A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region

<!DOCTYPE html> <html lang="en-US" xml:lang="en-US"> <head> <meta charset="utf-8"> <meta name="viewport" content="width=device-width, initial-scale=1.0"> <title> A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region | Zootaxa </title> <meta name="generator" content="Open Journal Systems 3.3.0.6"> <link rel="schema.DC" href="http://purl.org/dc/elements/1.1/" /> <meta name="DC.Creator.PersonalName" content="NATHANIEL N. LEVIA"/> <meta name="DC.Creator.PersonalName" content="CHARLES R. BARTLETT"/> <meta name="DC.Date.created" scheme="ISO8601" content="2025-02-21"/> <meta name="DC.Date.dateSubmitted" scheme="ISO8601" content="2025-02-20"/> <meta name="DC.Date.issued" scheme="ISO8601" content="2025-02-21"/> <meta name="DC.Date.modified" scheme="ISO8601" content="2025-02-21"/> <meta name="DC.Description" xml:lang="en" content="The genera Atracodes Melichar, and Pseudoflatoides Metcalf, are small Antillean genera in the flatid subfamily Flatoidinae. We found Pseudoflatoides to be a junior subjective synonym of Atracodes resulting in the transfer of four species (and seven subspecies). We revise the status of two subspecies of Atracodes fasciculosus comb. nov. to species level (as Atracodes fasciatus comb. nov. &amp; stat. nov. and Atracodes vittatus comb. nov. &amp; stat. nov.) based on characteristics of the male terminalia. Atracodes corrugatus Melichar (from Madagascar) is transferred to Lichenopsis Schmidt as Lichenopsis corrugata (Melichar), comb. nov. The resulting composition of Atracodes is 10 species, and four subspecies. We also compile available information on the two genera, provide a key to the American genera of Flatoidinae, along with a key to species of Atracodes, and suggest future work on the subfamily. "/> <meta name="DC.Format" scheme="IMT" content="application/pdf"/> <meta name="DC.Identifier" content="zootaxa.5590.2.4"/> <meta name="DC.Identifier.pageNumber" content="231-255 "/> <meta name="DC.Identifier.DOI" content="10.11646/zootaxa.5590.2.4"/> <meta name="DC.Identifier.URI" content="https://140.82.21.59/zt/article/view/zootaxa.5590.2.4"/> <meta name="DC.Language" scheme="ISO639-1" content="en"/> <meta name="DC.Rights" content="Copyright (c) 2025 "/> <meta name="DC.Rights" content=""/> <meta name="DC.Source" content="Zootaxa"/> <meta name="DC.Source.ISSN" content="1175-5334"/> <meta name="DC.Source.Issue" content="2"/> <meta name="DC.Source.Volume" content="5590"/> <meta name="DC.Source.URI" content="https://140.82.21.59/zt"/> <meta name="DC.Subject" xml:lang="en" content="iNaturalist"/> <meta name="DC.Title" content="<strong>A review of the genera <em>Atracodes</em> Melichar and <em>Pseudoflatoides</em> Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region</strong>"/> <meta name="DC.Type" content="Text.Serial.Journal"/> <meta name="DC.Type.articleType" content="Article"/> <meta name="gs_meta_revision" content="1.1"/> <meta name="citation_journal_title" content="Zootaxa"/> <meta name="citation_journal_abbrev" content="Zootaxa"/> <meta name="citation_issn" content="1175-5334"/> <meta name="citation_author" content="NATHANIEL N. LEVIA"/> <meta name="citation_author_institution" content="University of Delaware; Department of Entomology and Wildlife Ecology; 250 Townsend Hall; Newark; DE 19716-2160; USA."/> <meta name="citation_author" content="CHARLES R. BARTLETT"/> <meta name="citation_author_institution" content="University of Delaware; Department of Entomology and Wildlife Ecology; 250 Townsend Hall; Newark; DE 19716-2160; USA."/> <meta name="citation_title" content="<strong>A review of the genera <em>Atracodes</em> Melichar and <em>Pseudoflatoides</em> Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region</strong>"/> <meta name="citation_language" content="en"/> <meta name="citation_date" content="2025/02/21"/> <meta name="citation_volume" content="5590"/> <meta name="citation_issue" content="2"/> <meta name="citation_firstpage" content="231"/> <meta name="citation_lastpage" content="255"/> <meta name="citation_doi" content="10.11646/zootaxa.5590.2.4"/> <meta name="citation_abstract_html_url" content="https://140.82.21.59/zt/article/view/zootaxa.5590.2.4"/> <meta name="citation_keywords" xml:lang="en" content="Hemiptera"/> <meta name="citation_keywords" xml:lang="en" content="Flatoidini"/> <meta name="citation_keywords" xml:lang="en" content="Caribbean"/> <meta name="citation_keywords" xml:lang="en" content="new synonymy"/> <meta name="citation_keywords" xml:lang="en" content="new combination"/> <meta name="citation_keywords" xml:lang="en" content="iNaturalist"/> <meta name="citation_pdf_url" content="https://140.82.21.59/zt/article/download/zootaxa.5590.2.4/55693"/> <meta name="citation_reference" content="Amyot, C.J.-B. & Audinet-Serville, J.G. 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href="https://doi.org/10.11646/zootaxa.5590.2.4"> 10.11646/zootaxa.5590.2.4 </a> </span> </div> <div class="pages"> <span>Page range:</span> 231-255 </div> <div class="views"> <span>Abstract views:</span> 75 </div> <div class="views"> <span>PDF downloaded:</span> 9 </div> </div> <h1 class="jatsParser__meta-title"><strong>A review of the genera <em>Atracodes</em> Melichar and <em>Pseudoflatoides</em> Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region</strong></h1> <ul class="jatsParser__meta-authors"> <li><a class="jatsParser__meta-author-string-href" href="#author-1"><span class="jatsParser__meta-author">NATHANIEL N. LEVIA</span><sup class="jatsParser__meta-symbol jatsParser__symbol-plus">+</sup><sup class="jatsParser__meta-symbol jatsParser__symbol-minus jatsParser__hide">−</sup></a><a class="jatsParser__meta-orcidImage" href="https://orcid.org/0009-0006-8436-7660"><img src="https://140.82.21.59/plugins/themes/oldGregg/templates/images/orcid.png"></a></li> <li><a class="jatsParser__meta-author-string-href" href="#author-2"><span class="jatsParser__meta-author">CHARLES R. BARTLETT</span><sup class="jatsParser__meta-symbol jatsParser__symbol-plus">+</sup><sup class="jatsParser__meta-symbol jatsParser__symbol-minus jatsParser__hide">−</sup></a><a class="jatsParser__meta-orcidImage" href="https://orcid.org/0000-0001-9428-7337"><img src="https://140.82.21.59/plugins/themes/oldGregg/templates/images/orcid.png"></a></li> </ul> <div class="jatsParser__details-authors"> <div class="jatsParser__details-author jatsParser__hideAuthor" id="jatsParser__author-1"> <div class="jatsParser__details-author-affiliation">University of Delaware; Department of Entomology and Wildlife Ecology; 250 Townsend Hall; Newark; DE 19716-2160; USA.</div> </div> <div class="jatsParser__details-author jatsParser__hideAuthor" id="jatsParser__author-2"> <div class="jatsParser__details-author-affiliation">University of Delaware; Department of Entomology and Wildlife Ecology; 250 Townsend Hall; Newark; DE 19716-2160; USA.</div> </div> </div> <div class="jatsParser__keywords-wrapper"> <div class="jatsParser__keywords-row"> <span class="jatsParser__keyword">Hemiptera</span> <span class="jatsParser__keyword">Flatoidini</span> <span class="jatsParser__keyword">Caribbean</span> <span class="jatsParser__keyword">new synonymy</span> <span class="jatsParser__keyword">new combination</span> <span class="jatsParser__keyword">iNaturalist</span> </div> </div> </div> <div class="jatsParser__articleView"> <div class="jatsParser__left-article-block"> <div class="article-page__galleys"> <ul class="list-galleys primary-galleys"> <li> <a class="obj_galley_link pdf restricted" href="https://140.82.21.59/zt/article/view/zootaxa.5590.2.4/55693"> <span class="pkp_screen_reader"> Requires Subscription or Fee </span> PDF(40.28MB) </a> </li> </ul> </div> </div> <div class="jatsParser__center-article-block"> <div class="jatsParser__article-fulltext" id="jatsParserFullText"> <h2 class="article-section-title jatsParser__abstract">Abstract</h2> <p>The genera <em>Atracodes </em>Melichar, and <em>Pseudoflatoides </em>Metcalf, are small Antillean genera in the flatid subfamily Flatoidinae. We found <em>Pseudoflatoides </em>to be a junior subjective synonym of <em>Atracodes </em>resulting in the transfer of four species (and seven subspecies)<em>.</em> We revise the status of two subspecies of <em>Atracodes fasciculosus </em><strong>comb. nov.</strong> to species level (as <em>Atracodes fasciatus </em><strong>comb. nov. & stat. nov.</strong> and <em>Atracodes vittatus </em><strong>comb. nov. & stat. nov.</strong>) based on characteristics of the male terminalia. <em>Atracodes corrugatus </em>Melichar (from Madagascar) is transferred to <em>Lichenopsis </em>Schmidt as <em>Lichenopsis corrugata </em>(Melichar), <strong>comb</strong>. <strong>nov</strong>. The resulting composition of <em>Atracodes </em>is 10 species, and four subspecies. We also compile available information on the two genera, provide a key to the American genera of Flatoidinae, along with a key to species of <em>Atracodes</em>, and suggest future work on the subfamily. </p> <p> </p> <h2 class="article-section-title jatsParser__references">References</h2> <ol class="jatsParser__references-list"> <li>Amyot, C.J.-B. & Audinet-Serville, J.G. (1843) Histoire naturelle des Insectes. Hémiptères. Librairie Encyclopédique de Roret, Paris, lxxvi + 676 pp., 12 pls. <a href="https://doi.org/10.5962/bhl.title.8471">https://doi.org/10.5962/bhl.title.8471</a> </li> <li>Arthropod Easy Capture (2013) Arthropod Easy Capture. Version 1.34. Available from: <a href="https://sourceforge.net/projects/arthropodeasy">https://sourceforge.net/projects/arthropodeasy</a> (accessed 7 January 2025) </li> <li>Bartlett, C.R. (2000) An annotated list of planthoppers (Hemiptera: Fulgoroidea) of Guana Island (British West Indies). 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A review of the genera Atracodes Melichar and Pseudoflatoides Metcalf (Hemiptera: Flatidae: Flatoidinae) from the Caribbean region | Zootaxa