174; " /><meta name="citation_reference" content="citation_title=Foraging costs and accessibility of giving-up densities in a swan-pondweed system; Oikos; citation_year=2006; citation_volume=112; citation_pages=353-362; " /><meta name="citation_reference" content="citation_title=Using vigilance behaviour to test whether predation promotes habitat partitioning; Ecology; citation_year=1996; citation_volume=77; citation_issue=6; citation_pages=1880-1887; " /><meta name="citation_reference" content="citation_title=Why individual vigilance declines as group size increases; Anim Behav; citation_year=1996; citation_volume=51; citation_pages=1077-1086; " /><meta name="citation_reference" content="citation_title=Patch assessment in fox squirrels: the role of resource density, patch size and patch boundaries; Am Nat; citation_year=1996; citation_volume=147; citation_issue=3; citation_pages=360-390; " /><meta name="citation_reference" content="citation_title=Predators and prey lifestyles: an evolutionary and ecological overview; citation_publisher=University Press of New England, Hanover (NH); Predation, direct and indirect impacts on aquatic environments; citation_year=1987; citation_pages=203-224; " /><meta name="citation_reference" content="citation_title=Optimal behavior: can forages balance two conflicting demands?; Science; citation_year=1980; citation_volume=210; citation_issue=28; citation_pages=1041-1043; " /><meta name="citation_reference" content="citation_title=Biometry; citation_edition=2nd ed; citation_publisher=W.H. Freeman and Company, New York (NY); citation_year=1981; " /><meta name="citation_reference" content="citation_author=UNESCO; citation_publisher=MAB, Technical Note 7, Paris; Map of the world distribution of arid zones; citation_year=1977; " /><meta name="citation_fulltext_world_readable" content="" /><meta name="citation_pdf_url" content="https://academic.oup.com/beheco/article-pdf/18/2/368/525081/arl087.pdf" /><meta name="description" content="Abstract. Foraging theory predicts that animals will sacrifice feeding effort in order to reduce predation risk. 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class="name-role-wrap"> <div class="info-card-name"> Valeria Hochman </div> </div> <div class="info-card-affilitation"> <div class="aff">Mitriani Department for Desert Ecology, Blaustein Institute for Desert Research, Ben-Gurion University of the Negev, Midreshet Ben-Gurion 84990, Israel</div> </div> <div class="info-card-search-label"> Search for other works by this author on: </div> <div class="info-card-search info-card-search-internal"> <a href="/beheco/search-results?f_Authors=Valeria+Hochman" rel="nofollow">Oxford Academic</a> </div> <div class="info-card-search info-card-search-google"> <a href="http://scholar.google.com/scholar?q=author:%22Hochman Valeria%22">Google Scholar</a> </div> </div> </span> </span> <span class="al-author-name-more js-flyout-wrap"> <button type="button" class="linked-name js-linked-name-trigger btn-as-link">Burt P. Kotler</button><span class='delimiter'></span> <span class="al-author-info-wrap arrow-up"> <div class="info-card-author authorInfo_OUP_Abstract_ArticleTop_Info_Widget"> <div class="name-role-wrap"> <div class="info-card-name"> Burt P. Kotler </div> </div> <div class="info-card-affilitation"> <div class="aff">Mitriani Department for Desert Ecology, Blaustein Institute for Desert Research, Ben-Gurion University of the Negev, Midreshet Ben-Gurion 84990, Israel</div> </div> <div class="info-card-search-label"> Search for other works by this author on: </div> <div class="info-card-search info-card-search-internal"> <a href="/beheco/search-results?f_Authors=Burt+P.+Kotler" rel="nofollow">Oxford Academic</a> </div> <div class="info-card-search info-card-search-google"> <a href="http://scholar.google.com/scholar?q=author:%22Kotler Burt P.%22">Google Scholar</a> </div> </div> </span> </span> </div> </div> <div class="pub-history-wrap clearfix js-history-dropdown-wrap"> <div class="pub-history-row clearfix"> <div class="ww-citation-primary"><em>Behavioral Ecology</em>, Volume 18, Issue 2, March 2007, Pages 368–374, <a href='https://doi.org/10.1093/beheco/arl087'>https://doi.org/10.1093/beheco/arl087</a></div> </div> <div class="pub-history-row clearfix"> <div class="ww-citation-date-wrap"> <div class="citation-label">Published:</div> <div class="citation-date">14 December 2006</div> </div> <a href="javascript:;" class="history-label js-history-dropdown-trigger st-article-history at-ArticleHistory"> <span>Article history</span><i class="icon-general-arrow-filled-down arrow-icon"></i> </a> </div> <div class="ww-history js-history-entries-wrap at-history-entries-wrap"> <div class="history-entry at-history-entry"> <div class="wi-state">Received:</div> <div class="wi-date">18 February 2006</div> </div> <div class="history-entry at-history-entry"> <div class="wi-state">Revision received:</div> <div class="wi-date">11 November 2006</div> </div> <div class="history-entry at-history-entry"> <div class="wi-state">Accepted:</div> <div class="wi-date">17 November 2006</div> </div> <div class="history-entry at-history-entry"> <div class="wi-state">Published:</div> <div class="wi-date">14 December 2006</div> </div> </div> </div> </div> </div> <script> $(document).ready(function () { $('.article-top-widget').on('click', '.ati-toggle-trigger', function () { $(this).find('.icon-general-add, .icon-minus').toggleClass('icon-minus icon-general-add'); $(this).siblings('.ati-toggle-content').toggleClass('hide'); }); // In Chrome, an anchor tag with target="_blank" and a "mailto:" href opens a new tab/window as well as the email client // I suspect this behavior will be corrected in the future // Remove the target="_blank" $('ul.wi-affiliationList').find('a[href^="mailto:"]').each(function () { $(this).removeAttr('target'); }); }); </script> </div> <div class="widget widget-ArticleLinks widget-instance-OUP_Article_Links_Widget"> </div> <div class="article-body js-content-body"> <div class="toolbar-wrap js-toolbar-wrap"> <div class="toolbar-inner-wrap"> <ul id="Toolbar" role="navigation"> <li class="toolbar-item item-with-dropdown item-views"> <a class="at-views-dropdown drop-trigger" href="javascript:;" data-dropdown="FilterDrop" aria-haspopup="true"> <i class="icon-menu_views"></i> <div class="toolbar-label"> <div class="toolbar-text">Views</div> <i class="icon-general-arrow-filled-down arrow-icon"></i> </div> </a> <ul id="ViewsDrop" class="f-dropdown js-dropdown-content" data-dropdown-content aria-label="submenu"> <div class="arrow-up"></div> <li class="article-content-filter js-article-content-filter" data-content-filter="article-content"> <a href="javascript:;"><span>Article contents</span></a> </li> <li class="at-figures-tables article-content-filter js-article-content-filter" data-content-filter="figures-tables"> <a href="javascript:;"><span>Figures & tables</span></a> </li> <li class="article-content-filter js-article-content-filter" data-content-filter="video"> <a href="javascript:;"><span>Video</span></a> </li> <li class="article-content-filter js-article-content-filter" data-content-filter="audio"> <a href="javascript:;"><span>Audio</span></a> </li> <li class="article-content-filter js-article-content-filter" data-content-filter="supplementary-data"> <a href="javascript:;"><span>Supplementary Data</span></a> </li> </ul> </li> <li class="toolbar-item item-cite js-item-cite"> <div class="widget widget-ToolboxGetCitation widget-instance-OUP_Get_Citation"> <a href="#" class="js-cite-button at-CiteButton" data-reveal-id="getCitation" data-reveal> <i class="icon-read-more"></i> <span>Cite</span> </a> <div id="getCitation" class="reveal-modal js-citation-modal" data-reveal> <h3 class="modal-title">Cite</h3> <div class="oxford-citation-text"> <p>Valeria Hochman, Burt P. 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class="abstract-title js-splitscreen-abstract-title" >Abstract</h2> <section class="abstract"><p class="chapter-para">Foraging theory predicts that animals will sacrifice feeding effort in order to reduce predation risk. Once a forager chooses a habitat, it must decide how to allocate its foraging effort. Nubian Ibex are diurnal, social, cliff-dwelling herbivores. Many of their characteristics seem to have evolved as responses to predation risk. In order to assess the effects that perceived risk of predation might have on foraging behavior of free-ranging Nubian Ibex in the Negev Desert, Israel, we measured giving-up densities (GUDs) in artificial food patches and used them to gauge apprehension level. (Apprehension can be defined as a reduction in attention devoted to performing an activity as a consequence of reallocating attention to detecting or responding to predation risk. A forager can also be vigilant. Vigilance is often defined as time spent scanning the surroundings with the head up.) We also quantified time budgeting using focal observation of individual Nubian Ibex. Habitat preferences and patch selectivity as a measure of apprehension were considered. In particular, we tested the effect of distance from refuge on GUDs, the effect of micropatch structure on selectivity, and the effect of distance from the refuge and group size on Nubian Ibex vigilance level and apprehension. Nubian Ibex allocate their foraging effort more toward patches closer to the escape terrain. At the same time, Nubian Ibex are more apprehensive at intermediate distances from the cliff edge than nearer the cliff, and their use of vigilance increases with distance from the cliff edge. These results suggest that Nubian Ibex may switch from apprehension to a more extreme behavior of vigilance at greater distances from the refuge. This study demonstrated the use of antipredatory behaviors, apprehension, and vigilance by a forager. Estimating apprehension and vigilance levels of a forager simultaneously gives a more complete and accurate picture of how the habitat is perceived by them and combined with measurements of GUD allow a more accurate assessment of habitat quality.</p></section> <div class="article-metadata-panel clearfix at-ArticleMetadata"></div>  <div class="widget widget-ArticlePubStateInfo widget-instance-OUP_ArticlePubStateInfo"> </div> <div class="article-metadata-standalone-panel clearfix"></div> <div class="copyright copyright-statement">© The Author 2006. Published by Oxford University Press on behalf of the International Society for Behavioral Ecology. All rights reserved. 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Patch use, apprehension, and vigilance behavior of Nubian Ibex under perceived risk of predation | Behavioral Ecology | Oxford Academic