CINXE.COM
Edinger–Westphal nucleus - Wikipedia
<!DOCTYPE html> <html class="client-nojs vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available" lang="en" dir="ltr"> <head> <meta charset="UTF-8"> <title>Edinger–Westphal nucleus - Wikipedia</title> <script>(function(){var className="client-js vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available";var cookie=document.cookie.match(/(?:^|; )enwikimwclientpreferences=([^;]+)/);if(cookie){cookie[1].split('%2C').forEach(function(pref){className=className.replace(new RegExp('(^| )'+pref.replace(/-clientpref-\w+$|[^\w-]+/g,'')+'-clientpref-\\w+( |$)'),'$1'+pref+'$2');});}document.documentElement.className=className;}());RLCONF={"wgBreakFrames":false,"wgSeparatorTransformTable":["",""],"wgDigitTransformTable":["",""],"wgDefaultDateFormat":"dmy", "wgMonthNames":["","January","February","March","April","May","June","July","August","September","October","November","December"],"wgRequestId":"1770d781-c771-421b-84fb-4c7580284481","wgCanonicalNamespace":"","wgCanonicalSpecialPageName":false,"wgNamespaceNumber":0,"wgPageName":"Edinger–Westphal_nucleus","wgTitle":"Edinger–Westphal nucleus","wgCurRevisionId":1251473278,"wgRevisionId":1251473278,"wgArticleId":2062028,"wgIsArticle":true,"wgIsRedirect":false,"wgAction":"view","wgUserName":null,"wgUserGroups":["*"],"wgCategories":["CS1 maint: multiple names: authors list","Articles with short description","Short description matches Wikidata","Articles to be expanded from August 2022","All articles to be expanded","Cranial nerve nuclei","Oculomotor nerve","Midbrain"],"wgPageViewLanguage":"en","wgPageContentLanguage":"en","wgPageContentModel":"wikitext","wgRelevantPageName":"Edinger–Westphal_nucleus","wgRelevantArticleId":2062028,"wgIsProbablyEditable":true, "wgRelevantPageIsProbablyEditable":true,"wgRestrictionEdit":[],"wgRestrictionMove":[],"wgNoticeProject":"wikipedia","wgCiteReferencePreviewsActive":false,"wgFlaggedRevsParams":{"tags":{"status":{"levels":1}}},"wgMediaViewerOnClick":true,"wgMediaViewerEnabledByDefault":true,"wgPopupsFlags":0,"wgVisualEditor":{"pageLanguageCode":"en","pageLanguageDir":"ltr","pageVariantFallbacks":"en"},"wgMFDisplayWikibaseDescriptions":{"search":true,"watchlist":true,"tagline":false,"nearby":true},"wgWMESchemaEditAttemptStepOversample":false,"wgWMEPageLength":10000,"wgRelatedArticlesCompat":[],"wgCentralAuthMobileDomain":false,"wgEditSubmitButtonLabelPublish":true,"wgULSPosition":"interlanguage","wgULSisCompactLinksEnabled":false,"wgVector2022LanguageInHeader":true,"wgULSisLanguageSelectorEmpty":false,"wgWikibaseItemId":"Q165312","wgCheckUserClientHintsHeadersJsApi":["brands","architecture","bitness","fullVersionList","mobile","model","platform","platformVersion"],"GEHomepageSuggestedEditsEnableTopics": true,"wgGETopicsMatchModeEnabled":false,"wgGEStructuredTaskRejectionReasonTextInputEnabled":false,"wgGELevelingUpEnabledForUser":false};RLSTATE={"ext.globalCssJs.user.styles":"ready","site.styles":"ready","user.styles":"ready","ext.globalCssJs.user":"ready","user":"ready","user.options":"loading","ext.cite.styles":"ready","mediawiki.page.gallery.styles":"ready","skins.vector.search.codex.styles":"ready","skins.vector.styles":"ready","skins.vector.icons":"ready","jquery.makeCollapsible.styles":"ready","ext.wikimediamessages.styles":"ready","ext.visualEditor.desktopArticleTarget.noscript":"ready","ext.uls.interlanguage":"ready","wikibase.client.init":"ready","ext.wikimediaBadges":"ready"};RLPAGEMODULES=["ext.cite.ux-enhancements","site","mediawiki.page.ready","jquery.makeCollapsible","mediawiki.toc","skins.vector.js","ext.centralNotice.geoIP","ext.centralNotice.startUp","ext.gadget.ReferenceTooltips","ext.gadget.switcher","ext.urlShortener.toolbar","ext.centralauth.centralautologin", "mmv.bootstrap","ext.popups","ext.visualEditor.desktopArticleTarget.init","ext.visualEditor.targetLoader","ext.echo.centralauth","ext.eventLogging","ext.wikimediaEvents","ext.navigationTiming","ext.uls.interface","ext.cx.eventlogging.campaigns","ext.cx.uls.quick.actions","wikibase.client.vector-2022","ext.checkUser.clientHints","ext.growthExperiments.SuggestedEditSession","wikibase.sidebar.tracking"];</script> <script>(RLQ=window.RLQ||[]).push(function(){mw.loader.impl(function(){return["user.options@12s5i",function($,jQuery,require,module){mw.user.tokens.set({"patrolToken":"+\\","watchToken":"+\\","csrfToken":"+\\"}); }];});});</script> <link rel="stylesheet" href="/w/load.php?lang=en&modules=ext.cite.styles%7Cext.uls.interlanguage%7Cext.visualEditor.desktopArticleTarget.noscript%7Cext.wikimediaBadges%7Cext.wikimediamessages.styles%7Cjquery.makeCollapsible.styles%7Cmediawiki.page.gallery.styles%7Cskins.vector.icons%2Cstyles%7Cskins.vector.search.codex.styles%7Cwikibase.client.init&only=styles&skin=vector-2022"> <script async="" src="/w/load.php?lang=en&modules=startup&only=scripts&raw=1&skin=vector-2022"></script> <meta name="ResourceLoaderDynamicStyles" content=""> <link rel="stylesheet" href="/w/load.php?lang=en&modules=site.styles&only=styles&skin=vector-2022"> <meta name="generator" content="MediaWiki 1.44.0-wmf.4"> <meta name="referrer" content="origin"> <meta name="referrer" content="origin-when-cross-origin"> <meta name="robots" content="max-image-preview:standard"> <meta name="format-detection" content="telephone=no"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Cn3nucleus-en.svg/1200px-Cn3nucleus-en.svg.png"> <meta property="og:image:width" content="1200"> <meta property="og:image:height" content="732"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Cn3nucleus-en.svg/800px-Cn3nucleus-en.svg.png"> <meta property="og:image:width" content="800"> <meta property="og:image:height" content="488"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Cn3nucleus-en.svg/640px-Cn3nucleus-en.svg.png"> <meta property="og:image:width" content="640"> <meta property="og:image:height" content="391"> <meta name="viewport" content="width=1120"> <meta property="og:title" content="Edinger–Westphal nucleus - Wikipedia"> <meta property="og:type" content="website"> <link rel="preconnect" href="//upload.wikimedia.org"> <link rel="alternate" media="only screen and (max-width: 640px)" href="//en.m.wikipedia.org/wiki/Edinger%E2%80%93Westphal_nucleus"> <link rel="alternate" type="application/x-wiki" title="Edit this page" href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit"> <link rel="apple-touch-icon" href="/static/apple-touch/wikipedia.png"> <link rel="icon" href="/static/favicon/wikipedia.ico"> <link rel="search" type="application/opensearchdescription+xml" href="/w/rest.php/v1/search" title="Wikipedia (en)"> <link rel="EditURI" type="application/rsd+xml" href="//en.wikipedia.org/w/api.php?action=rsd"> <link rel="canonical" href="https://en.wikipedia.org/wiki/Edinger%E2%80%93Westphal_nucleus"> <link rel="license" href="https://creativecommons.org/licenses/by-sa/4.0/deed.en"> <link rel="alternate" type="application/atom+xml" title="Wikipedia Atom feed" href="/w/index.php?title=Special:RecentChanges&feed=atom"> <link rel="dns-prefetch" href="//meta.wikimedia.org" /> <link rel="dns-prefetch" href="//login.wikimedia.org"> </head> <body class="skin--responsive skin-vector skin-vector-search-vue mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject mw-editable page-Edinger–Westphal_nucleus rootpage-Edinger–Westphal_nucleus skin-vector-2022 action-view"><a class="mw-jump-link" href="#bodyContent">Jump to content</a> <div class="vector-header-container"> <header class="vector-header mw-header"> <div class="vector-header-start"> <nav class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-dropdown" class="vector-dropdown vector-main-menu-dropdown vector-button-flush-left vector-button-flush-right" > <input type="checkbox" id="vector-main-menu-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-main-menu-dropdown" class="vector-dropdown-checkbox " aria-label="Main menu" > <label id="vector-main-menu-dropdown-label" for="vector-main-menu-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-menu mw-ui-icon-wikimedia-menu"></span> <span class="vector-dropdown-label-text">Main menu</span> </label> <div class="vector-dropdown-content"> <div id="vector-main-menu-unpinned-container" class="vector-unpinned-container"> <div id="vector-main-menu" class="vector-main-menu vector-pinnable-element"> <div class="vector-pinnable-header vector-main-menu-pinnable-header vector-pinnable-header-unpinned" data-feature-name="main-menu-pinned" data-pinnable-element-id="vector-main-menu" data-pinned-container-id="vector-main-menu-pinned-container" data-unpinned-container-id="vector-main-menu-unpinned-container" > <div class="vector-pinnable-header-label">Main menu</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-main-menu.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-main-menu.unpin">hide</button> </div> <div id="p-navigation" class="vector-menu mw-portlet mw-portlet-navigation" > <div class="vector-menu-heading"> Navigation </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-mainpage-description" class="mw-list-item"><a href="/wiki/Main_Page" title="Visit the main page [z]" accesskey="z"><span>Main page</span></a></li><li id="n-contents" class="mw-list-item"><a href="/wiki/Wikipedia:Contents" title="Guides to browsing Wikipedia"><span>Contents</span></a></li><li id="n-currentevents" class="mw-list-item"><a href="/wiki/Portal:Current_events" title="Articles related to current events"><span>Current events</span></a></li><li id="n-randompage" class="mw-list-item"><a href="/wiki/Special:Random" title="Visit a randomly selected article [x]" accesskey="x"><span>Random article</span></a></li><li id="n-aboutsite" class="mw-list-item"><a href="/wiki/Wikipedia:About" title="Learn about Wikipedia and how it works"><span>About Wikipedia</span></a></li><li id="n-contactpage" class="mw-list-item"><a href="//en.wikipedia.org/wiki/Wikipedia:Contact_us" title="How to contact Wikipedia"><span>Contact us</span></a></li> </ul> </div> </div> <div id="p-interaction" class="vector-menu mw-portlet mw-portlet-interaction" > <div class="vector-menu-heading"> Contribute </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-help" class="mw-list-item"><a href="/wiki/Help:Contents" title="Guidance on how to use and edit Wikipedia"><span>Help</span></a></li><li id="n-introduction" class="mw-list-item"><a href="/wiki/Help:Introduction" title="Learn how to edit Wikipedia"><span>Learn to edit</span></a></li><li id="n-portal" class="mw-list-item"><a href="/wiki/Wikipedia:Community_portal" title="The hub for editors"><span>Community portal</span></a></li><li id="n-recentchanges" class="mw-list-item"><a href="/wiki/Special:RecentChanges" title="A list of recent changes to Wikipedia [r]" accesskey="r"><span>Recent changes</span></a></li><li id="n-upload" class="mw-list-item"><a href="/wiki/Wikipedia:File_upload_wizard" title="Add images or other media for use on Wikipedia"><span>Upload file</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> <a href="/wiki/Main_Page" class="mw-logo"> <img class="mw-logo-icon" src="/static/images/icons/wikipedia.png" alt="" aria-hidden="true" height="50" width="50"> <span class="mw-logo-container skin-invert"> <img class="mw-logo-wordmark" alt="Wikipedia" src="/static/images/mobile/copyright/wikipedia-wordmark-en.svg" style="width: 7.5em; height: 1.125em;"> <img class="mw-logo-tagline" alt="The Free Encyclopedia" src="/static/images/mobile/copyright/wikipedia-tagline-en.svg" width="117" height="13" style="width: 7.3125em; height: 0.8125em;"> </span> </a> </div> <div class="vector-header-end"> <div id="p-search" role="search" class="vector-search-box-vue vector-search-box-collapses vector-search-box-show-thumbnail vector-search-box-auto-expand-width vector-search-box"> <a href="/wiki/Special:Search" class="cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only search-toggle" title="Search Wikipedia [f]" accesskey="f"><span class="vector-icon mw-ui-icon-search mw-ui-icon-wikimedia-search"></span> <span>Search</span> </a> <div class="vector-typeahead-search-container"> <div class="cdx-typeahead-search cdx-typeahead-search--show-thumbnail cdx-typeahead-search--auto-expand-width"> <form action="/w/index.php" id="searchform" class="cdx-search-input cdx-search-input--has-end-button"> <div id="simpleSearch" class="cdx-search-input__input-wrapper" data-search-loc="header-moved"> <div class="cdx-text-input cdx-text-input--has-start-icon"> <input class="cdx-text-input__input" type="search" name="search" placeholder="Search Wikipedia" aria-label="Search Wikipedia" autocapitalize="sentences" title="Search Wikipedia [f]" accesskey="f" id="searchInput" > <span class="cdx-text-input__icon cdx-text-input__start-icon"></span> </div> <input type="hidden" name="title" value="Special:Search"> </div> <button class="cdx-button cdx-search-input__end-button">Search</button> </form> </div> </div> </div> <nav class="vector-user-links vector-user-links-wide" aria-label="Personal tools"> <div class="vector-user-links-main"> <div id="p-vector-user-menu-preferences" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-userpage" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-dropdown" class="vector-dropdown " title="Change the appearance of the page's font size, width, and color" > <input type="checkbox" id="vector-appearance-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-appearance-dropdown" class="vector-dropdown-checkbox " aria-label="Appearance" > <label id="vector-appearance-dropdown-label" for="vector-appearance-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-appearance mw-ui-icon-wikimedia-appearance"></span> <span class="vector-dropdown-label-text">Appearance</span> </label> <div class="vector-dropdown-content"> <div id="vector-appearance-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <div id="p-vector-user-menu-notifications" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-overflow" class="vector-menu mw-portlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="https://donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_en.wikipedia.org&uselang=en" class=""><span>Donate</span></a> </li> <li id="pt-createaccount-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:CreateAccount&returnto=Edinger%E2%80%93Westphal+nucleus" title="You are encouraged to create an account and log in; however, it is not mandatory" class=""><span>Create account</span></a> </li> <li id="pt-login-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:UserLogin&returnto=Edinger%E2%80%93Westphal+nucleus" title="You're encouraged to log in; however, it's not mandatory. [o]" accesskey="o" class=""><span>Log in</span></a> </li> </ul> </div> </div> </div> <div id="vector-user-links-dropdown" class="vector-dropdown vector-user-menu vector-button-flush-right vector-user-menu-logged-out" title="Log in and more options" > <input type="checkbox" id="vector-user-links-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-user-links-dropdown" class="vector-dropdown-checkbox " aria-label="Personal tools" > <label id="vector-user-links-dropdown-label" for="vector-user-links-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-ellipsis mw-ui-icon-wikimedia-ellipsis"></span> <span class="vector-dropdown-label-text">Personal tools</span> </label> <div class="vector-dropdown-content"> <div id="p-personal" class="vector-menu mw-portlet mw-portlet-personal user-links-collapsible-item" title="User menu" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport" class="user-links-collapsible-item mw-list-item"><a href="https://donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_en.wikipedia.org&uselang=en"><span>Donate</span></a></li><li id="pt-createaccount" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:CreateAccount&returnto=Edinger%E2%80%93Westphal+nucleus" title="You are encouraged to create an account and log in; however, it is not mandatory"><span class="vector-icon mw-ui-icon-userAdd mw-ui-icon-wikimedia-userAdd"></span> <span>Create account</span></a></li><li id="pt-login" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:UserLogin&returnto=Edinger%E2%80%93Westphal+nucleus" title="You're encouraged to log in; however, it's not mandatory. [o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>Log in</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> Pages for logged out editors <a href="/wiki/Help:Introduction" aria-label="Learn more about editing"><span>learn more</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/Special:MyContributions" title="A list of edits made from this IP address [y]" accesskey="y"><span>Contributions</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/Special:MyTalk" title="Discussion about edits from this IP address [n]" accesskey="n"><span>Talk</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"><!-- CentralNotice --></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="Contents" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Contents</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">hide</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">(Top)</div> </a> </li> <li id="toc-Structure" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Structure"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Structure</span> </div> </a> <button aria-controls="toc-Structure-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Structure subsection</span> </button> <ul id="toc-Structure-sublist" class="vector-toc-list"> <li id="toc-Anatomical_relations" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Anatomical_relations"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.1</span> <span>Anatomical relations</span> </div> </a> <ul id="toc-Anatomical_relations-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Function" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Function"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Function</span> </div> </a> <button aria-controls="toc-Function-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Function subsection</span> </button> <ul id="toc-Function-sublist" class="vector-toc-list"> <li id="toc-Neurophysiology" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Neurophysiology"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1</span> <span>Neurophysiology</span> </div> </a> <ul id="toc-Neurophysiology-sublist" class="vector-toc-list"> <li id="toc-Pupillary_light_reflex" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Pupillary_light_reflex"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1.1</span> <span>Pupillary light reflex</span> </div> </a> <ul id="toc-Pupillary_light_reflex-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> </ul> </li> <li id="toc-Research" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Research"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Research</span> </div> </a> <ul id="toc-Research-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Eponym" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Eponym"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Eponym</span> </div> </a> <ul id="toc-Eponym-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Additional_images" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Additional_images"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Additional images</span> </div> </a> <ul id="toc-Additional_images-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Edinger–Westphal nucleus</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 15 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-15" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">15 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%86%D9%88%D8%A7%D8%A9_%D8%A5%D9%8A%D8%AF%D9%86%D8%BA%D8%B1%E2%80%93%D9%88%D9%8A%D8%B3%D8%AA%D9%81%D8%A7%D9%84" title="نواة إيدنغر–ويستفال – Arabic" lang="ar" hreflang="ar" data-title="نواة إيدنغر–ويستفال" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Edinger-Westphal-Kern" title="Edinger-Westphal-Kern – German" lang="de" hreflang="de" data-title="Edinger-Westphal-Kern" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/N%C3%BAcleo_de_Edinger-Westphal" title="Núcleo de Edinger-Westphal – Spanish" lang="es" hreflang="es" data-title="Núcleo de Edinger-Westphal" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Edinger%E2%80%93Westphal_nukleo" title="Edinger–Westphal nukleo – Basque" lang="eu" hreflang="eu" data-title="Edinger–Westphal nukleo" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Noyau_d%27Edinger-Westphal" title="Noyau d'Edinger-Westphal – French" lang="fr" hreflang="fr" data-title="Noyau d'Edinger-Westphal" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%97%90%EB%94%A9%EA%B1%B0-%EB%B2%A0%EC%8A%A4%ED%8A%B8%ED%8C%94_%ED%95%B5" title="에딩거-베스트팔 핵 – Korean" lang="ko" hreflang="ko" data-title="에딩거-베스트팔 핵" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Nucleo_di_Edinger-Westphal" title="Nucleo di Edinger-Westphal – Italian" lang="it" hreflang="it" data-title="Nucleo di Edinger-Westphal" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Nucleus_accessorius_nervi_oculomotorii" title="Nucleus accessorius nervi oculomotorii – Latin" lang="la" hreflang="la" data-title="Nucleus accessorius nervi oculomotorii" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Kern_van_Edinger-Westphal" title="Kern van Edinger-Westphal – Dutch" lang="nl" hreflang="nl" data-title="Kern van Edinger-Westphal" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%A8%E3%83%87%E3%82%A3%E3%83%B3%E3%82%AC%E3%83%BC%E3%83%BB%E3%82%A6%E3%82%A7%E3%82%B9%E3%83%88%E3%83%95%E3%82%A1%E3%83%AB%E6%A0%B8" title="エディンガー・ウェストファル核 – Japanese" lang="ja" hreflang="ja" data-title="エディンガー・ウェストファル核" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/J%C4%85dro_Edingera-Westphala" title="Jądro Edingera-Westphala – Polish" lang="pl" hreflang="pl" data-title="Jądro Edingera-Westphala" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/N%C3%BAcleo_de_Edinger-Westphal" title="Núcleo de Edinger-Westphal – Portuguese" lang="pt" hreflang="pt" data-title="Núcleo de Edinger-Westphal" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%AF%D0%B4%D1%80%D0%BE_%D0%AD%D0%B4%D0%B8%D0%BD%D0%B3%D0%B5%D1%80%D0%B0-%D0%92%D0%B5%D1%81%D1%82%D1%84%D0%B0%D0%BB%D1%8F" title="Ядро Эдингера-Вестфаля – Russian" lang="ru" hreflang="ru" data-title="Ядро Эдингера-Вестфаля" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%AF%D0%B4%D1%80%D0%BE_%D0%95%D0%B4%D1%96%D0%BD%D0%B3%D0%B5%D1%80%D0%B0-%D0%92%D0%B5%D1%81%D1%82%D1%84%D0%B0%D0%BB%D1%8F" title="Ядро Едінгера-Вестфаля – Ukrainian" lang="uk" hreflang="uk" data-title="Ядро Едінгера-Вестфаля" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%8B%95%E7%9C%BC%E7%A5%9E%E7%B6%93%E5%89%AF%E6%A0%B8" title="動眼神經副核 – Chinese" lang="zh" hreflang="zh" data-title="動眼神經副核" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q165312#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Edinger%E2%80%93Westphal_nucleus" title="View the content page [c]" accesskey="c"><span>Article</span></a></li><li id="ca-talk" class="vector-tab-noicon mw-list-item"><a href="/wiki/Talk:Edinger%E2%80%93Westphal_nucleus" rel="discussion" title="Discuss improvements to the content page [t]" accesskey="t"><span>Talk</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="Change language variant" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">English</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="Views"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Edinger%E2%80%93Westphal_nucleus"><span>Read</span></a></li><li id="ca-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit" title="Edit this page [e]" accesskey="e"><span>Edit</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=history" title="Past revisions of this page [h]" accesskey="h"><span>View history</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="Tools" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Tools</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">Tools</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">hide</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="More options" > <div class="vector-menu-heading"> Actions </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/Edinger%E2%80%93Westphal_nucleus"><span>Read</span></a></li><li id="ca-more-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit" title="Edit this page [e]" accesskey="e"><span>Edit</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=history"><span>View history</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> General </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/Special:WhatLinksHere/Edinger%E2%80%93Westphal_nucleus" title="List of all English Wikipedia pages containing links to this page [j]" accesskey="j"><span>What links here</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Special:RecentChangesLinked/Edinger%E2%80%93Westphal_nucleus" rel="nofollow" title="Recent changes in pages linked from this page [k]" accesskey="k"><span>Related changes</span></a></li><li id="t-upload" class="mw-list-item"><a href="/wiki/Wikipedia:File_Upload_Wizard" title="Upload files [u]" accesskey="u"><span>Upload file</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Special:SpecialPages" title="A list of all special pages [q]" accesskey="q"><span>Special pages</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&oldid=1251473278" title="Permanent link to this revision of this page"><span>Permanent link</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=info" title="More information about this page"><span>Page information</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=Special:CiteThisPage&page=Edinger%E2%80%93Westphal_nucleus&id=1251473278&wpFormIdentifier=titleform" title="Information on how to cite this page"><span>Cite this page</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=Special:UrlShortener&url=https%3A%2F%2Fen.wikipedia.org%2Fwiki%2FEdinger%25E2%2580%2593Westphal_nucleus"><span>Get shortened URL</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=Special:QrCode&url=https%3A%2F%2Fen.wikipedia.org%2Fwiki%2FEdinger%25E2%2580%2593Westphal_nucleus"><span>Download QR code</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> Print/export </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=Special:DownloadAsPdf&page=Edinger%E2%80%93Westphal_nucleus&action=show-download-screen" title="Download this page as a PDF file"><span>Download as PDF</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&printable=yes" title="Printable version of this page [p]" accesskey="p"><span>Printable version</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> In other projects </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q165312" title="Structured data on this page hosted by Wikidata [g]" accesskey="g"><span>Wikidata item</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Appearance</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">hide</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Not to be confused with <a href="/wiki/Accessory_oculomotor_nuclei" title="Accessory oculomotor nuclei">Accessory oculomotor nuclei</a>.</div><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">One of two nuclei of the oculomotor nerve</div> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox"><tbody><tr><th colspan="2" class="infobox-above" style="background-color:lemonchiffon">Edinger–Westphal nucleus</th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Cn3nucleus-en.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Cn3nucleus-en.svg/250px-Cn3nucleus-en.svg.png" decoding="async" width="250" height="153" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Cn3nucleus-en.svg/375px-Cn3nucleus-en.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Cn3nucleus-en.svg/500px-Cn3nucleus-en.svg.png 2x" data-file-width="770" data-file-height="470" /></a></span><div class="infobox-caption">Section through superior colliculus (unlabeled) showing path of <a href="/wiki/Oculomotor_nerve" title="Oculomotor nerve">oculomotor nerve</a>.</div></td></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Gray762.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Gray762.png/250px-Gray762.png" decoding="async" width="250" height="339" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/a/a9/Gray762.png 1.5x" data-file-width="295" data-file-height="400" /></a></span><div class="infobox-caption">Figure showing the different groups of cells, which constitute, according to Perlia, the nucleus of origin of the oculomotor nerve.<br />1. Posterior dorsal nucleus.<br />1’. Posterior ventral nucleus.<br />2. Anterior dorsal nucleus.<br />2’. Anterior ventral nucleus.<br />3. Central nucleus.<br />4. <b>Nucleus of Edinger and Westphal</b>.<br />5. Antero-internal nucleus.<br />6. Antero-external nucleus.<br />8. Crossed fibers.<br />9. <a href="/wiki/Trochlear_nerve" title="Trochlear nerve">Trochlear nerve</a>, with 9’, its nucleus of origin, and 9", its <a href="/wiki/Decussation" title="Decussation">decussation</a>.<br />10. <a href="/wiki/Third_ventricle" title="Third ventricle">Third ventricle</a>.<br />M, M. Median line.</div></td></tr><tr><th colspan="2" class="infobox-header" style="background-color: #efefef">Details</th></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em">Parts</th><td class="infobox-data">Provides input to <a href="/wiki/Parasympathetic_root_of_ciliary_ganglion" class="mw-redirect" title="Parasympathetic root of ciliary ganglion">parasympathetic root of ciliary ganglion</a></td></tr><tr><th colspan="2" class="infobox-header" style="background-color: #efefef">Identifiers</th></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Latin" title="Latin">Latin</a></th><td class="infobox-data"><i>nuclei accessorii nervi oculomotorii</i></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Medical_Subject_Headings" title="Medical Subject Headings">MeSH</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://meshb.nlm.nih.gov/record/ui?ui=D065839">D065839</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/NeuroNames" title="NeuroNames">NeuroNames</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="http://braininfo.rprc.washington.edu/centraldirectory.aspx?ID=498">498</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/NeuroLex" title="NeuroLex">NeuroLex</a> ID</th><td class="infobox-data"><a rel="nofollow" class="external text" href="http://uri.neuinfo.org/nif/nifstd/birnlex_822">birnlex_822</a></td></tr><tr><td colspan="2" class="infobox-below"><a href="/wiki/Anatomical_terms_of_neuroanatomy" title="Anatomical terms of neuroanatomy">Anatomical terms of neuroanatomy</a><div style="text-align: right;"><small class="noprint">[<a href="https://www.wikidata.org/wiki/Q165312" class="extiw" title="d:Q165312">edit on Wikidata</a>]</small></div></td></tr></tbody></table> <p>The <b>Edinger–Westphal nucleus</b> also called the <b>accessory</b> or <b>visceral oculomotor nerve</b>, is one of the two <a href="/wiki/Oculomotor_nerve#Nuclei" title="Oculomotor nerve">nuclei of the oculomotor nerve</a> (CN III) located in the <a href="/wiki/Midbrain" title="Midbrain">midbrain</a>.<sup id="cite_ref-Joyce-2022_1-0" class="reference"><a href="#cite_note-Joyce-2022-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Sonne-2022_2-0" class="reference"><a href="#cite_note-Sonne-2022-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Heiland_Hogan-2022_3-0" class="reference"><a href="#cite_note-Heiland_Hogan-2022-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> It receives afferents from both <a href="/wiki/Pretectal_area" title="Pretectal area">pretectal nuclei</a> (which have in turn received afferents from the <a href="/wiki/Optic_tract" title="Optic tract">optic tract</a>).<sup id="cite_ref-Yoo-2023_4-0" class="reference"><a href="#cite_note-Yoo-2023-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> It contains <a href="/wiki/Parasympathetic_nervous_system" title="Parasympathetic nervous system">parasympathetic</a> pre-ganglionic neuron cell bodies that synapse in the <a href="/wiki/Ciliary_ganglion" title="Ciliary ganglion">ciliary ganglion</a>.<sup id="cite_ref-Heiland_Hogan-2022_3-1" class="reference"><a href="#cite_note-Heiland_Hogan-2022-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Sonne-2022_2-1" class="reference"><a href="#cite_note-Sonne-2022-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> It contributes the autonomic, parasympathetic component to the oculomotor nerve (CN III),<sup id="cite_ref-Yoo-2023_4-1" class="reference"><a href="#cite_note-Yoo-2023-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> ultimately providing innervation to the <a href="/wiki/Iris_sphincter_muscle" title="Iris sphincter muscle">iris sphincter muscle</a> and <a href="/wiki/Ciliary_muscle" title="Ciliary muscle">ciliary muscle</a> to mediate the <a href="/wiki/Pupillary_light_reflex" title="Pupillary light reflex">pupillary light reflex</a> and <a href="/wiki/Accommodation_(eye)" class="mw-redirect" title="Accommodation (eye)">accommodation</a>, respectively.<sup id="cite_ref-Sonne-2022_2-2" class="reference"><a href="#cite_note-Sonne-2022-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Heiland_Hogan-2022_3-2" class="reference"><a href="#cite_note-Heiland_Hogan-2022-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p>The Edinger–Westphal nucleus has two parts. The first is of preganglionic fibers (EWpg) that terminate in the ciliary ganglion. The second is of centrally projecting cells (EWcp) that project to a number of <a href="/wiki/Brainstem" title="Brainstem">brainstem</a> structures.<sup id="cite_ref-Haines_2018_5-0" class="reference"><a href="#cite_note-Haines_2018-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Heiland_Hogan-2022_3-3" class="reference"><a href="#cite_note-Heiland_Hogan-2022-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-pmid21452224_6-0" class="reference"><a href="#cite_note-pmid21452224-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Structure">Structure</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=1" title="Edit section: Structure"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The Edinger–Westphal nucleus refers to the adjacent population of non-preganglionic neurons that do not project to the <a href="/wiki/Ciliary_ganglion" title="Ciliary ganglion">ciliary ganglion</a>, but rather project to the <a href="/wiki/Spinal_cord" title="Spinal cord">spinal cord</a>, <a href="/wiki/Dorsal_raphe_nucleus" title="Dorsal raphe nucleus">dorsal raphe nucleus</a>, lateral <a href="/wiki/Septal_nuclei" class="mw-redirect" title="Septal nuclei">septal nuclei</a>, <a href="/wiki/Lateral_hypothalamic_area" class="mw-redirect" title="Lateral hypothalamic area">lateral hypothalamic area</a> and the <a href="/wiki/Central_nucleus_of_the_amygdala" title="Central nucleus of the amygdala">central nucleus of the amygdala</a>, among other regions.<sup id="cite_ref-pmid21452224_6-1" class="reference"><a href="#cite_note-pmid21452224-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Dos_Santos_2015_7-0" class="reference"><a href="#cite_note-Dos_Santos_2015-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p>Unlike the classical preganglionic neurons that contain <a href="/wiki/Choline_acetyltransferase" title="Choline acetyltransferase">choline acetyltransferase</a>, neurons of the Edinger–Westphal nucleus contain various <a href="/wiki/Neuropeptide" title="Neuropeptide">neuropeptides</a> such as <a href="/wiki/Urocortin" title="Urocortin">urocortin</a> and <a href="/wiki/Cocaine-_and_amphetamine-regulated_transcript" class="mw-redirect" title="Cocaine- and amphetamine-regulated transcript">cocaine- and amphetamine-regulated transcript</a>.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p>Previously, it has been proposed to rename this group of non-preganglionic, neuropeptide-containing neurons to <i>perioculomotor subgriseal neuronal stream</i> (<i>pIII<sub>SG</sub></i>).<sup id="cite_ref-pmid18186029_9-0" class="reference"><a href="#cite_note-pmid18186029-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> It has also been suggested that the preganglionic oculomotor neurons within the Edinger–Westphal nucleus be referred to as the <i>EWpg</i>, and the neuropeptide-containing neurons be known as the centrally-projecting Edinger Westphal nucleus, or <i>EWcp</i>.<sup id="cite_ref-pmid21452224_6-2" class="reference"><a href="#cite_note-pmid21452224-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Anatomical_relations">Anatomical relations</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=2" title="Edit section: Anatomical relations"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The paired nuclei are <a href="/wiki/Posterior_(anatomy)" class="mw-redirect" title="Posterior (anatomy)">posterior</a> to the main motor nucleus (<a href="/wiki/Oculomotor_nucleus" title="Oculomotor nucleus">oculomotor nucleus</a>) and anterolateral to the <a href="/wiki/Cerebral_aqueduct" title="Cerebral aqueduct">cerebral aqueduct</a> in the <a href="/wiki/Anatomical_terms_of_location#Directional_terms" title="Anatomical terms of location">rostral</a> <a href="/wiki/Midbrain" title="Midbrain">midbrain</a> at the level of the <a href="/wiki/Superior_colliculus" title="Superior colliculus">superior colliculus</a>. </p><p>It is the most rostral of the parasympathetic nuclei in the <a href="/wiki/Brain_stem" class="mw-redirect" title="Brain stem">brain stem</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Function">Function</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=3" title="Edit section: Function"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The Edinger–Westphal nucleus supplies <a href="/wiki/Preganglionic" class="mw-redirect" title="Preganglionic">preganglionic</a> parasympathetic fibers to the <a href="/wiki/Human_eye" title="Human eye">eye</a>, <a href="/wiki/Pupillary_response" title="Pupillary response">constricting the pupil</a>, <a href="/wiki/Accommodation_reflex" title="Accommodation reflex">accommodating</a> the <a href="/wiki/Lens_(anatomy)" class="mw-redirect" title="Lens (anatomy)">lens</a>, and convergence of the eyes.<sup id="cite_ref-KaufmanLevin2011_10-0" class="reference"><a href="#cite_note-KaufmanLevin2011-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Neurophysiology">Neurophysiology</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=4" title="Edit section: Neurophysiology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading4"><h4 id="Pupillary_light_reflex">Pupillary light reflex</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=5" title="Edit section: Pupillary light reflex"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1251242444">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output .ambox-content{border-left:10px solid #f28500}.mw-parser-output .ambox-style{border-left:10px solid #fc3}.mw-parser-output .ambox-move{border-left:10px solid #9932cc}.mw-parser-output .ambox-protection{border-left:10px solid #a2a9b1}.mw-parser-output .ambox .mbox-text{border:none;padding:0.25em 0.5em;width:100%}.mw-parser-output .ambox .mbox-image{border:none;padding:2px 0 2px 0.5em;text-align:center}.mw-parser-output .ambox .mbox-imageright{border:none;padding:2px 0.5em 2px 0;text-align:center}.mw-parser-output .ambox .mbox-empty-cell{border:none;padding:0;width:1px}.mw-parser-output .ambox .mbox-image-div{width:52px}@media(min-width:720px){.mw-parser-output .ambox{margin:0 10%}}@media print{body.ns-0 .mw-parser-output .ambox{display:none!important}}</style><table class="box-Expand_section plainlinks metadata ambox mbox-small-left ambox-content" role="presentation"><tbody><tr><td class="mbox-image"><span typeof="mw:File"><a href="/wiki/File:Wiki_letter_w_cropped.svg" class="mw-file-description"><img alt="[icon]" src="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/20px-Wiki_letter_w_cropped.svg.png" decoding="async" width="20" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/30px-Wiki_letter_w_cropped.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/40px-Wiki_letter_w_cropped.svg.png 2x" data-file-width="44" data-file-height="31" /></a></span></td><td class="mbox-text"><div class="mbox-text-span">This section <b>needs expansion</b>. You can help by <a class="external text" href="https://en.wikipedia.org/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=">adding to it</a>. <span class="date-container"><i>(<span class="date">August 2022</span>)</i></span></div></td></tr></tbody></table> <p>The EWN receives feedback from the <a href="/wiki/Locus_coeruleus" title="Locus coeruleus">locus coeruleus</a> about illumination of the retina in order for the EWN to adjust the size of the pupil in response to varying ambient brightness.<sup id="cite_ref-Heiland_Hogan-2022_3-4" class="reference"><a href="#cite_note-Heiland_Hogan-2022-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p><br /> </p> <div class="mw-heading mw-heading2"><h2 id="Research">Research</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=6" title="Edit section: Research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>It has also been implicated in the <a href="/wiki/Mirror_neuron" title="Mirror neuron">mirroring</a> of pupil size in sad facial expressions. When seeing a sad face, participants' pupils dilated or constricted to mirror the face they saw, which predicted both how sad they perceived the face to be, as well as activity within this region.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Eponym">Eponym</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=7" title="Edit section: Eponym"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The nucleus is named for both <a href="/wiki/Ludwig_Edinger" title="Ludwig Edinger">Ludwig Edinger</a>, from <a href="/wiki/Frankfurt" title="Frankfurt">Frankfurt</a>, who demonstrated it in the <a href="/wiki/Fetus" title="Fetus">fetus</a> in 1885, and for <a href="/wiki/Karl_Friedrich_Otto_Westphal" title="Karl Friedrich Otto Westphal">Karl Friedrich Otto Westphal</a>, from <a href="/wiki/Berlin" title="Berlin">Berlin</a>, who demonstrated it in the <a href="/wiki/Adult" title="Adult">adult</a> in 1887.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Additional_images">Additional images</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=8" title="Edit section: Additional images"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul class="gallery mw-gallery-traditional"> <li class="gallerybox" style="width: 155px"> <div class="thumb" style="width: 150px; height: 150px;"><span typeof="mw:File"><a href="/wiki/File:Gray696.png" class="mw-file-description" title="The cranial nerve nuclei schematically represented; dorsal view. Motor nuclei in red; sensory in blue."><img alt="The cranial nerve nuclei schematically represented; dorsal view. Motor nuclei in red; sensory in blue." src="//upload.wikimedia.org/wikipedia/commons/thumb/3/30/Gray696.png/67px-Gray696.png" decoding="async" width="67" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/30/Gray696.png/101px-Gray696.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/30/Gray696.png/134px-Gray696.png 2x" data-file-width="336" data-file-height="600" /></a></span></div> <div class="gallerytext">The cranial nerve nuclei schematically represented; dorsal view. Motor nuclei in red; sensory in blue.</div> </li> <li class="gallerybox" style="width: 155px"> <div class="thumb" style="width: 150px; height: 150px;"><span typeof="mw:File"><a href="/wiki/File:Gray697.png" class="mw-file-description" title="Nuclei of origin of cranial motor nerves schematically represented; lateral view."><img alt="Nuclei of origin of cranial motor nerves schematically represented; lateral view." src="//upload.wikimedia.org/wikipedia/commons/thumb/8/82/Gray697.png/111px-Gray697.png" decoding="async" width="111" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/82/Gray697.png/167px-Gray697.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/82/Gray697.png/222px-Gray697.png 2x" data-file-width="450" data-file-height="486" /></a></span></div> <div class="gallerytext">Nuclei of origin of cranial motor nerves schematically represented; lateral view.</div> </li> <li class="gallerybox" style="width: 155px"> <div class="thumb" style="width: 150px; height: 150px;"><span typeof="mw:File"><a href="/wiki/File:Gray698.png" class="mw-file-description" title="Primary terminal nuclei of the afferent (sensory) cranial nerves schematically represented; lateral view."><img alt="Primary terminal nuclei of the afferent (sensory) cranial nerves schematically represented; lateral view." src="//upload.wikimedia.org/wikipedia/commons/thumb/6/67/Gray698.png/116px-Gray698.png" decoding="async" width="116" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/67/Gray698.png/174px-Gray698.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/67/Gray698.png/232px-Gray698.png 2x" data-file-width="450" data-file-height="466" /></a></span></div> <div class="gallerytext">Primary terminal nuclei of the afferent (sensory) cranial nerves schematically represented; lateral view. </div> </li> </ul> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=9" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Perlia%27s_nucleus" title="Perlia's nucleus">Perlia's nucleus</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=10" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-Joyce-2022-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-Joyce-2022_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFJoyceLePeterson2022" class="citation cs2">Joyce, Christopher; Le, Patrick H.; Peterson, Diana C. (2022), <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/books/NBK537126/">"Neuroanatomy, Cranial Nerve 3 (Oculomotor)"</a>, <i>StatPearls</i>, Treasure Island (FL): StatPearls Publishing, <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/30725811">30725811</a><span class="reference-accessdate">, retrieved <span class="nowrap">2022-08-08</span></span></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=StatPearls&rft.atitle=Neuroanatomy%2C+Cranial+Nerve+3+%28Oculomotor%29&rft.date=2022&rft_id=info%3Apmid%2F30725811&rft.aulast=Joyce&rft.aufirst=Christopher&rft.au=Le%2C+Patrick+H.&rft.au=Peterson%2C+Diana+C.&rft_id=http%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fbooks%2FNBK537126%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-Sonne-2022-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-Sonne-2022_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Sonne-2022_2-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Sonne-2022_2-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSonneLopez-Ojeda2022" class="citation cs2">Sonne, James; Lopez-Ojeda, Wilfredo (2022), <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/books/NBK470353/">"Neuroanatomy, Cranial Nerve"</a>, <i>StatPearls</i>, Treasure Island (FL): StatPearls Publishing, <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/29261885">29261885</a><span class="reference-accessdate">, retrieved <span class="nowrap">2022-08-08</span></span></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=StatPearls&rft.atitle=Neuroanatomy%2C+Cranial+Nerve&rft.date=2022&rft_id=info%3Apmid%2F29261885&rft.aulast=Sonne&rft.aufirst=James&rft.au=Lopez-Ojeda%2C+Wilfredo&rft_id=http%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fbooks%2FNBK470353%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-Heiland_Hogan-2022-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-Heiland_Hogan-2022_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Heiland_Hogan-2022_3-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Heiland_Hogan-2022_3-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Heiland_Hogan-2022_3-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Heiland_Hogan-2022_3-4"><sup><i><b>e</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHeiland_HoganSubramanianM_Das2022" class="citation cs2">Heiland Hogan, Madeline B.; Subramanian, Suriya; M Das, Joe (2022), <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/books/NBK554555/">"Neuroanatomy, Edinger–Westphal Nucleus (Accessory Oculomotor Nucleus)"</a>, <i>StatPearls</i>, Treasure Island (FL): StatPearls Publishing, <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/32119442">32119442</a><span class="reference-accessdate">, retrieved <span class="nowrap">2022-08-08</span></span></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=StatPearls&rft.atitle=Neuroanatomy%2C+Edinger%E2%80%93Westphal+Nucleus+%28Accessory+Oculomotor+Nucleus%29&rft.date=2022&rft_id=info%3Apmid%2F32119442&rft.aulast=Heiland+Hogan&rft.aufirst=Madeline+B.&rft.au=Subramanian%2C+Suriya&rft.au=M+Das%2C+Joe&rft_id=http%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fbooks%2FNBK554555%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-Yoo-2023-4"><span class="mw-cite-backlink">^ <a href="#cite_ref-Yoo-2023_4-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Yoo-2023_4-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFYooMihaila2023" class="citation cs2">Yoo, Hannah; Mihaila, Dana M. (2023), <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/books/NBK553169/">"Neuroanatomy, Pupillary Light Reflexes and Pathway"</a>, <i>StatPearls</i>, Treasure Island (FL): StatPearls Publishing, <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/31985962">31985962</a><span class="reference-accessdate">, retrieved <span class="nowrap">2023-08-15</span></span></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=StatPearls&rft.atitle=Neuroanatomy%2C+Pupillary+Light+Reflexes+and+Pathway&rft.date=2023&rft_id=info%3Apmid%2F31985962&rft.aulast=Yoo&rft.aufirst=Hannah&rft.au=Mihaila%2C+Dana+M.&rft_id=http%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fbooks%2FNBK553169%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-Haines_2018-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-Haines_2018_5-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHaines2018" class="citation book cs1">Haines, Duane (2018). <i>Fundamental neuroscience for basic and clinical applications</i> (Fifth ed.). Philadelphia, PA: Elsevier. p. 188. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780323396325" title="Special:BookSources/9780323396325"><bdi>9780323396325</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Fundamental+neuroscience+for+basic+and+clinical+applications&rft.place=Philadelphia%2C+PA&rft.pages=188&rft.edition=Fifth&rft.pub=Elsevier&rft.date=2018&rft.isbn=9780323396325&rft.aulast=Haines&rft.aufirst=Duane&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-pmid21452224-6"><span class="mw-cite-backlink">^ <a href="#cite_ref-pmid21452224_6-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-pmid21452224_6-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-pmid21452224_6-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKoziczBittencourtMayReiner2011" class="citation journal cs1">Kozicz T, Bittencourt JC, May PJ, Reiner A, Gamlin PD, Palkovits M, Horn AK, Toledo CA, Ryabinin AE (2011). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3675228">"The Edinger–Westphal nucleus: A historical, structural and functional perspective on a dichotomous terminology"</a>. <i>J. Comp. Neurol</i>. <b>519</b> (8): 1413–34. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1002%2Fcne.22580">10.1002/cne.22580</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3675228">3675228</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21452224">21452224</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=J.+Comp.+Neurol.&rft.atitle=The+Edinger%E2%80%93Westphal+nucleus%3A+A+historical%2C+structural+and+functional+perspective+on+a+dichotomous+terminology&rft.volume=519&rft.issue=8&rft.pages=1413-34&rft.date=2011&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3675228%23id-name%3DPMC&rft_id=info%3Apmid%2F21452224&rft_id=info%3Adoi%2F10.1002%2Fcne.22580&rft.aulast=Kozicz&rft.aufirst=T&rft.au=Bittencourt%2C+JC&rft.au=May%2C+PJ&rft.au=Reiner%2C+A&rft.au=Gamlin%2C+PD&rft.au=Palkovits%2C+M&rft.au=Horn%2C+AK&rft.au=Toledo%2C+CA&rft.au=Ryabinin%2C+AE&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3675228&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-Dos_Santos_2015-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-Dos_Santos_2015_7-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDos_Santos_Junior,_ED,_Da_Silva_AV,_Da_Silva_KR,_Haemmerle_CA,_Batagello_DS,_Da_Silva_JM,_Lima_LB,_Da_Silva_RJ,_Diniz_GB,_Sita_LV,_Elias_CF,_Bittencourt_JC2015" class="citation journal cs1">Dos Santos Junior, ED, Da Silva AV, Da Silva KR, Haemmerle CA, Batagello DS, Da Silva JM, Lima LB, Da Silva RJ, Diniz GB, Sita LV, Elias CF, Bittencourt JC (2015). <a rel="nofollow" class="external text" href="https://repositorio.unesp.br/bitstream/11449/160880/1/WOS000362309000003.pdf">"The centrally projecting Edinger-Westphal nucleus-I: Efferents in the rat brain"</a> <span class="cs1-format">(PDF)</span>. <i>J. Chem. Neuroanat</i>. <b>68</b>: 22–38. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.jchemneu.2015.07.002">10.1016/j.jchemneu.2015.07.002</a>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/11449%2F160880">11449/160880</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/26206178">26206178</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:205769513">205769513</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=J.+Chem.+Neuroanat.&rft.atitle=The+centrally+projecting+Edinger-Westphal+nucleus-I%3A+Efferents+in+the+rat+brain&rft.volume=68&rft.pages=22-38&rft.date=2015&rft_id=info%3Ahdl%2F11449%2F160880&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A205769513%23id-name%3DS2CID&rft_id=info%3Apmid%2F26206178&rft_id=info%3Adoi%2F10.1016%2Fj.jchemneu.2015.07.002&rft.au=Dos+Santos+Junior%2C+ED%2C+Da+Silva+AV%2C+Da+Silva+KR%2C+Haemmerle+CA%2C+Batagello+DS%2C+Da+Silva+JM%2C+Lima+LB%2C+Da+Silva+RJ%2C+Diniz+GB%2C+Sita+LV%2C+Elias+CF%2C+Bittencourt+JC&rft_id=https%3A%2F%2Frepositorio.unesp.br%2Fbitstream%2F11449%2F160880%2F1%2FWOS000362309000003.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_journal" title="Template:Cite journal">cite journal</a>}}</code>: CS1 maint: multiple names: authors list (<a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">link</a>)</span>.</span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKozicz2003" class="citation journal cs1">Kozicz, T. (2003). "Neurons colocalizing urocortin and cocaine and amphetamine-regulated transcript immunoreactivities are induced by acute lipopolysaccharide stress in the Edinger–Westphal nucleus in the rat". <i>Neuroscience</i>. <b>116</b> (2): 315–20. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0306-4522%2802%2900772-8">10.1016/S0306-4522(02)00772-8</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/12559087">12559087</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:11251849">11251849</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Neuroscience&rft.atitle=Neurons+colocalizing+urocortin+and+cocaine+and+amphetamine-regulated+transcript+immunoreactivities+are+induced+by+acute+lipopolysaccharide+stress+in+the+Edinger%E2%80%93Westphal+nucleus+in+the+rat&rft.volume=116&rft.issue=2&rft.pages=315-20&rft.date=2003&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A11251849%23id-name%3DS2CID&rft_id=info%3Apmid%2F12559087&rft_id=info%3Adoi%2F10.1016%2FS0306-4522%2802%2900772-8&rft.aulast=Kozicz&rft.aufirst=T.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-pmid18186029-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-pmid18186029_9-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMayReinerRyabinin2008" class="citation journal cs1">May PJ, Reiner AJ, Ryabinin AE (March 2008). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2863095">"Comparison of the Distributions of Urocortin Containing and Cholinergic Neurons in the Perioculomotor Midbrain of the Cat and Macaque"</a>. <i>J. Comp. Neurol</i>. <b>507</b> (3): 1300–16. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1002%2Fcne.21514">10.1002/cne.21514</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2863095">2863095</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18186029">18186029</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=J.+Comp.+Neurol.&rft.atitle=Comparison+of+the+Distributions+of+Urocortin+Containing+and+Cholinergic+Neurons+in+the+Perioculomotor+Midbrain+of+the+Cat+and+Macaque&rft.volume=507&rft.issue=3&rft.pages=1300-16&rft.date=2008-03&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2863095%23id-name%3DPMC&rft_id=info%3Apmid%2F18186029&rft_id=info%3Adoi%2F10.1002%2Fcne.21514&rft.aulast=May&rft.aufirst=PJ&rft.au=Reiner%2C+AJ&rft.au=Ryabinin%2C+AE&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2863095&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-KaufmanLevin2011-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-KaufmanLevin2011_10-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKaufmanLevinAlm2011" class="citation book cs1">Kaufman, Paul L.; Levin, Leonard A.; Alm, Albert (2011). <a rel="nofollow" class="external text" href="https://books.google.com/books?id=1oIlvnXnJvEC&pg=PA508"><i>Adler's Physiology of the Eye</i></a>. Elsevier Health Sciences. p. 508. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-323-05714-1" title="Special:BookSources/978-0-323-05714-1"><bdi>978-0-323-05714-1</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Adler%27s+Physiology+of+the+Eye&rft.pages=508&rft.pub=Elsevier+Health+Sciences&rft.date=2011&rft.isbn=978-0-323-05714-1&rft.aulast=Kaufman&rft.aufirst=Paul+L.&rft.au=Levin%2C+Leonard+A.&rft.au=Alm%2C+Albert&rft_id=https%3A%2F%2Fbooks.google.com%2Fbooks%3Fid%3D1oIlvnXnJvEC%26pg%3DPA508&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHarrisonWilsonCritchley2007" class="citation journal cs1">Harrison, NA; Wilson, CE; Critchley, HD (2007). "Processing of observed pupil size modulates perception of sadness and predicts empathy". <i>Emotion</i>. <b>7</b> (4): 724–9. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1037%2F1528-3542.7.4.724">10.1037/1528-3542.7.4.724</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18039039">18039039</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Emotion&rft.atitle=Processing+of+observed+pupil+size+modulates+perception+of+sadness+and+predicts+empathy&rft.volume=7&rft.issue=4&rft.pages=724-9&rft.date=2007&rft_id=info%3Adoi%2F10.1037%2F1528-3542.7.4.724&rft_id=info%3Apmid%2F18039039&rft.aulast=Harrison&rft.aufirst=NA&rft.au=Wilson%2C+CE&rft.au=Critchley%2C+HD&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHarrisonSingerRotshteinDolan2006" class="citation journal cs1">Harrison, NA; Singer, T; Rotshtein, P; Dolan, RJ; Critchley, HD (2006). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1716019">"Pupillary contagion: central mechanisms engaged in sadness processing"</a>. <i>Social Cognitive and Affective Neuroscience</i>. <b>1</b> (1): 5–17. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fscan%2Fnsl006">10.1093/scan/nsl006</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1716019">1716019</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/17186063">17186063</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Social+Cognitive+and+Affective+Neuroscience&rft.atitle=Pupillary+contagion%3A+central+mechanisms+engaged+in+sadness+processing&rft.volume=1&rft.issue=1&rft.pages=5-17&rft.date=2006&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1716019%23id-name%3DPMC&rft_id=info%3Apmid%2F17186063&rft_id=info%3Adoi%2F10.1093%2Fscan%2Fnsl006&rft.aulast=Harrison&rft.aufirst=NA&rft.au=Singer%2C+T&rft.au=Rotshtein%2C+P&rft.au=Dolan%2C+RJ&rft.au=Critchley%2C+HD&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1716019&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text"><i><a rel="nofollow" class="external text" href="http://www.whonamedit.com/synd.cfm/893.html">synd/893</a></i> at <a href="/wiki/Who_Named_It%3F" class="mw-redirect" title="Who Named It?">Who Named It?</a></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&action=edit&section=11" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="https://www1.columbia.edu/sec/itc/hs/medical/anatomy_resources/anatomy/lecture6/lecture6_files/Slide0009.gif">Diagram at Columbia</a></li> <li><a rel="nofollow" class="external text" href="http://brainmaps.org/index.php?q=Edinger-Westphal%20nucleus">Stained brain slice images which include the "Edinger–Westphal nucleus"</a> at the <a href="/wiki/BrainMaps" title="BrainMaps">BrainMaps project</a></li> <li><a href="/wiki/Loyola_University_Chicago" title="Loyola University Chicago">MedEd at Loyola</a> <i><a rel="nofollow" class="external text" href="http://www.meddean.luc.edu/Lumen/MedEd/Neuro/frames/nlBSs/nl18fr.htm">Neuro/frames/nlBSs/nl18fr.htm</a></i></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20160303222022/http://www.yale.edu/cnerves/cn3/cn3_12.html">"3-12"</a>. <i>Cranial Nerves</i>. <a href="/wiki/Yale_School_of_Medicine" title="Yale School of Medicine">Yale School of Medicine</a>. Archived from <a rel="nofollow" class="external text" href="http://www.yale.edu/cnerves/cn3/cn3_12.html">the original</a> on 2016-03-03.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Cranial+Nerves&rft.atitle=3-12&rft_id=http%3A%2F%2Fwww.yale.edu%2Fcnerves%2Fcn3%2Fcn3_12.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AEdinger%E2%80%93Westphal+nucleus" class="Z3988"></span></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist dt::after{content:": "}.mw-parser-output .hlist dd::after,.mw-parser-output .hlist li::after{content:" · ";font-weight:bold}.mw-parser-output .hlist dd:last-child::after,.mw-parser-output .hlist dt:last-child::after,.mw-parser-output .hlist li:last-child::after{content:none}.mw-parser-output .hlist dd dd:first-child::before,.mw-parser-output .hlist dd dt:first-child::before,.mw-parser-output .hlist dd li:first-child::before,.mw-parser-output .hlist dt dd:first-child::before,.mw-parser-output .hlist dt dt:first-child::before,.mw-parser-output .hlist dt li:first-child::before,.mw-parser-output .hlist li dd:first-child::before,.mw-parser-output .hlist li dt:first-child::before,.mw-parser-output .hlist li li:first-child::before{content:" (";font-weight:normal}.mw-parser-output .hlist dd dd:last-child::after,.mw-parser-output .hlist dd dt:last-child::after,.mw-parser-output .hlist dd li:last-child::after,.mw-parser-output .hlist dt dd:last-child::after,.mw-parser-output .hlist dt dt:last-child::after,.mw-parser-output .hlist dt li:last-child::after,.mw-parser-output .hlist li dd:last-child::after,.mw-parser-output .hlist li dt:last-child::after,.mw-parser-output .hlist li li:last-child::after{content:")";font-weight:normal}.mw-parser-output .hlist ol{counter-reset:listitem}.mw-parser-output .hlist ol>li{counter-increment:listitem}.mw-parser-output .hlist ol>li::before{content:" "counter(listitem)"\a0 "}.mw-parser-output .hlist dd ol>li:first-child::before,.mw-parser-output .hlist dt ol>li:first-child::before,.mw-parser-output .hlist li ol>li:first-child::before{content:" ("counter(listitem)"\a0 "}</style><style data-mw-deduplicate="TemplateStyles:r1236075235">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output .navbox-subgroup{width:100%}.mw-parser-output .navbox-group,.mw-parser-output .navbox-title,.mw-parser-output .navbox-abovebelow{padding:0.25em 1em;line-height:1.5em;text-align:center}.mw-parser-output .navbox-group{white-space:nowrap;text-align:right}.mw-parser-output .navbox,.mw-parser-output .navbox-subgroup{background-color:#fdfdfd}.mw-parser-output .navbox-list{line-height:1.5em;border-color:#fdfdfd}.mw-parser-output .navbox-list-with-group{text-align:left;border-left-width:2px;border-left-style:solid}.mw-parser-output tr+tr>.navbox-abovebelow,.mw-parser-output tr+tr>.navbox-group,.mw-parser-output tr+tr>.navbox-image,.mw-parser-output tr+tr>.navbox-list{border-top:2px solid #fdfdfd}.mw-parser-output .navbox-title{background-color:#ccf}.mw-parser-output .navbox-abovebelow,.mw-parser-output .navbox-group,.mw-parser-output .navbox-subgroup .navbox-title{background-color:#ddf}.mw-parser-output .navbox-subgroup .navbox-group,.mw-parser-output .navbox-subgroup .navbox-abovebelow{background-color:#e6e6ff}.mw-parser-output .navbox-even{background-color:#f7f7f7}.mw-parser-output .navbox-odd{background-color:transparent}.mw-parser-output .navbox .hlist td dl,.mw-parser-output .navbox .hlist td ol,.mw-parser-output .navbox .hlist td ul,.mw-parser-output .navbox td.hlist dl,.mw-parser-output .navbox td.hlist ol,.mw-parser-output .navbox td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Anatomy_of_the_midbrain" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Mesencephalon" title="Template:Mesencephalon"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Mesencephalon" title="Template talk:Mesencephalon"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Mesencephalon" title="Special:EditPage/Template:Mesencephalon"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Anatomy_of_the_midbrain" style="font-size:114%;margin:0 4em">Anatomy of the <a href="/wiki/Midbrain" title="Midbrain">midbrain</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Midbrain_tectum" class="mw-redirect" title="Midbrain tectum">Tectum</a><br />(Dorsal)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Corpora_quadrigemina" title="Corpora quadrigemina">Corpora quadrigemina</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Inferior_colliculus" title="Inferior colliculus">Inferior colliculus</a> <ul><li><a href="/wiki/Inferior_colliculus#Input_and_output_connections_of_IC" title="Inferior colliculus">Inferior brachium</a></li></ul></li> <li><a href="/wiki/Superior_colliculus" title="Superior colliculus">Superior colliculus</a> <ul><li><a href="/wiki/Superior_colliculus#Structure" title="Superior colliculus">Superior brachium</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grey_matter" title="Grey matter">Grey matter</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pretectal_area" title="Pretectal area">Pretectal area</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/White_matter" title="White matter">White matter</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Sensory / ascending</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Spinotectal_tract" class="mw-redirect" title="Spinotectal tract">Spinotectal tract</a></li> <li><a href="/wiki/Central_tegmental_tract" title="Central tegmental tract">Central tegmental tract</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Motor / descending</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Tectospinal_tract" title="Tectospinal tract">Tectospinal tract</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ventricular_system" title="Ventricular system">CSF</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cerebral_aqueduct" title="Cerebral aqueduct">Cerebral aqueduct</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cerebral_peduncle" title="Cerebral peduncle">Peduncle</a><br />(Ventral)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Midbrain_tegmentum" title="Midbrain tegmentum">Tegmentum</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/White_matter" title="White matter">White matter</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Sensory / ascending</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Lemnisci <ul><li><a href="/wiki/Medial_lemniscus" title="Medial lemniscus">Medial</a></li> <li><a href="/wiki/Lateral_lemniscus" title="Lateral lemniscus">Lateral</a></li></ul></li> <li><a href="/wiki/Medial_longitudinal_fasciculus" title="Medial longitudinal fasciculus">Ascending MLF</a> <ul><li><a href="/wiki/Vestibulo-oculomotor_fibers" title="Vestibulo-oculomotor fibers">Vestibulo-oculomotor fibers</a></li></ul></li> <li><a href="/wiki/Spinothalamic_tract" title="Spinothalamic tract">Spinothalamic tract</a></li> <li><a href="/wiki/Ventral_trigeminal_tract" title="Ventral trigeminal tract">Ventral trigeminal tract</a></li> <li><a href="/wiki/Dentatothalamic_tract" title="Dentatothalamic tract">Dentatothalamic tract</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Motor / descending</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rubrospinal_tract" title="Rubrospinal tract">Rubrospinal tract</a></li> <li><a href="/wiki/Rubro-olivary_tract" title="Rubro-olivary tract">Rubro-olivary tract</a></li> <li><a href="/wiki/Medial_longitudinal_fasciculus" title="Medial longitudinal fasciculus">Descending MLF</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grey_matter" title="Grey matter">Grey matter</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cranial_nerve_nucleus" title="Cranial nerve nucleus">cranial nuclei</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/General_somatic_afferent_fibers" class="mw-redirect" title="General somatic afferent fibers">GSA</a> <ul><li><a href="/wiki/Mesencephalic_nucleus_of_trigeminal_nerve" title="Mesencephalic nucleus of trigeminal nerve">Mesencephalic</a></li></ul></li> <li><a href="/wiki/General_somatic_efferent_fibers" class="mw-redirect" title="General somatic efferent fibers">GSE</a> <ul><li><a href="/wiki/Oculomotor_nucleus" title="Oculomotor nucleus">Oculomotor nucleus</a></li> <li><a href="/wiki/Trochlear_nucleus" title="Trochlear nucleus">Trochlear nucleus</a></li></ul></li> <li><a href="/wiki/General_visceral_efferent_fibers" class="mw-redirect" title="General visceral efferent fibers">GVE</a> <ul><li><a class="mw-selflink selflink">Edinger–Westphal nucleus</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ventral_tegmental_area" title="Ventral tegmental area">Ventral tegmental area</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rostromedial_tegmental_nucleus" title="Rostromedial tegmental nucleus">Rostromedial tegmental nucleus</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Periaqueductal_gray" title="Periaqueductal gray">Periaqueductal gray</a></li> <li><a href="/wiki/Red_nucleus" title="Red nucleus">Red nucleus</a></li> <li><a href="/wiki/Rostral_interstitial_nucleus_of_medial_longitudinal_fasciculus" title="Rostral interstitial nucleus of medial longitudinal fasciculus">Rostral interstitial nucleus of medial longitudinal fasciculus</a></li> <li><a href="/wiki/Parabrachial_area" class="mw-redirect" title="Parabrachial area">Parabrachial area</a></li> <li><a href="/wiki/Interpeduncular_nucleus" title="Interpeduncular nucleus">Interpeduncular nucleus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Midbrain_reticular_formation" title="Midbrain reticular formation">Midbrain reticular formation</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dorsal_tegmental_nucleus" title="Dorsal tegmental nucleus">Dorsal tegmental nucleus</a></li> <li><a href="/wiki/Raphe_nuclei" title="Raphe nuclei">Raphe nuclei</a> <ul><li><a href="/wiki/Dorsal_raphe_nucleus" title="Dorsal raphe nucleus">dorsal</a></li></ul></li> <li><a href="/wiki/Reticulotegmental_nucleus" title="Reticulotegmental nucleus">Reticulotegmental nucleus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Base</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/White_matter" title="White matter">White</a> / <a href="/wiki/Cerebral_crus" title="Cerebral crus">Cerebral crus</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pyramidal_tracts" title="Pyramidal tracts">Pyramidal tracts</a> <ul><li><a href="/wiki/Corticospinal_tract" title="Corticospinal tract">Corticospinal tract</a></li> <li><a href="/wiki/Corticobulbar_tract" title="Corticobulbar tract">Corticobulbar tract</a></li></ul></li> <li><a href="/wiki/Corticopontine_fibers" title="Corticopontine fibers">Corticopontine tract</a> <ul><li><a href="/wiki/Frontopontine_fibers" title="Frontopontine fibers">Frontopontine fibers</a></li> <li><a href="/wiki/Temporopontine_fibers" title="Temporopontine fibers">Temporopontine fibers</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grey_matter" title="Grey matter">Grey</a> / <a href="/wiki/Substantia_nigra" title="Substantia nigra">Substantia nigra</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pars_compacta" title="Pars compacta">Pars compacta</a></li> <li><a href="/wiki/Pars_reticulata" title="Pars reticulata">Pars reticulata</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Surface</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Superior_cerebellar_peduncle" title="Superior cerebellar peduncle">Superior cerebellar peduncle</a> <ul><li><a href="/wiki/Decussation_of_superior_cerebellar_peduncles" class="mw-redirect" title="Decussation of superior cerebellar peduncles">Decussation</a></li></ul></li> <li><a href="/wiki/Interpeduncular_fossa" title="Interpeduncular fossa">Interpeduncular fossa</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="The_cranial_nerves" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Cranial_nerves" title="Template:Cranial nerves"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Cranial_nerves" title="Template talk:Cranial nerves"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Cranial_nerves" title="Special:EditPage/Template:Cranial nerves"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="The_cranial_nerves" style="font-size:114%;margin:0 4em">The <a href="/wiki/Cranial_nerves" title="Cranial nerves">cranial nerves</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Terminal_nerve" title="Terminal nerve">Terminal</a> (CN 0)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Nuclei <ul><li><a href="/wiki/Septal_nuclei" class="mw-redirect" title="Septal nuclei">septal nuclei</a></li></ul></li> <li>Course <ul><li><i>no significant branches</i></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Olfactory_nerve" title="Olfactory nerve">Olfactory</a> (CN I)</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Nuclei <ul><li><a href="/wiki/Anterior_olfactory_nucleus" title="Anterior olfactory nucleus">anterior olfactory nucleus</a></li></ul></li> <li>Course <ul><li><a href="/wiki/Olfactory_bulb" title="Olfactory bulb">olfactory bulb</a></li> <li><a href="/wiki/Olfactory_tract" title="Olfactory tract">olfactory tract</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Optic_nerve" title="Optic nerve">Optic</a> (CN II)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Nuclei <ul><li><a href="/wiki/Lateral_geniculate_nucleus" title="Lateral geniculate nucleus">lateral geniculate nucleus</a></li></ul></li> <li>Course <ul><li><a href="/wiki/Optic_chiasm" title="Optic chiasm">optic chiasm</a></li> <li><a href="/wiki/Optic_tract" title="Optic tract">optic tract</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Oculomotor_nerve" title="Oculomotor nerve">Oculomotor</a> (CN III)</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Nuclei <ul><li><a href="/wiki/Oculomotor_nucleus" title="Oculomotor nucleus">oculomotor nucleus</a></li> <li><a class="mw-selflink selflink">Edinger–Westphal nucleus</a></li></ul></li> <li>Branches <ul><li><a href="/wiki/Superior_branch_of_oculomotor_nerve" class="mw-redirect" title="Superior branch of oculomotor nerve">superior</a></li> <li><a href="/wiki/Parasympathetic_root_of_ciliary_ganglion" class="mw-redirect" title="Parasympathetic root of ciliary ganglion">parasympathetic root</a> of <a href="/wiki/Ciliary_ganglion" title="Ciliary ganglion">ciliary ganglion</a></li> <li><a href="/wiki/Inferior_branch_of_oculomotor_nerve" class="mw-redirect" title="Inferior branch of oculomotor nerve">inferior</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Trochlear_nerve" title="Trochlear nerve">Trochlear</a> (CN IV)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Trochlear_nucleus" title="Trochlear nucleus">Nucleus</a></li> <li>Branches <ul><li><i>no significant branches</i></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Trigeminal_nerve" title="Trigeminal nerve">Trigeminal</a> (CN V)</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Nuclei <ul><li><a href="/wiki/Principal_sensory_nucleus_of_trigeminal_nerve" title="Principal sensory nucleus of trigeminal nerve">PSN</a></li> <li><a href="/wiki/Spinal_trigeminal_nucleus" title="Spinal trigeminal nucleus">spinal trigeminal nucleus</a></li> <li><a href="/wiki/Mesencephalic_nucleus_of_trigeminal_nerve" title="Mesencephalic nucleus of trigeminal nerve">MN</a></li> <li><a href="/wiki/Trigeminal_motor_nucleus" title="Trigeminal motor nucleus">TMN</a></li></ul></li> <li>Course <ul><li><a href="/wiki/Trigeminal_ganglion" title="Trigeminal ganglion">trigeminal ganglion</a></li></ul></li> <li>Branches <ul><li><a href="/wiki/Ophthalmic_nerve" title="Ophthalmic nerve">ophthalmic</a></li> <li><a href="/wiki/Maxillary_nerve" title="Maxillary nerve">maxillary</a></li> <li><a href="/wiki/Mandibular_nerve" title="Mandibular nerve">mandibular</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Abducens_nerve" title="Abducens nerve">Abducens</a> (CN VI)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abducens_nucleus" title="Abducens nucleus">Nucleus</a></li> <li>Branches <ul><li><i>no significant branches</i></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Facial_nerve" title="Facial nerve">Facial</a> (CN VII)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Near origin</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Intermediate_nerve" title="Intermediate nerve">Intermediate nerve</a></li> <li><a href="/wiki/Geniculate_ganglion" title="Geniculate ganglion">Geniculate</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Inside<br /><a href="/wiki/Facial_canal" title="Facial canal">facial canal</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Greater_petrosal_nerve" title="Greater petrosal nerve">Greater petrosal</a> <ul><li><a href="/wiki/Pterygopalatine_ganglion" title="Pterygopalatine ganglion">pterygopalatine ganglion</a></li></ul></li> <li><a href="/wiki/Nerve_to_the_stapedius" title="Nerve to the stapedius">Nerve to the stapedius</a></li> <li><a href="/wiki/Chorda_tympani" title="Chorda tympani">Chorda tympani</a> <ul><li><a href="/wiki/Lingual_nerve" title="Lingual nerve">lingual nerve</a></li> <li><a href="/wiki/Submandibular_ganglion" title="Submandibular ganglion">submandibular ganglion</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">At <a href="/wiki/Stylomastoid_foramen" title="Stylomastoid foramen">stylomastoid<br />foramen</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Posterior_auricular_nerve" title="Posterior auricular nerve">Posterior auricular</a></li> <li><a href="/wiki/Suprahyoid_muscles" title="Suprahyoid muscles">Suprahyoid</a> <ul><li><a href="/wiki/Digastric_branch_of_facial_nerve" title="Digastric branch of facial nerve">digastric</a></li> <li><a href="/wiki/Stylohyoid_branch_of_facial_nerve" title="Stylohyoid branch of facial nerve">stylohyoid</a></li></ul></li> <li><a href="/wiki/Parotid_plexus" title="Parotid plexus">Parotid plexus</a> <ul><li><a href="/wiki/Temporal_branches_of_the_facial_nerve" title="Temporal branches of the facial nerve">temporal</a></li> <li><a href="/wiki/Zygomatic_branches_of_the_facial_nerve" title="Zygomatic branches of the facial nerve">zygomatic</a></li> <li><a href="/wiki/Buccal_branch_of_the_facial_nerve" class="mw-redirect" title="Buccal branch of the facial nerve">buccal</a></li> <li><a href="/wiki/Marginal_mandibular_branch_of_facial_nerve" class="mw-redirect" title="Marginal mandibular branch of facial nerve">mandibular</a></li> <li><a href="/wiki/Cervical_branch_of_the_facial_nerve" title="Cervical branch of the facial nerve">cervical</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Nuclei</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Facial_motor_nucleus" title="Facial motor nucleus">Facial motor nucleus</a></li> <li><a href="/wiki/Solitary_nucleus" title="Solitary nucleus">Solitary nucleus</a></li> <li><a href="/wiki/Superior_salivary_nucleus" class="mw-redirect" title="Superior salivary nucleus">Superior salivary nucleus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Vestibulocochlear_nerve" title="Vestibulocochlear nerve">Vestibulocochlear</a> (CN VIII)</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Nuclei <ul><li><a href="/wiki/Vestibular_nuclei" title="Vestibular nuclei">vestibular nuclei</a></li> <li><a href="/wiki/Cochlear_nuclei" class="mw-redirect" title="Cochlear nuclei">cochlear nuclei</a></li></ul></li> <li><a href="/wiki/Cochlear_nerve" title="Cochlear nerve">Cochlear nerve</a> <ul><li><a href="/wiki/Striae_medullares" class="mw-redirect" title="Striae medullares">striae medullares</a></li> <li><a href="/wiki/Lateral_lemniscus" title="Lateral lemniscus">lateral lemniscus</a></li></ul></li> <li><a href="/wiki/Vestibular_nerve" title="Vestibular nerve">Vestibular</a> <ul><li><a href="/wiki/Scarpa%27s_ganglion" class="mw-redirect" title="Scarpa's ganglion">Scarpa's ganglion</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Glossopharyngeal_nerve" title="Glossopharyngeal nerve">Glossopharyngeal</a> (CN IX)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Before <a href="/wiki/Jugular_fossa" title="Jugular fossa">jugular fossa</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Ganglia <ul><li><a href="/wiki/Superior_ganglion_of_glossopharyngeal_nerve" title="Superior ganglion of glossopharyngeal nerve">superior</a></li> <li><a href="/wiki/Inferior_ganglion_of_glossopharyngeal_nerve" title="Inferior ganglion of glossopharyngeal nerve">inferior</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">After <a href="/wiki/Jugular_fossa" title="Jugular fossa">jugular fossa</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Tympanic_nerve" title="Tympanic nerve">Tympanic</a> <ul><li><a href="/wiki/Tympanic_plexus" title="Tympanic plexus">tympanic plexus</a></li> <li><a href="/wiki/Lesser_petrosal_nerve" title="Lesser petrosal nerve">lesser petrosal</a></li> <li><a href="/wiki/Otic_ganglion" title="Otic ganglion">otic ganglion</a></li></ul></li> <li><a href="/wiki/Stylopharyngeal_branch_of_glossopharyngeal_nerve" title="Stylopharyngeal branch of glossopharyngeal nerve">Stylopharyngeal branch</a></li> <li><a href="/wiki/Pharyngeal_branches_of_glossopharyngeal_nerve" title="Pharyngeal branches of glossopharyngeal nerve">Pharyngeal branches</a></li> <li><a href="/wiki/Tonsillar_branches_of_glossopharyngeal_nerve" title="Tonsillar branches of glossopharyngeal nerve">Tonsillar branches</a></li> <li><a href="/wiki/Lingual_branches_of_glossopharyngeal_nerve" title="Lingual branches of glossopharyngeal nerve">Lingual branches</a></li> <li><a href="/wiki/Branch_of_glossopharyngeal_nerve_to_carotid_sinus" class="mw-redirect" title="Branch of glossopharyngeal nerve to carotid sinus">Carotid sinus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Nuclei</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nucleus_ambiguus" title="Nucleus ambiguus">Nucleus ambiguus</a></li> <li><a href="/wiki/Inferior_salivatory_nucleus" class="mw-redirect" title="Inferior salivatory nucleus">Inferior salivatory nucleus</a></li> <li><a href="/wiki/Solitary_nucleus" title="Solitary nucleus">Solitary nucleus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Vagus_nerve" title="Vagus nerve">Vagus</a> (CN X)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Before <a href="/wiki/Jugular_fossa" title="Jugular fossa">jugular fossa</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Ganglia <ul><li><a href="/wiki/Superior_ganglion_of_vagus_nerve" title="Superior ganglion of vagus nerve">superior</a></li> <li><a href="/wiki/Inferior_ganglion_of_vagus_nerve" title="Inferior ganglion of vagus nerve">inferior</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">After <a href="/wiki/Jugular_fossa" title="Jugular fossa">jugular fossa</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Meningeal_branch_of_vagus_nerve" title="Meningeal branch of vagus nerve">Meningeal branch</a></li> <li><a href="/wiki/Auricular_branch_of_vagus_nerve" title="Auricular branch of vagus nerve">Auricular branch</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Neck" title="Neck">Neck</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pharyngeal_branch_of_vagus_nerve" title="Pharyngeal branch of vagus nerve">Pharyngeal branch</a> <ul><li><a href="/wiki/Pharyngeal_plexus_of_vagus_nerve" title="Pharyngeal plexus of vagus nerve">pharyngeal plexus</a></li></ul></li> <li><a href="/wiki/Superior_laryngeal_nerve" title="Superior laryngeal nerve">Superior laryngeal</a> <ul><li><a href="/wiki/External_laryngeal_nerve" class="mw-redirect" title="External laryngeal nerve">external</a></li> <li><a href="/wiki/Internal_laryngeal_nerve" class="mw-redirect" title="Internal laryngeal nerve">internal</a></li></ul></li> <li><a href="/wiki/Recurrent_laryngeal_nerve" title="Recurrent laryngeal nerve">Recurrent laryngeal</a></li> <li><a href="/wiki/Superior_cervical_cardiac_branches_of_vagus_nerve" class="mw-redirect" title="Superior cervical cardiac branches of vagus nerve">Superior cervical cardiac</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Thorax" title="Thorax">Thorax</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Inferior_cardiac_branches_of_the_vagus_nerve" class="mw-redirect" title="Inferior cardiac branches of the vagus nerve">Inferior cardiac</a></li> <li><a href="/wiki/Pulmonary_branches_of_vagus_nerve" title="Pulmonary branches of vagus nerve">Pulmonary</a></li> <li>Vagal trunks <ul><li><a href="/wiki/Anterior_vagal_trunk" title="Anterior vagal trunk">anterior</a></li> <li><a href="/wiki/Posterior_vagal_trunk" title="Posterior vagal trunk">posterior</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Abdomen" title="Abdomen">Abdomen</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Celiac_branches_of_vagus_nerve" title="Celiac branches of vagus nerve">Celiac</a></li> <li><a href="/wiki/Renal_branches_of_vagus_nerve" title="Renal branches of vagus nerve">Renal</a></li> <li><a href="/wiki/Hepatic_branches_of_anterior_vagal_trunk" title="Hepatic branches of anterior vagal trunk">Hepatic</a></li> <li><a href="/wiki/Anterior_gastric_branches_of_anterior_vagal_trunk" title="Anterior gastric branches of anterior vagal trunk">Anterior gastric</a></li> <li><a href="/wiki/Posterior_gastric_branches_of_posterior_vagal_trunk" title="Posterior gastric branches of posterior vagal trunk">Posterior gastric</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Nuclei</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nucleus_ambiguus" title="Nucleus ambiguus">Nucleus ambiguus</a></li> <li><a href="/wiki/Dorsal_nucleus_of_vagus_nerve" title="Dorsal nucleus of vagus nerve">Dorsal nucleus of vagus nerve</a></li> <li><a href="/wiki/Solitary_nucleus" title="Solitary nucleus">Solitary nucleus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Accessory_nerve" title="Accessory nerve">Accessory</a> (CN XI)</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Nuclei <ul><li><a href="/wiki/Nucleus_ambiguus" title="Nucleus ambiguus">nucleus ambiguus</a></li> <li><a href="/wiki/Spinal_accessory_nucleus" title="Spinal accessory nucleus">spinal accessory nucleus</a></li></ul></li> <li><a href="/wiki/Cranial_root_of_accessory_nerves" title="Cranial root of accessory nerves">Cranial</a></li> <li><a href="/wiki/Spinal_root_of_accessory_nerve" title="Spinal root of accessory nerve">Spinal</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Hypoglossal_nerve" title="Hypoglossal nerve">Hypoglossal</a> (CN XII)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hypoglossal_nucleus" title="Hypoglossal nucleus">Nucleus</a></li> <li>Branches <ul><li><a href="/wiki/Lingual_branches_of_hypoglossal_nerve" class="mw-redirect" title="Lingual branches of hypoglossal nerve">lingual</a></li></ul></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Optical_illusions_(list)" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Optical_illusions" title="Template:Optical illusions"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Optical_illusions" title="Template talk:Optical illusions"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Optical_illusions" title="Special:EditPage/Template:Optical illusions"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Optical_illusions_(list)" style="font-size:114%;margin:0 4em"><a href="/wiki/Optical_illusion" title="Optical illusion">Optical illusions</a> (<a href="/wiki/List_of_optical_illusions" title="List of optical illusions">list</a>)</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Illusions</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Afterimage" title="Afterimage">Afterimage</a></li> <li><a href="/wiki/Ambigram" title="Ambigram">Ambigram</a></li> <li><a href="/wiki/Ambiguous_image" title="Ambiguous image">Ambiguous image</a></li> <li><a href="/wiki/Ames_room" title="Ames room">Ames room</a></li> <li><a href="/wiki/Autostereogram" title="Autostereogram">Autostereogram</a></li> <li><a href="/wiki/Barberpole_illusion" title="Barberpole illusion">Barberpole</a></li> <li><a href="/wiki/Bezold_effect" title="Bezold effect">Bezold</a></li> <li><a href="/wiki/Caf%C3%A9_wall_illusion" title="Café wall illusion">Café wall</a></li> <li><a href="/wiki/Checker_shadow_illusion" title="Checker shadow illusion">Checker shadow</a></li> <li><a href="/wiki/Chubb_illusion" title="Chubb illusion">Chubb</a></li> <li><a href="/wiki/Cornsweet_illusion" title="Cornsweet illusion">Cornsweet</a></li> <li><a href="/wiki/Delboeuf_illusion" title="Delboeuf illusion">Delboeuf</a></li> <li><a href="/wiki/Ebbinghaus_illusion" title="Ebbinghaus illusion">Ebbinghaus</a></li> <li><a href="/wiki/Ehrenstein_illusion" title="Ehrenstein illusion">Ehrenstein</a></li> <li><a href="/wiki/Flash_lag_illusion" title="Flash lag illusion">Flash lag</a></li> <li><a href="/wiki/Fraser_spiral_illusion" title="Fraser spiral illusion">Fraser spiral</a></li> <li><a href="/wiki/Gravity_hill" title="Gravity hill">Gravity hill</a></li> <li><a href="/wiki/Grid_illusion" title="Grid illusion">Grid</a></li> <li><a href="/wiki/Hering_illusion" title="Hering illusion">Hering</a></li> <li><a href="/wiki/Impossible_trident" title="Impossible trident">Impossible trident</a></li> <li><a href="/wiki/Irradiation_illusion" title="Irradiation illusion">Irradiation</a></li> <li><a href="/wiki/Jastrow_illusion" title="Jastrow illusion">Jastrow</a></li> <li><a href="/wiki/Lilac_chaser" title="Lilac chaser">Lilac chaser</a></li> <li><a href="/wiki/Mach_bands" title="Mach bands">Mach bands</a></li> <li><a href="/wiki/McCollough_effect" title="McCollough effect">McCollough</a></li> <li><a href="/wiki/M%C3%BCller-Lyer_illusion" title="Müller-Lyer illusion">Müller-Lyer</a></li> <li><a href="/wiki/Necker_cube" title="Necker cube">Necker cube</a></li> <li><a href="/wiki/Oppel-Kundt_illusion" title="Oppel-Kundt illusion">Oppel-Kundt</a></li> <li><a href="/wiki/Orbison_illusion" title="Orbison illusion">Orbison</a></li> <li><a href="/wiki/Penrose_stairs" title="Penrose stairs">Penrose stairs</a></li> <li><a href="/wiki/Penrose_triangle" title="Penrose triangle">Penrose triangle</a></li> <li><a href="/wiki/Peripheral_drift_illusion" title="Peripheral drift illusion">Peripheral drift</a></li> <li><a href="/wiki/Poggendorff_illusion" title="Poggendorff illusion">Poggendorff</a></li> <li><a href="/wiki/Ponzo_illusion" title="Ponzo illusion">Ponzo</a></li> <li><a href="/wiki/Rubin_vase" title="Rubin vase">Rubin vase</a></li> <li><a href="/wiki/Sander_illusion" title="Sander illusion">Sander</a></li> <li><a href="/wiki/Schroeder_stairs" title="Schroeder stairs">Schroeder stairs</a></li> <li><a href="/wiki/Shepard_tables" title="Shepard tables">Shepard tables</a></li> <li><a href="/wiki/Spinning_dancer" title="Spinning dancer">Spinning dancer</a></li> <li><a href="/wiki/Ternus_illusion" title="Ternus illusion">Ternus</a></li> <li><a href="/wiki/Vertical%E2%80%93horizontal_illusion" title="Vertical–horizontal illusion">Vertical–horizontal</a></li> <li><a href="/wiki/White%27s_illusion" title="White's illusion">White's</a></li> <li><a href="/wiki/Wundt_illusion" title="Wundt illusion">Wundt</a></li> <li><a href="/wiki/Z%C3%B6llner_illusion" title="Zöllner illusion">Zöllner</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="3" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Optical_Illustion-Ambiguous_Patterns.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/51/Optical_Illustion-Ambiguous_Patterns.svg/30px-Optical_Illustion-Ambiguous_Patterns.svg.png" decoding="async" width="30" height="112" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/51/Optical_Illustion-Ambiguous_Patterns.svg/45px-Optical_Illustion-Ambiguous_Patterns.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/51/Optical_Illustion-Ambiguous_Patterns.svg/60px-Optical_Illustion-Ambiguous_Patterns.svg.png 2x" data-file-width="117" data-file-height="438" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Popular culture</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Op_art" title="Op art">Op art</a></li> <li><i><a href="/wiki/Trompe-l%27%C5%93il" title="Trompe-l'œil">Trompe-l'œil</a></i></li> <li><i><a href="/wiki/Spectropia" title="Spectropia">Spectropia</a></i> (1864 book)</li> <li><i><a href="/wiki/Ascending_and_Descending" title="Ascending and Descending">Ascending and Descending</a></i> (1960 drawing)</li> <li><i><a href="/wiki/Waterfall_(M._C._Escher)" title="Waterfall (M. C. Escher)">Waterfall</a></i> (1961 drawing)</li> <li><i><a href="/wiki/The_dress" title="The dress">The dress</a></i> (2015 photograph)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Related</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Accidental_viewpoint" title="Accidental viewpoint">Accidental viewpoint</a></li> <li><a href="/wiki/Auditory_illusion" title="Auditory illusion">Auditory illusions</a></li> <li><a href="/wiki/Illusion" title="Illusion">Illusions</a></li> <li><a href="/wiki/Tactile_illusion" title="Tactile illusion">Tactile illusions</a></li> <li><a href="/wiki/Time_perception" title="Time perception">Temporal illusion</a></li></ul> </div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐api‐ext.codfw.main‐7556f8b5dd‐mh5wj Cached time: 20241122150901 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 0.696 seconds Real time usage: 0.876 seconds Preprocessor visited node count: 2376/1000000 Post‐expand include size: 152617/2097152 bytes Template argument size: 2586/2097152 bytes Highest expansion depth: 10/100 Expensive parser function count: 4/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 83171/5000000 bytes Lua time usage: 0.433/10.000 seconds Lua memory usage: 6858062/52428800 bytes Number of Wikibase entities loaded: 1/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 752.677 1 -total 20.02% 150.658 13 Template:Navbox 18.83% 141.703 1 Template:Infobox_brain 18.23% 137.201 1 Template:Infobox_anatomy 17.26% 129.914 1 Template:Infobox 17.12% 128.864 4 Template:Citation 14.50% 109.168 1 Template:Mesencephalon 12.23% 92.075 1 Template:Short_description 8.95% 67.357 1 Template:Expand_section 8.06% 60.697 1 Template:Ambox --> <!-- Saved in parser cache with key enwiki:pcache:idhash:2062028-0!canonical and timestamp 20241122150901 and revision id 1251473278. Rendering was triggered because: api-parse --> </div><!--esi <esi:include src="/esitest-fa8a495983347898/content" /> --><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Edinger–Westphal_nucleus&oldid=1251473278">https://en.wikipedia.org/w/index.php?title=Edinger–Westphal_nucleus&oldid=1251473278</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Categories</a>: <ul><li><a href="/wiki/Category:Cranial_nerve_nuclei" title="Category:Cranial nerve nuclei">Cranial nerve nuclei</a></li><li><a href="/wiki/Category:Oculomotor_nerve" title="Category:Oculomotor nerve">Oculomotor nerve</a></li><li><a href="/wiki/Category:Midbrain" title="Category:Midbrain">Midbrain</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks mw-hidden-cats-hidden">Hidden categories: <ul><li><a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">CS1 maint: multiple names: authors list</a></li><li><a href="/wiki/Category:Articles_with_short_description" title="Category:Articles with short description">Articles with short description</a></li><li><a href="/wiki/Category:Short_description_matches_Wikidata" title="Category:Short description matches Wikidata">Short description matches Wikidata</a></li><li><a href="/wiki/Category:Articles_to_be_expanded_from_August_2022" title="Category:Articles to be expanded from August 2022">Articles to be expanded from August 2022</a></li><li><a href="/wiki/Category:All_articles_to_be_expanded" title="Category:All articles to be expanded">All articles to be expanded</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> This page was last edited on 16 October 2024, at 09:32<span class="anonymous-show"> (UTC)</span>.</li> <li id="footer-info-copyright">Text is available under the <a href="/wiki/Wikipedia:Text_of_the_Creative_Commons_Attribution-ShareAlike_4.0_International_License" title="Wikipedia:Text of the Creative Commons Attribution-ShareAlike 4.0 International License">Creative Commons Attribution-ShareAlike 4.0 License</a>; additional terms may apply. By using this site, you agree to the <a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Terms_of_Use" class="extiw" title="foundation:Special:MyLanguage/Policy:Terms of Use">Terms of Use</a> and <a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy" class="extiw" title="foundation:Special:MyLanguage/Policy:Privacy policy">Privacy Policy</a>. Wikipedia® is a registered trademark of the <a rel="nofollow" class="external text" href="https://wikimediafoundation.org/">Wikimedia Foundation, Inc.</a>, a non-profit organization.</li> </ul> <ul id="footer-places"> <li id="footer-places-privacy"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy">Privacy policy</a></li> <li id="footer-places-about"><a href="/wiki/Wikipedia:About">About Wikipedia</a></li> <li id="footer-places-disclaimers"><a href="/wiki/Wikipedia:General_disclaimer">Disclaimers</a></li> <li id="footer-places-contact"><a href="//en.wikipedia.org/wiki/Wikipedia:Contact_us">Contact Wikipedia</a></li> <li id="footer-places-wm-codeofconduct"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Universal_Code_of_Conduct">Code of Conduct</a></li> <li id="footer-places-developers"><a href="https://developer.wikimedia.org">Developers</a></li> <li id="footer-places-statslink"><a href="https://stats.wikimedia.org/#/en.wikipedia.org">Statistics</a></li> <li id="footer-places-cookiestatement"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Cookie_statement">Cookie statement</a></li> <li id="footer-places-mobileview"><a href="//en.m.wikipedia.org/w/index.php?title=Edinger%E2%80%93Westphal_nucleus&mobileaction=toggle_view_mobile" class="noprint stopMobileRedirectToggle">Mobile view</a></li> </ul> <ul id="footer-icons" class="noprint"> <li id="footer-copyrightico"><a href="https://wikimediafoundation.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/static/images/footer/wikimedia-button.svg" width="84" height="29" alt="Wikimedia Foundation" loading="lazy"></a></li> <li id="footer-poweredbyico"><a href="https://www.mediawiki.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/w/resources/assets/poweredby_mediawiki.svg" alt="Powered by MediaWiki" width="88" height="31" loading="lazy"></a></li> </ul> </footer> </div> </div> </div> <div class="vector-settings" id="p-dock-bottom"> <ul></ul> </div><script>(RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgHostname":"mw-web.codfw.main-f69cdc8f6-55zsq","wgBackendResponseTime":158,"wgPageParseReport":{"limitreport":{"cputime":"0.696","walltime":"0.876","ppvisitednodes":{"value":2376,"limit":1000000},"postexpandincludesize":{"value":152617,"limit":2097152},"templateargumentsize":{"value":2586,"limit":2097152},"expansiondepth":{"value":10,"limit":100},"expensivefunctioncount":{"value":4,"limit":500},"unstrip-depth":{"value":1,"limit":20},"unstrip-size":{"value":83171,"limit":5000000},"entityaccesscount":{"value":1,"limit":400},"timingprofile":["100.00% 752.677 1 -total"," 20.02% 150.658 13 Template:Navbox"," 18.83% 141.703 1 Template:Infobox_brain"," 18.23% 137.201 1 Template:Infobox_anatomy"," 17.26% 129.914 1 Template:Infobox"," 17.12% 128.864 4 Template:Citation"," 14.50% 109.168 1 Template:Mesencephalon"," 12.23% 92.075 1 Template:Short_description"," 8.95% 67.357 1 Template:Expand_section"," 8.06% 60.697 1 Template:Ambox"]},"scribunto":{"limitreport-timeusage":{"value":"0.433","limit":"10.000"},"limitreport-memusage":{"value":6858062,"limit":52428800}},"cachereport":{"origin":"mw-api-ext.codfw.main-7556f8b5dd-mh5wj","timestamp":"20241122150901","ttl":2592000,"transientcontent":false}}});});</script> <script type="application/ld+json">{"@context":"https:\/\/schema.org","@type":"Article","name":"Edinger\u2013Westphal nucleus","url":"https:\/\/en.wikipedia.org\/wiki\/Edinger%E2%80%93Westphal_nucleus","sameAs":"http:\/\/www.wikidata.org\/entity\/Q165312","mainEntity":"http:\/\/www.wikidata.org\/entity\/Q165312","author":{"@type":"Organization","name":"Contributors to Wikimedia projects"},"publisher":{"@type":"Organization","name":"Wikimedia Foundation, Inc.","logo":{"@type":"ImageObject","url":"https:\/\/www.wikimedia.org\/static\/images\/wmf-hor-googpub.png"}},"datePublished":"2005-06-17T11:33:47Z","dateModified":"2024-10-16T09:32:00Z","image":"https:\/\/upload.wikimedia.org\/wikipedia\/commons\/a\/a4\/Cn3nucleus-en.svg","headline":"one of two nuclei of the oculomotor nerve"}</script> </body> </html>