CINXE.COM
Grey wolf | Canids
<!DOCTYPE HTML> <html> <head> <title>Grey wolf | Canids</title> <meta http-equiv="Content-Type" content="text/html; charset=UTF-8"> <meta name="keywords" content="Grey wolf"> <meta name="description" content="Grey wolf"> <meta name="alberon-uptime-monitor-tag" content="ok"> <link href='//fonts.googleapis.com/css?family=Orienta' rel='stylesheet' type='text/css'> <!-- base css, generic styles that can be overridden --> <link rel="stylesheet" type="text/css" href="/app/css/base.css" /> <link rel="stylesheet" href="/app/layouts/brochure/canids/layout.css?ver=4" type="text/css"> <script type="text/javascript" src="/jquery/jquery.min.js"></script> <script type="text/javascript" src="/jquery/ui.core.js"></script> <script type="text/javascript" src="/jquery/ui.draggable.js"></script> <script type="text/javascript" src="/jquery/ui.resizable.js"></script> <script type="text/javascript" src="/jquery/ui.dialog.js"></script> <script type="text/javascript" src="/jquery/ui.droppable.js"></script> <script type="text/javascript" src="/jquery/rating/jquery.rating.js"></script> <script type="text/javascript" src="/jquery/hoverbutton.oww.js"></script> <script type="text/javascript" src="/jquery/search.js"></script> <link rel="stylesheet" type="text/css" href="/jquery/dialog/jquery-ui-themeroller.css" > <link rel="stylesheet" type="text/css" href="/jquery/rating/jquery.rating.css" > <script type="text/javascript"> var DDSPEED = 5; var DDTIMER = 15; // main function to handle the mouse events // // d=direction of fade, 1=fade in, -1 = fade out. function ddMenu(id,d){ var h = document.getElementById(id + '-ddheader'); var c = document.getElementById(id + '-ddcontent'); clearInterval(c.timer); if(d == 1){ clearTimeout(h.timer); if(c.maxh && c.maxh <= c.offsetHeight){return} else if(!c.maxh){ c.style.display = 'block'; c.style.height = 'auto'; c.maxh = c.offsetHeight; c.style.height = '0px'; } c.timer = setInterval(function(){ddSlide(c,1)},DDTIMER); }else{ h.timer = setTimeout(function(){ddCollapse(c)},50); } } // collapse the menu // function ddCollapse(c){ c.timer = setInterval(function(){ddSlide(c,-1)},DDTIMER); } // cancel the collapse if a user rolls over the dropdown // function cancelHide(id){ var h = document.getElementById(id + '-ddheader'); var c = document.getElementById(id + '-ddcontent'); clearTimeout(h.timer); clearInterval(c.timer); if(c.offsetHeight < c.maxh){ c.timer = setInterval(function(){ddSlide(c,1)},DDTIMER); } } // incrementally expand/contract the dropdown and change the opacity // function ddSlide(c,d){ var currh = parseInt(c.style.height) ; //offsetHeight; var dist; if(d == 1){ dist = (Math.round((c.maxh - currh) / DDSPEED)); }else{ dist = (Math.round(currh / DDSPEED)); } if(dist <= 1 ) { dist = 1; } c.style.height = currh + (dist * d) + 'px'; c.style.opacity = currh / c.maxh; c.style.filter = 'alpha(opacity=' + (currh * 100 / c.maxh) + ')'; if((currh < 2 && d != 1) || (currh > (c.maxh - 2) && d == 1)){ clearInterval(c.timer); } } </script> </head> <body onload=""> <div id="container"> <div id="header"><img style=" height: 60px; margin-left: 10px; margin-top: 5px;" alt="Canids Specialist Group" src="/app/images/structure/canids-logo-brown.jpg"><div class="housekeepingmenu"> <div class="menuitem"><a class="off" href="/support-" >Support us</a></div><div class="separator">|</div> <div class="menuitem"><a class="off" href="/tools-1" >Tools</a></div><div class="separator">|</div> <div class="menuitem"><a class="off" href="/photo-gallery" >Photo Gallery</a></div><div class="separator">|</div> <div class="menuitem"><a class="off" href="/useful-links" >Useful Links</a></div><div class="separator">|</div> <div class="menuitem"><a class="off" href="/contact" >Contact</a></div></div><!-- housekeepingmenu --> </div><!-- header --> <div id="navigation"> <div id="nav"><ul class="dropdown"><li id="home-ddheader" class=""><a href="/home">Home</a></li></ul><ul class="dropdown"><li id="PNKHEJ389685-ddheader" onmouseout="ddMenu('PNKHEJ389685', -1)" onmouseover="ddMenu('PNKHEJ389685', 1)" class=""><a href="/PNKHEJ389685">About</a></li><div id="PNKHEJ389685-ddcontent" onmouseout="ddMenu('PNKHEJ389685', -1)" onmouseover="cancelHide('PNKHEJ389685')" class="submenu"><div class="submenuitem"><a href="/groups">Working Groups</a></div><div class="submenuitem"><a href="/what-we-do">What We Do</a></div><div class="submenuitem"><a href="/who-we-are">Who We Are</a></div><div class="submenuitem"><a href="/histor">History</a></div><div class="submenuitem"><a href="/sponsors-1">Sponsors</a></div></div></ul><ul class="dropdown"><li id="speciespage-ddheader" onmouseout="ddMenu('speciespage', -1)" onmouseover="ddMenu('speciespage', 1)" class=""><a href="/speciespage">Species</a></li><div id="speciespage-ddcontent" onmouseout="ddMenu('speciespage', -1)" onmouseover="cancelHide('speciespage')" class="submenu"><div class="submenuitem fatMenu"><div style="vertical-align: top; display: inline-block; margin-right: 10px;"><div class="name">South America</div><div class="species"><a href="/species/view/PREKFN688091">Short-eared dog</a></div><div class="species"><a href="/species/view/PREKFY360401">Crab-eating fox</a></div><div class="species"><a href="/species/view/PREKIV875331">Bush dog</a></div><div class="species"><a href="/species/view/PREKIN694741">Hoary fox</a></div><div class="species"><a href="/species/view/PREKI2182561">Pampas Fox</a></div><div class="species"><a href="/species/view/PREKIA411651">Sechuran Fox</a></div><div class="species"><a href="/species/view/PREKGA855611">Maned wolf</a></div><div class="species"><a href="/species/view/PREKHM366471">Darwin's fox</a></div><div class="species"><a href="/species/view/PREKGI446591">Culpeo</a></div><div class="species"><a href="/species/view/PREKHU172531">Chilla</a></div></div></div><div class="submenuitem fatMenu"><div style="vertical-align: top; display: inline-block; margin-right: 10px;"><div class="name">Central & North America</div><div class="species"><a href="/species/view/PREKJA587951">Coyote</a></div><div class="species"><a href="/species/view/PREKJI659751">Red wolf</a></div><div class="species"><a href="/species/view/PREKJP597461">Gray fox</a></div><div class="species"><a href="/species/view/PREKJY135341">Island fox</a></div><div class="species"><a href="/species/view/PREKK8422331">Kit fox</a></div><div class="species"><a href="/species/view/PREKKG784471">Swift fox</a></div></div></div><div class="submenuitem fatMenu"><div style="vertical-align: top; display: inline-block; margin-right: 10px;"><div class="name">Europe & North/Central Asia</div><div class="species"><a href="/species/view/PREKKS663901">Arctic fox</a></div><div class="species"><a href="/species/view/PREKLD895731">Grey wolf</a></div><div class="species"><a href="/species/view/PREKLP237241">Red fox</a></div><div class="species"><a href="/species/view/PREKLY462191">Raccoon dog</a></div><div class="species"><a href="/species/view/PREKM6521571">Corsac</a></div><div class="species"><a href="/species/view/PREKMF443401">Tibetan fox</a></div></div></div><div class="submenuitem fatMenu"><div style="vertical-align: top; display: inline-block; margin-right: 10px;"><div class="name">Sub-Saharan Africa</div><div class="species"><a href="/species/view/PREKMO428071">Side-striped jackal</a></div><div class="species"><a href="/species/view/PREKMW872861">Golden jackal</a></div><div class="species"><a href="/species/view/PREKN8360151">Black-backed jackal</a></div><div class="species"><a href="/species/view/PREKNL433181">Ethiopian wolf</a></div><div class="species"><a href="/species/view/PREKNU724601">African wild dog</a></div><div class="species"><a href="/species/view/PREKO4241281">Bat-eared fox</a></div><div class="species"><a href="/species/view/PREKOC822211">Cape fox</a></div></div></div><div class="submenuitem fatMenu"><div style="vertical-align: top; display: inline-block; margin-right: 10px;"><div class="name">North Africa & the Middle East</div><div class="species"><a href="/species/view/PREKON160201">Blanford's fox</a></div><div class="species"><a href="/species/view/PREKOV287451">Pale fox</a></div><div class="species"><a href="/species/view/PREKP7952371">Ruppell's fox</a></div><div class="species"><a href="/species/view/PREKPF521541">Fennec fox</a></div></div></div><div class="submenuitem fatMenu"><div style="vertical-align: top; display: inline-block; margin-right: 10px;"><div class="name">South Asia</div><div class="species"><a href="/species/view/PREKQW171301">Dhole</a></div><div class="species"><a href="/species/view/PREKR4188521">Bengal fox</a></div><div class="species"><a href="/species/view/WCD6DU963961">Dingo </a></div></div></div></div></ul><ul class="dropdown"><li id="members-ddheader" class=""><a href="/members">Members</a></li></ul><ul class="dropdown"><li id="projects-ddheader" class=""><a href="/projects">Projects</a></li></ul><ul class="dropdown"><li id="publications-ddheader" class=""><a href="/publications">Publications</a></li></ul><ul class="dropdown"><li id="c-b-c-ddheader" onmouseout="ddMenu('c-b-c', -1)" onmouseover="ddMenu('c-b-c', 1)" class=""><a href="/c-b-c">Journal</a></li><div id="c-b-c-ddcontent" onmouseout="ddMenu('c-b-c', -1)" onmouseover="cancelHide('c-b-c')" class="submenu"><div class="submenuitem"><a href="/about-canid-biology-and-conservation">About CBC</a></div><div class="submenuitem"><a href="/editorial-board">Editorial Board</a></div><div class="submenuitem"><a href="/call-for-papers">Call for Papers</a></div><div class="submenuitem"><a href="/instructions-for-authors">Instructions to authors</a></div><div class="submenuitem"><a href="/all-articles">Previous Issues</a></div><div class="submenuitem"><a href="/recent-articles">Recent Issues</a></div><div class="submenuitem"><a href="/Q6WN21890762">----------------------------</a></div></div></ul><ul class="dropdown"><li id="news-and-events-ddheader" class=""><a href="/news-and-events">News</a></li></ul><ul class="dropdown"><li id="working-groups-ddheader" class=""><a href="/working-groups">Working Groups</a></li></ul></div><script type="text/javascript"> $(document).ready(function() { $('.fatMenu').css('display', 'inline-block'); $('.dropdown').each(function() { var menuContainer = $(this); var topMenuItem = menuContainer.children('li'); var subMenu = menuContainer.children('.submenu'); var fatMenuElements = subMenu.children('.fatMenu'); if (fatMenuElements.size() == 0) return; menuContainer.data('fatMenu', { _data : { menu : menuContainer, topMenuItem : topMenuItem, subMenu : subMenu, fatMenuElements : fatMenuElements }, setup : function() { var noElements = this._data.fatMenuElements.size(); this._data.subMenu.width(noElements * 133); this._data.subMenu.find('div').css('border', 'none'); } }); menuContainer.data('fatMenu').setup(); }); }); </script><div style="clear:both;"></div></div><div id="headerpic"></div> <div id="outercontent"><div class="contentSpacer"></div><div id="leftPane"><div class="searchBox"><form method="post" action="/sitesearch">Search <input name="q" type="text" class="search" /><input type="submit" style="display: none;" /></form></div><script type="text/javascript"> $(document).ready(function() { $('.search').keyup(function(e) { var searchObject = $(this); if (e.which == 13) return true; if (searchObject.data('jqXhtmlSearch') != null) { searchObject.data('jqXhtmlSearch').abort(); searchObject.data('jqXhtmlSearch', null); } if (searchObject.data('searchResults') != null) { searchObject.data('searchResults').remove(); searchObject.data('searchResults', null); } var criteria = searchObject.val(); if (criteria.length < 3) return; var uri = '/sitesearch'; searchObject.data('jqXhtmlSearch', $.get(uri, { ajax : 1, q : criteria, limit : 10 }, function(data) { var searchResults = $('<div class="searchResults"/>'); searchObject.data('searchResults', searchResults); searchResults.css('position', 'absolute'); searchResults.css('top', searchObject.position().top + searchObject.height() + 6); searchResults.css('left', searchObject.position().left + 2); searchResults.html(data); searchResults.find('a').click(function() { searchResults.hide(); }); $('body').append(searchResults); })); }); }); </script><ul class="sideMenu"><li class=""><a href="/species/region/PREKDB770131">South America</a></li><li class=""><a href="/species/region/PREKDL133611">Central & North America</a></li><li class="selected"><a href="/species/region/PREKDX515331">Europe & North/Central Asia</a></li><li class=""><a href="/species/region/PREKE8430791">Sub-Saharan Africa</a></li><li class=""><a href="/species/region/PREKEJ362401">North Africa & the Middle East</a></li><li class=""><a href="/species/region/PREKQ6133971">South Asia</a></li></ul><div class="subscribeBox">Subscribe to <a href="/subscribe">Canids-L mailing List</a></div><p style="text-align: center;"><a href="https://www.youtube.com/channel/UCgZH3Opj68FzAThu75cEkgw/feed" target="_blank"><img style="margin-left: 5px; margin-right: 5px; border: 0px;" title="Follow our Youtube channel" src="/app/images/icons/youtube.png" alt="Follow our Youtube channel" width="79" height="33" /></a><a href="https://www.facebook.com/Canidconservation" target="_blank"><img style="border: 0px;" title="Find us on Facebook" src="/app/images/icons/facebook-icon.png" alt="Find us on Facebook" width="33" height="33" /></a> <a href="https://twitter.com/WildCanids" target="_blank"><img src="/app/images/icons/twitter-icon.png" alt="" width="32" height="33" /></a></p></div><div id="innercontent"> <h1><span class="mainMenuItemBread">Europe & North/Central Asia</span><span class="subMenuItem">Grey wolf</span><span class="scientificName"> Canis lupus</span></h1><div class="pageImages"><img src="/app/images/canids/grey-wolf/greywolf.jpg" title="Endangered "Mexican wolf" subspecies (C.l.baileyi) - © Rurik List"><p>Endangered "Mexican wolf" subspecies (<em>C.l.baileyi</em>) - © Rurik List</p><p><a href="/resources/view/RESNRI805921"><strong>Wolf Working Group</strong></a> - The Wolf working group (formally the Wolf specialist group) is an international organisation of experts on wolves. With wolf conservation matters of internal significance, cooperation across the key geographic areas is paramount. The working group plays a key role in this by allowing for the joint planning of conservation programs, exchange of experiences, research and publications and an assembly of knowledgeable personnel across the globe. </p><strong>Projects</strong><ul class="resources"><li><a href="/resources/view/UFC1BN880851">The Wood River Wolf Project, Idaho USA</a></li><li><a href="/resources/view/RDXG0Q621951">Wolf-deer research in the Superior National Forest of Minnesota</a></li><li><a href="/resources/view/RDXNSR234511">Context dependent landscapes of fear with the grey wolf </a></li><li><a href="/resources/view/RDXVU2336051">Using citizen science to better understand Yellowstone’s wolves</a></li><li><a href="/resources/view/UCARMB449641">Himalayan Wolves Project</a></li><li><a href="/resources/view/RDXS5U296011">Canid conservation in the Gobi-Steppe ecosystem of Mongolia</a></li><li><a href="/resources/view/UNROOJ589581">Project Coyote - Ranching with Wildlife</a></li><li><a href="/resources/view/UO64NP155651">Iberian Wolf Portuguese National Census</a></li><li><a href="/resources/view/UO64OX700051">Wind Farms & Iberian Wolf</a></li><li><a href="/resources/view/UPBEM7907981">Coastal Wolf Ecology</a></li></ul><strong>Publications</strong><ul class="resources"><li><a href="https://www.canids.org/resources/CSG%20France%202021%20Wolf%20Culling%20Proposal%2010Nov2020.pdf">Letter addressing the proposed French wolf quota for 2021</a></li><li><a href="/resources/view/WDC7OF772031">Letter to the Swiss Government objecting the Wolf Management Plans for the Winter 2023-2024</a></li><li><a href="https://www.canids.org/resources/LCIE%20-%20statement%20on%20wolf%20downlisting%20proposal(1).pdf">LCIE 2024. Statement on the proposed downlisting of the wolf under the Bern Convention and the EU Habitats Directive</a></li></ul><strong>Relevant Links</strong><ul class="resources"><li><a href="http://www.flickr.com/groups/canidguide/">Flickr: Wild Canids of the World</a></li><li><a href="https://www.iucnredlist.org/species/3746/163508960">2018 IUCN Red List Assessment - Grey Wolf</a></li><li><a href="http://www.wolf.org/wolves/">International Wolf Centre</a></li><li><a href="http://fohn.net/wolf-pictures-facts/">The Wolf Den: Wolf Pictures and Facts</a></li><li><a href="http://www.pbs.org/wgbh/nova/wolves/">NOVA Online - Wild Wolves</a></li><li><a href="http://www.lcie.org/">IUCN / SSC Large Carnivore Initiative for Europe</a></li><li><a href="http://www.iberianature.com/material/wolf.html">Iberian Wolf - Wolves in Spain</a></li><li><a href="http://loup.org/spip/">Sur la piste du loup/On the wolf track</a></li><li><a href="http://www.canislupus.it">CSDL (Centro per lo Studio e la Documentazione sul Lupo)</a></li><li><a href="http://www.defenders.org/gray-wolf/basic-facts">Defenders of Wildlife - Gray Wolf</a></li><li><a href="http://www.wolvesontario.org">Wolves Ontario Project - An Earthroots Project</a></li></ul><strong>Reports / Papers</strong><ul class="resources"><li><a href="/resources/view/Q7NT8M406991">2004 Status Survey & Conservation Action Plan - Europe & North/Central Asia</a></li></ul><strong>Other Names</strong><h5><span style="font-size: small;">English: <span style="font-weight: normal;">Grey Wolf, Arctic Wolf, Common Wolf, Gray Wolf, Mexican Wolf, Plains Wolf, Timber Wolf, Tundra Wolf, Wolf</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">French: <span style="font-weight: normal;">Loup, Loup Gris, Loup Vulgaire</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Spanish; Castilian: <span style="font-weight: normal;">Lobo</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Albanian:<span style="font-weight: normal;"> Ujku</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Arabic: <span style="font-weight: normal;">Dheeb, ????? ??????</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Croatian: <span style="font-weight: normal;">Vuk</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Czech: <span style="font-weight: normal;">Vlk</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Estonian: <span style="font-weight: normal;">Hunt</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Finnish: <span style="font-weight: normal;">Susi</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">German: <span style="font-weight: normal;">Wolf</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Greek, Modern: <span style="font-weight: normal;">λ?κος</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Hungarian: <span style="font-weight: normal;">Farkas</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Italian: <span style="font-weight: normal;">Lupo</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Latvian: <span style="font-weight: normal;">Vilks</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Lithuanian: <span style="font-weight: normal;">Vilkas</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Mongolian: <span style="font-weight: normal;">Saaral Chono</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Norwegian: <span style="font-weight: normal;">Ulv</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Polish: <span style="font-weight: normal;">Wilk</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Romanian: <span style="font-weight: normal;">Lup</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Russian: <span style="font-weight: normal;">????</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Serbian: <span style="font-weight: normal;">???</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Slovak: <span style="font-weight: normal;">Vlk</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Slovenian: <span style="font-weight: normal;">Volk</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Swedish: <span style="font-weight: normal;">Varg</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Turkish: <span style="font-weight: normal;">Kurt</span></span></h5> <p><span style="font-size: small;"> </span></p> <h5><span style="font-size: small;">Ukrainian: <span style="font-weight: normal;">????</span></span></h5></div><a class="statusImage" href="http://www.redlist.org/static/categories_criteria_3_1#categories"><img style="width:404px;" alt="Least Concern" src="/app/images/iucn-categories/least-concern.jpg" /></a><h4>Taxonomic Notes</h4> <p>In Europe, two subspecies are recognized: <em>C. l. signatus</em> (Iberia) and <em>C. l. italicus</em> (Italy, France and Switzerland).<br /><br />In Asia, two subspecies are recognized: <em>C. l. pallipes</em> (most of the Asian range from Israel to China) and <em>C. l. arabs</em> (Arabian peninsula). Furthermore, wolves of the Himalayan range have been proposed as a distinct subspecies (<em>C. l. chanco</em>).<br /><br />In North America, five subspecies are often recognized: <em>C. l. arctos</em> (Arctic wolf), <em>C. l. lycaon</em> (Eastern wolf), which Chambers <em>et al</em>. (2012) consider a distinct species, <em>C. l. nubilus</em> (Plains wolf), <em>C. l. occidentalis</em> (Northwestern wolf or Northern timber wolf), and <em>C. l. baileyi </em>(Mexican wolf).<br /><br />Note that this assessment follows Jackson <em>et al</em>. (2017) in regarding the Dingo, sometimes considered a subspecies of Grey Wolf (<em>C. l. dingo),</em> as a feral dog population derived from a domesticate, and hence as <em>C. familiaris</em>, along with all other free-ranging dogs.</p> <h4>Justification</h4> <p>Originally, the Grey Wolf was the world's most widely distributed mammal. It has become extinct in much of Western Europe, in Mexico and much of the USA, and their present distribution is more restricted; wolves occur primarily but not exclusively in wilderness and remote areas. Their original worldwide range has been reduced by about one-third by deliberate persecution due to depredation on livestock and fear of attacks on humans. Since about 1970, legal protection, land-use changes, rural human population shifts to cities and increasing prey populations have arrested wolf population declines and fostered natural recolonization in parts of its range and reintroduction in three areas of USA. Continued threats include competition with humans for livestock and game species, exaggerated concern by the public regarding the threat and danger of wolves, and fragmentation of habitat, with resulting areas becoming too small for populations with long-term viability.<br /><br />Although the Grey Wolf still faces some threats, its relatively widespread range and stable population trend mean that the species, at global level, does not meet, or nearly meet, any of the criteria for the threatened categories. The global population is estimated to be in the order of 200-250 thousands individuals. Therefore, it is assessed as Least Concern. However, at regional level, several wolf populations, such as those in Europe, are seriously threatened (<a href="http://www.lcie.org/">http://www.lcie.org</a>).</p> <h4>Geographic Range Information</h4> <p>Originally, the Grey Wolf was the world's most widely distributed mammal, living throughout the northern hemisphere north of 15°N latitude in North America and 12°N in India. It became extinct in much of Western Europe (Boitani 1995), Mexico and much of the USA (Mech 1970) and Japan. Their present distribution is more restricted: wolves occur primarily in wilderness and remote areas, especially in Canada, Alaska and northern USA, Europe, and Asia from about 75°N to 12°N (Mech and Boitani 2004), but they are found also in human-dominated landscapes where there is sufficient prey base and human-caused mortality is low. Distribution is highly dynamic as wolf populations are currently increasing in range and numbers in north-central and western United States and much of Europe. Dispersing individuals can be found in almost any European country except the United Kingdom and Ireland.</p><p><strong>Population trend:</strong>Stable</p><a href="https://f002.backblazeb2.com/file/cached-individual-maps/T3746A163508960.jpg?Authorization=3_20210112200937_b7545942ac8b1912be56d2b0_613675df446437f211c7e12806baf6efa254c9a8_002_20210119200937_0019_dnld" target="_blank"><div><img class="rangeMap" src="/app/images/grey-wolf-map.jpg" alt="(Grey wolf range map)" /></div></a><div>(Click on map for more detail)</div><h4>Population Information</h4> <p>Because of the diversity in climate, topography, vegetation, human settlement and development of wolf range, wolf populations in various parts of the original range vary from extinct to relatively pristine. Wolf densities vary from about one/12 km² to one/120 km².<br /><br />Mech and Boitani (2004) provide details, for each range country, on subspecies present, population status, approximate numbers, the percentage of former range occupied at present, main prey (where known), legal status, and cause of decline. For Europe, the wolf population is essentially a large meta-population with several distinct subpopulations; the status and trends for each population is provided by the Large Carnivore Initiative for Europe (<a href="http://www.lcie.org/">http://www.lcie.org/</a>) and in Chapron <em>et al</em>. (2014). The status and trends for wolf populations in the United States are provided by the U.S. Fish and Wildlife Service and game agencies in several US states, including Minnesota, Wisconsin, Michigan, Montana, Idaho, Wyoming, Oregon, Washington, Arizona, and New Mexico. In general, population increases in North America and Europe are likely to be off-set by localized declines in other parts of the range.</p> <h4>Habitat and Ecology Information</h4> <p>Ranges in all northern habitats where there is suitable food (Mech 1970), densities being highest where prey biomass is highest (Fuller 1989). Food is extremely variable, but the majority comprises large ungulates (moose, caribou, deer, elk, wild boar, etc.). Wolves will also eat smaller prey items, livestock, carrion, and garbage.</p> <h4>Threats Information</h4> <p>The historical range of the Grey Wolf has been reduced by about one-third, primarily in developed areas of Europe, Asia, Mexico, and the United States by poisoning and deliberate persecution due to depredation on livestock. Since about 1970, legal protection, land-use changes, and rural human population shifts to cities have arrested wolf population declines and fostered natural recolonization in parts of Western Europe and the United States, and reintroduction in the western United States and Mexico. Continued threats include competition with humans for livestock, especially in developing countries, exaggerated concern by the public concerning the threat and danger of wolves. Fragmentation of habitat, with resulting areas becoming too small for populations with long-term viability can be a threat in parts of the range with highest density of human activities.</p> <h4>Use and Trade Information</h4> <p>There is sustainable utilization of the species' fur in Canada, Alaska, Montana and the former Soviet Union and Mongolia, although illegal hunting for fur does occur in some range states.</p> <h4>Conservation Actions Information</h4> <p><strong>Legislation</strong><br />The species is included in CITES Appendix II, except populations from Bhutan, India, Nepal and Pakistan, which are listed on Appendix I. The species is strictly protected under the Bern Convention (Appendix II) and Habitats Directive (Annex II and IV), and there is extensive legal protection in many European countries; however, there are national exceptions, and enforcement is variable and often non-existent. <br /><strong><br /></strong><strong>Presence in protected areas</strong><br />The Grey Wolf occurs in many protected areas across its range. <br /><br />Occurrence in captivity<br />The species lives and breeds well in captivity and is common in many zoological gardens.<br /><br />Gaps in knowledge<br />One of the most important questions still remaining about wolves involves the nature of their interaction with prey populations. The conditions under which wolves limit, regulate, or control their population is still open and important (Mech and Boitani 2003). Of recent and increasing importance is clarifying the role that wolves play in precipitating ecosystem effects, particularly trophic cascades (Mech and Boitani 2003, Ripple and Betscha 2004, Peterson <em>et al</em>. 2014). Of more academic interest are questions involving wolf genetics, scent-marking behaviour, pseudo pregnancy and diseases (Mech 1995). Of recent and increasing importance is the issue of hybridization with dogs (and indeed other <em>Canis</em> spp) in several parts of the range in Europe and Asia, with unknown impact on medium- and long-term genetic integrity of the species.</p></div></div><div style="clear: both;"></div></div><div id="footer"><div class="copyrightNotice" style="width:610px"><div> The Canid Specialist Group is housed by WildCRU at Tubney House, Oxford, UK. Claudio Sillero's position at WildCRU is funded by <a target="_blank" href="http://www.bornfree.org.uk/">The Born Free Foundation.</a> Copyright 漏 <?php echo date('Y'); ?> by the IUCN/SSC Canid Specialist Group. All rights reserved. <a href="http://www.oxford-web.co.uk">Website developed by oxford web</a> and designed by Teresa Fuertes. </div></div><div class="sponsors"> <img alt="Wildlife Conservation Research Unit" src="/app/images/structure/wild-cru.png" /> <img src="/app/images/structure/born-free.png" alt="Born Free Foundation"/></div><div style="clear: both;"></div></div></body></html>