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John Edward Sulston - Wikipedia
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class="firstHeading mw-first-heading"><span class="mw-page-title-main">John Edward Sulston</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Ga naar een artikel in een andere taal. Beschikbaar in 51 talen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-51" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">51 talen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AC%D9%88%D9%86_%D8%B3%D8%A7%D9%84%D8%B3%D8%AA%D9%88%D9%86" title="جون سالستون – Arabisch" lang="ar" hreflang="ar" data-title="جون سالستون" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%AC%D9%88%D9%86_%D8%B3%D8%A7%D9%84%D8%B3%D8%AA%D9%88%D9%86" title="جون سالستون – Egyptisch Arabisch" lang="arz" hreflang="arz" data-title="جون سالستون" data-language-autonym="مصرى" data-language-local-name="Egyptisch Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%AC%D8%A7%D9%86_%D8%B3%D8%A7%D9%84%D8%B3%D8%AA%D9%86" title="جان سالستن – South Azerbaijani" lang="azb" hreflang="azb" data-title="جان سالستن" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D0%BD_%D0%A1%D0%B0%D0%BB%D1%81%D1%82%D0%B0%D0%BD" title="Джон Салстан – Belarussisch" lang="be" hreflang="be" data-title="Джон Салстан" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D0%BD_%D0%A1%D1%8A%D0%BB%D1%81%D1%82%D1%8A%D0%BD" title="Джон Сълстън – Bulgaars" lang="bg" hreflang="bg" data-title="Джон Сълстън" data-language-autonym="Български" data-language-local-name="Bulgaars" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%9C%E0%A6%A8_%E0%A6%B8%E0%A6%BE%E0%A6%B2%E0%A6%B8%E0%A7%8D%E0%A6%9F%E0%A6%A8" title="জন সালস্টন – Bengaals" lang="bn" hreflang="bn" data-title="জন সালস্টন" data-language-autonym="বাংলা" data-language-local-name="Bengaals" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/John_Edward_Sulston" title="John Edward Sulston – Catalaans" lang="ca" hreflang="ca" data-title="John Edward Sulston" data-language-autonym="Català" data-language-local-name="Catalaans" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/John_Sulston" title="John Sulston – Tsjechisch" lang="cs" hreflang="cs" data-title="John Sulston" data-language-autonym="Čeština" data-language-local-name="Tsjechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Duits" lang="de" hreflang="de" data-title="John E. Sulston" data-language-autonym="Deutsch" data-language-local-name="Duits" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/John_Sulston" title="John Sulston – Engels" lang="en" hreflang="en" data-title="John Sulston" data-language-autonym="English" data-language-local-name="Engels" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/John_Sulston" title="John Sulston – Spaans" lang="es" hreflang="es" data-title="John Sulston" data-language-autonym="Español" data-language-local-name="Spaans" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/John_Sulston" title="John Sulston – Estisch" lang="et" hreflang="et" data-title="John Sulston" data-language-autonym="Eesti" data-language-local-name="Estisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Baskisch" lang="eu" hreflang="eu" data-title="John E. Sulston" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%AC%D8%A7%D9%86_%D8%B3%D8%A7%D9%84%D8%B3%D8%AA%D9%86" title="جان سالستن – Perzisch" lang="fa" hreflang="fa" data-title="جان سالستن" data-language-autonym="فارسی" data-language-local-name="Perzisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/John_Sulston" title="John Sulston – Fins" lang="fi" hreflang="fi" data-title="John Sulston" data-language-autonym="Suomi" data-language-local-name="Fins" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/John_Sulston" title="John Sulston – Frans" lang="fr" hreflang="fr" data-title="John Sulston" data-language-autonym="Français" data-language-local-name="Frans" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/John_Sulston" title="John Sulston – Iers" lang="ga" hreflang="ga" data-title="John Sulston" data-language-autonym="Gaeilge" data-language-local-name="Iers" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gd mw-list-item"><a href="https://gd.wikipedia.org/wiki/John_Sulston" title="John Sulston – Schots-Gaelisch" lang="gd" hreflang="gd" data-title="John Sulston" data-language-autonym="Gàidhlig" data-language-local-name="Schots-Gaelisch" class="interlanguage-link-target"><span>Gàidhlig</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%92%27%D7%95%D7%9F_%D7%A1%D7%9C%D7%A1%D7%98%D7%95%D7%9F" title="ג'ון סלסטון – Hebreeuws" lang="he" hreflang="he" data-title="ג'ון סלסטון" data-language-autonym="עברית" data-language-local-name="Hebreeuws" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%9C%E0%A5%89%E0%A4%A8_%E0%A4%B8%E0%A4%B2%E0%A5%8D%E0%A4%B8%E0%A4%9F%E0%A4%A8" title="जॉन सल्सटन – Hindi" lang="hi" hreflang="hi" data-title="जॉन सल्सटन" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Kroatisch" lang="hr" hreflang="hr" data-title="John E. Sulston" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Hongaars" lang="hu" hreflang="hu" data-title="John E. Sulston" data-language-autonym="Magyar" data-language-local-name="Hongaars" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/John_Edward_Sulston" title="John Edward Sulston – Indonesisch" lang="id" hreflang="id" data-title="John Edward Sulston" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/John_Edward_Sulston" title="John Edward Sulston – Ido" lang="io" hreflang="io" data-title="John Edward Sulston" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/John_Sulston" title="John Sulston – Italiaans" lang="it" hreflang="it" data-title="John Sulston" data-language-autonym="Italiano" data-language-local-name="Italiaans" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B8%E3%83%A7%E3%83%B3%E3%83%BB%E3%82%B5%E3%83%AB%E3%82%B9%E3%83%88%E3%83%B3" title="ジョン・サルストン – Japans" lang="ja" hreflang="ja" data-title="ジョン・サルストン" data-language-autonym="日本語" data-language-local-name="Japans" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D0%BD_%D0%A1%D0%B0%D0%BB%D1%81%D1%82%D0%BE%D0%BD" title="Джон Салстон – Kazachs" lang="kk" hreflang="kk" data-title="Джон Салстон" data-language-autonym="Қазақша" data-language-local-name="Kazachs" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%A1%B4_%EC%84%A4%EC%8A%A4%ED%84%B4" title="존 설스턴 – Koreaans" lang="ko" hreflang="ko" data-title="존 설스턴" data-language-autonym="한국어" data-language-local-name="Koreaans" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Ioannes_Eduardus_Sulston" title="Ioannes Eduardus Sulston – Latijn" lang="la" hreflang="la" data-title="Ioannes Eduardus Sulston" data-language-autonym="Latina" data-language-local-name="Latijn" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/John_Sulston" title="John Sulston – Litouws" lang="lt" hreflang="lt" data-title="John Sulston" data-language-autonym="Lietuvių" data-language-local-name="Litouws" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Maleis" lang="ms" hreflang="ms" data-title="John E. Sulston" data-language-autonym="Bahasa Melayu" data-language-local-name="Maleis" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%AC%D8%A7%D9%86_%D8%B3%D8%A7%D9%84%D8%B3%D8%AA%D9%86" title="جان سالستن – Mazanderani" lang="mzn" hreflang="mzn" data-title="جان سالستن" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/John_Edward_Sulston" title="John Edward Sulston – Nedersaksisch" lang="nds" hreflang="nds" data-title="John Edward Sulston" data-language-autonym="Plattdüütsch" data-language-local-name="Nedersaksisch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Noors - Bokmål" lang="nb" hreflang="nb" data-title="John E. Sulston" data-language-autonym="Norsk bokmål" data-language-local-name="Noors - Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Occitaans" lang="oc" hreflang="oc" data-title="John E. Sulston" data-language-autonym="Occitan" data-language-local-name="Occitaans" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/John_Sulston" title="John Sulston – Pools" lang="pl" hreflang="pl" data-title="John Sulston" data-language-autonym="Polski" data-language-local-name="Pools" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%AC%D8%A7%D9%86_%D8%B3%D9%84%D8%B3%D8%AA%D9%86" title="جان سلستن – Western Punjabi" lang="pnb" hreflang="pnb" data-title="جان سلستن" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/John_Sulston" title="John Sulston – Portugees" lang="pt" hreflang="pt" data-title="John Sulston" data-language-autonym="Português" data-language-local-name="Portugees" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A1%D0%B0%D0%BB%D1%81%D1%82%D0%BE%D0%BD,_%D0%94%D0%B6%D0%BE%D0%BD" title="Салстон, Джон – Russisch" lang="ru" hreflang="ru" data-title="Салстон, Джон" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/John_Sulston" title="John Sulston – Simple English" lang="en-simple" hreflang="en-simple" data-title="John Sulston" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/John_Sulston" title="John Sulston – Slowaaks" lang="sk" hreflang="sk" data-title="John Sulston" data-language-autonym="Slovenčina" data-language-local-name="Slowaaks" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Zweeds" lang="sv" hreflang="sv" data-title="John E. Sulston" data-language-autonym="Svenska" data-language-local-name="Zweeds" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/John_Sulston" title="John Sulston – Swahili" lang="sw" hreflang="sw" data-title="John Sulston" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Turks" lang="tr" hreflang="tr" data-title="John E. Sulston" data-language-autonym="Türkçe" data-language-local-name="Turks" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D0%BD_%D0%A1%D0%B0%D0%BB%D1%81%D1%82%D0%BE%D0%BD" title="Джон Салстон – Oekraïens" lang="uk" hreflang="uk" data-title="Джон Салстон" data-language-autonym="Українська" data-language-local-name="Oekraïens" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%AC%D8%A7%D9%86_%D8%B3%D9%84%D8%B3%D9%B9%D9%86" title="جان سلسٹن – Urdu" lang="ur" hreflang="ur" data-title="جان سلسٹن" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/John_Sulston" title="John Sulston – Vietnamees" lang="vi" hreflang="vi" data-title="John Sulston" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamees" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Yoruba" lang="yo" hreflang="yo" data-title="John E. Sulston" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E7%BA%A6%E7%BF%B0%C2%B7%E8%8B%8F%E5%B0%94%E6%96%AF%E9%A1%BF" title="约翰·苏尔斯顿 – Chinees" lang="zh" hreflang="zh" data-title="约翰·苏尔斯顿" data-language-autonym="中文" data-language-local-name="Chinees" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/John_E._Sulston" title="John E. Sulston – Minnanyu" lang="nan" hreflang="nan" data-title="John E. Sulston" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnanyu" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E8%98%87%E7%88%BE%E6%96%AF%E9%A0%93" title="蘇爾斯頓 – Kantonees" lang="yue" hreflang="yue" data-title="蘇爾斯頓" data-language-autonym="粵語" data-language-local-name="Kantonees" class="interlanguage-link-target"><span>粵語</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q271532#sitelinks-wikipedia" title="Taalkoppelingen bewerken" class="wbc-editpage">Koppelingen bewerken</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Naamruimten"> <div id="p-associated-pages" class="vector-menu 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font-size:120%;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Nobel_prize_medal.svg" class="mw-file-description" title="Nobelprijswinnaar"><img alt="Nobelprijswinnaar" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/25px-Nobel_prize_medal.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/38px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/50px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span>  John Edward Sulston </th></tr> <tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center;" colspan="3">27 maart 1942 – 6 maart 2018 </th></tr> <tr> <td style="text-align:center; padding:0;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:John_Sulston.jpg" class="mw-file-description"><img alt="John Edward Sulston" src="//upload.wikimedia.org/wikipedia/commons/thumb/b/bc/John_Sulston.jpg/266px-John_Sulston.jpg" decoding="async" width="266" height="261" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/bc/John_Sulston.jpg/399px-John_Sulston.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/bc/John_Sulston.jpg/532px-John_Sulston.jpg 2x" data-file-width="1102" data-file-height="1083" /></a></span> </td></tr> <tr> <th>Geboorteland </th> <td colspan="2"><a href="/wiki/Verenigd_Koninkrijk" title="Verenigd Koninkrijk">Verenigd Koninkrijk</a> </td></tr> <tr> <th>Geboorteplaats </th> <td colspan="2"><a href="/wiki/Cambridge_(Verenigd_Koninkrijk)" title="Cambridge (Verenigd Koninkrijk)">Cambridge</a> </td></tr> <tr> <th>Overlijdensplaats </th> <td colspan="2"><a href="/wiki/Stapleford_(Cambridgeshire)" title="Stapleford (Cambridgeshire)">Stapleford (Cambridgeshire)</a> </td></tr> <tr> <th>Nobelprijs </th> <td colspan="2">Fysiologie of Geneeskunde </td></tr> <tr> <th>Jaar </th> <td colspan="2">2002 </td></tr> <tr> <th>Reden </th> <td colspan="2">Voor hun ontdekkingen betreffende de genetische regulatie van de orgaanontwikkeling en de <a href="/wiki/Apoptose" title="Apoptose">geprogrammeerde celdood</a> </td></tr> <tr> <th>Samen met </th> <td colspan="2"><a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a><br /><a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> </td></tr> <tr> <th><span class="nowrap">Voorganger(s)</span> </th> <td colspan="2"><a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Leland H. Hartwell</a><br /><a href="/wiki/Tim_Hunt" title="Tim Hunt">Tim Hunt</a><br /><a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a> </td></tr> <tr> <th>Opvolger(s) </th> <td colspan="2"><a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a><br /><a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a> </td></tr> <tr> <td class="infobox-kop notheme center" style="background-color:lightsteelblue; color:inherit;" colspan="3"> <table cellspacing="0" cellpadding="0"> <tbody><tr> <td style="vertical-align:middle;"><b>Portaal</b>  <span class="noviewer" typeof="mw:File"><a href="/wiki/Bestand:Portal.svg" class="mw-file-description" title="Portaalicoon"><img alt="Portaalicoon" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/22px-Portal.svg.png" decoding="async" width="22" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/33px-Portal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/44px-Portal.svg.png 2x" data-file-width="36" data-file-height="32" /></a></span>   </td> <td class="ta-left"><b><a href="/wiki/Portaal:Geneeskunde" title="Portaal:Geneeskunde">Geneeskunde</a> </b> </td></tr></tbody></table> </td></tr> </tbody></table> <p><b>John Edward Sulston</b>, <a href="/wiki/Royal_Society" title="Royal Society">FRS</a> (<a href="/wiki/Cambridge_(Verenigd_Koninkrijk)" title="Cambridge (Verenigd Koninkrijk)">Cambridge</a>, <a href="/wiki/27_maart" title="27 maart">27 maart</a> <a href="/wiki/1942" title="1942">1942</a> – <a href="/wiki/Stapleford_(Cambridgeshire)" title="Stapleford (Cambridgeshire)">Stapleford</a>, <a href="/wiki/6_maart" title="6 maart">6 maart</a> <a href="/wiki/2018" title="2018">2018</a><sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>) was een Brits <a href="/wiki/Biologie" title="Biologie">bioloog</a>. In 2002 won hij samen met <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> en <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a> voor hun ontdekkingen betreffende de genetische regulatie van de orgaanontwikkeling en de <a href="/wiki/Apoptose" title="Apoptose">geprogrammeerde celdood</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Biografie">Biografie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=John_Edward_Sulston&veaction=edit&section=1" title="Bewerk dit kopje: Biografie" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=John_Edward_Sulston&action=edit&section=1" title="De broncode bewerken van de sectie: Biografie"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Sulston werd geboren in Cambridge als zoon van de Anglicaanse priester Theodore Sulston en de onderwijzeres Muriel Sulston.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Hij studeerde aan het <a href="/w/index.php?title=Pembroke_College_(Cambridge)&action=edit&redlink=1" class="new" title="Pembroke College (Cambridge) (de pagina bestaat niet)">Pembroke College</a>, alwaar hij in 1963 afstudeerde. Daarna ging hij naar het scheikundedepartement van de <a href="/wiki/Universiteit_van_Cambridge" title="Universiteit van Cambridge">Universiteit van Cambridge</a>, waar hij in 1966 zijn <a href="/wiki/Doctor_of_Philosophy" title="Doctor of Philosophy">Ph.D.</a> haalde met zijn onderzoek naar <a href="/wiki/Nucleotide" title="Nucleotide">nucleotiden</a>. Na zijn studie focuste hij zich geheel op biologisch onderzoek, vooral op het gebied van <a href="/wiki/Moleculaire_biologie" title="Moleculaire biologie">moleculaire biologie</a>. Eerst werkte hij als <a href="/wiki/Postdoc" title="Postdoc">postdoc</a> bij het <a href="/w/index.php?title=Salk_Institute&action=edit&redlink=1" class="new" title="Salk Institute (de pagina bestaat niet)">Salk Institute</a> in de Verenigde Staten, alvorens terug te keren naar Cambridge om te gaan werken onder <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> bij het MRC Laboratory of Molecular Biology. </p> <div class="mw-heading mw-heading2"><h2 id="Werk">Werk</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=John_Edward_Sulston&veaction=edit&section=2" title="Bewerk dit kopje: Werk" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=John_Edward_Sulston&action=edit&section=2" title="De broncode bewerken van de sectie: Werk"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Sulston speelde een centrale rol in zowel het onderzoek naar de rondworm <i><a href="/wiki/Caenorhabditis_elegans" title="Caenorhabditis elegans">C. elegans</a></i> als het <a href="/wiki/Menselijkgenoomproject" title="Menselijkgenoomproject">menselijk genoom</a> sequentieproject. Met behulp van een microscoop voorzien van speciale lenzen was hij de eerste wetenschapper die waarnam dat van 1090 cellen die door celdeling ontstaan tijdens de ontwikkeling van de rondworm uit één enkele cel er altijd 131 cellen reproduceerbaar geëlimineerd worden in een ordelijk, sequentieel proces. Uiteindelijk blijven in de volgroeide worm 959 cellen over. Dit afsterven, wat <a href="/wiki/Apoptose" title="Apoptose">apoptose</a> of 'geprogrammeerde celdood' wordt genoemd, is belangrijk voor de normale ontwikkeling van een organisme. </p><p>Hij zorgde voor de sequentie van <i>C. elegans</i> om aan te tonen dat sequentieprojecten op grote schaal mogelijk was. Tijdens het onderzoek werd hij in 1992 benoemd tot directeur van het Sanger Centre. In samenwerking met anderen heeft hij ongeveer een derde van het menselijk genoom de DNA-sequentie bepaald. Na het voltooien van de sequentie van het menselijk genoom, trok Sulston zich terug als directeur. Sindsdien zet hij zich actief in dat het werk rond het bepalen van de genetische blauwdruk publiekelijk en vrij beschikbaar blijft voor toekomstig onderzoek. </p><p>In 2007 werd bekend dat Sulston zou gaan werken bij de faculteit van Life Science aan de <a href="/wiki/Universiteit_van_Manchester" title="Universiteit van Manchester">Universiteit van Manchester</a>. Ook werd hij voorzitter van het Institute for Science, Ethics and Innovation, een nieuw onderzoeksinstituut dat zich focust op ethische vragen die worden opgeroepen door de wetenschap. </p><p>John Edward Sulston overleed in 2018 op 75-jarige leeftijd.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Externe_links">Externe links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=John_Edward_Sulston&veaction=edit&section=3" title="Bewerk dit kopje: Externe links" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=John_Edward_Sulston&action=edit&section=3" title="De broncode bewerken van de sectie: Externe links"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="http://biology.plosjournals.org/perlserv?request=get-document&doi=10.1371/journal.pbio.0040198">PLoS Biology: All for All</a></li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20130728042628/http://genome.wellcome.ac.uk/doc_wtd021047.html">John Sulston biography from the Wellcome Trust</a></li> <li><style data-mw-deduplicate="TemplateStyles:r67679320">.mw-parser-output .taalaanduiding{font-family:sans-serif;font-size:85%;cursor:help;color:var(--color-subtle,#555)}.mw-parser-output .taalaanduiding span{border-bottom:1px dotted var(--color-subtle,#555)}</style><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/2002/summary/">The Nobel Prize in Physiology or Medicine 2002</a>. NobelPrize.org. Nobel Media AB 2018.</li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20071121061007/http://www.npg.org.uk/live/search/person.asp?LinkID=mp59340">Portraits of John Sulston</a> from the <a href="/w/index.php?title=National_Portrait_Gallery_(London)&action=edit&redlink=1" class="new" title="National Portrait Gallery (London) (de pagina bestaat niet)">National Portrait Gallery (United Kingdom)</a></li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20120716181331/http://www.channel4.com/learning/microsites/G/genetics/lectures/about.html">John Sulston profile from Channel4</a></li> <li><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?db=pubmed&cmd=search&term=Sulston+S">John Sulston scientific publications</a> from <a href="/wiki/PubMed" title="PubMed">PubMed</a></li> <li><a rel="nofollow" class="external text" href="https://www.sanger.ac.uk/Info/Press/2002/021007.shtml">Sir John Sulston awarded the 2002 Nobel Prize (press release from the Sanger Centre)</a></li> <li><a rel="nofollow" class="external text" href="http://news.bbc.co.uk/2/hi/in_depth/sci_tech/2000/human_genome/753685.stm">The public servant: John Sulston</a></li></ul> <div class="toccolours appendix" role="presentation" style="font-size:90%; margin:1em 0 -0.5em; clear:both;"> <div><span style="font-weight:bold">Bronnen, noten en/of referenties</span></div> <ul><li><span class="citation" id="CITEREFC.D._Dijkstra2002"><span style="font-variant:small-caps;">C.D. Dijkstra</span> (2002). <cite>Nobelprijs Fysiologie of Geneeskunde 2002 voor onderzoek naar de genetische regulatie van orgaanontwikkeling en geprogrammeerde celdood</cite>. <i>Nederlands Tijdschrift voor Geneeskunde</i> <b>146</b> (52): 2525-2527. </span></li></ul> <div class="reflist" style="list-style-type: decimal;"><div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Nederlands">(<span>nl</span>) </span><span class="citation" id="CITEREFFout:_ongeldige_tijd."><a rel="nofollow" class="external text" href="https://www.telegraaf.nl/nieuws/1771033/genoompionier-john-sulston-overleden">Genoompionier John Sulston overleden</a>. De Telegraaf (<span class="mw-formatted-date" title="2018-03-09">9 maart 2018</span>). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20220628053020/https://www.telegraaf.nl/nieuws/1771033/genoompionier-john-sulston-overleden">Gearchiveerd</a> op <span class="mw-formatted-date" title="2022-06-28">28 juni 2022</span>. </span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> <a rel="nofollow" class="external text" href="http://www.nndb.com/people/426/000136018/">Biografie John E. Sulston</a> op Notable Names Database (NNDb).<small> Geraadpleegd op 21 juni 2015</small></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.sanger.ac.uk/news/view/professor-sir-john-sulston-1942-2018">John Sulston (1942-2018) – Founding Director of the Wellcome Sanger Institute</a>, Sanger Institute, 9 maart 2018.</span> </li> </ol></div></div> </div> <style data-mw-deduplicate="TemplateStyles:r67837862">.mw-parser-output .navigatie{position:relative;clear:both;overflow:auto;margin:1em auto -0.5em;padding:2px;background-color:var(--background-color-neutral-subtle,#f8f9fa);border:1px solid var(--border-color-base,#a2a9b1);text-align:center;font-size:87%}.mw-parser-output .navigatie-bewerken{margin-left:0.5em}.mw-parser-output .navigatie-bewerken .mw-ui-icon::before{background-size:0.9em}.mw-parser-output .navigatie-afb-links,.mw-parser-output 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href="/wiki/Bibliotheek_van_het_Japanse_parlement" title="Bibliotheek van het Japanse parlement">Bibliotheek van het Japanse parlement</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://id.ndl.go.jp/auth/ndlna/00924845">00924845</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Tsjechi%C3%AB" title="Nationale Bibliotheek van Tsjechië">Nationale Bibliotheek van Tsjechië</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=xx0076957&CON_LNG=ENG">xx0076957</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Isra%C3%ABl" title="Nationale Bibliotheek van Israël">Nationale Bibliotheek van Israël</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://olduli.nli.org.il/F/?func=find-b&local_base=NLX10&find_code=UID&request=987007268623305171">987007268623305171</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Polen" title="Nationale Bibliotheek van Polen">Nationale Bibliotheek van Polen</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://dbn.bn.org.pl/descriptor-details/a0000003021723">a0000003021723</a></span></li><li><b><a href="https://www.wikidata.org/wiki/Property:P1006" class="extiw" title="d:Property:P1006">Nederlandse Thesaurus van Auteursnamen</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p239988531">239988531</a></span></li><li><b><a href="/wiki/LIBRIS" title="LIBRIS">LIBRIS</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://libris.kb.se/auth/240045">240045</a></span></li><li><b><a href="/wiki/Syst%C3%A8me_universitaire_de_documentation" title="Système universitaire de documentation">Système universitaire de documentation</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/072520787">072520787</a></span></li><li><b><a href="/wiki/Virtual_International_Authority_File" title="Virtual International Authority File">Virtual International Authority File</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/18364938">18364938</a></span></li><li><b></b> <b><a href="/wiki/WorldCat" title="WorldCat">WorldCat</a></b> (via VIAF): <a rel="nofollow" class="external text" href="https://www.worldcat.org/identities/containsVIAFID/18364938">18364938</a></li></ul></div></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67837862"> <div class="navigatie mw-collapsible mw-collapsed" role="navigation" aria-labelledby="Winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" data-expandtext="Uitklappen" data-collapsetext="Inklappen"> <div class="navigatie-afb-links noviewer"><div class="navigatie-bewerken show-autoconfirmed nomobile"><span class="nowrap skin-invert-image"><span typeof="mw:File"><a href="/wiki/Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Sjabloon:Navigatie winnaars van de Nobelprijs voor Fysiologie of Geneeskunde"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/16px-OOjs_UI_icon_article-ltr.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/24px-OOjs_UI_icon_article-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/32px-OOjs_UI_icon_article-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> · <span class="ui-state-disabled" typeof="mw:File"><span title="Overleg sjabloon (de pagina bestaat niet)"><img alt="Overleg sjabloon (de pagina bestaat niet)" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/40/OOjs_UI_icon_speechBubbles-ltr.svg/16px-OOjs_UI_icon_speechBubbles-ltr.svg.png" decoding="async" 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title="Santiago Ramón y Cajal">Ramón y Cajal</a> · 1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Laveran</a> · 1908: <a href="/wiki/Ilja_Iljitsj_Metsjnikov" title="Ilja Iljitsj Metsjnikov">Mechnikov</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> · 1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> · 1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> · 1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> · 1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> · 1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> · 1914: <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> · 1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> · 1920: <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> · 1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a>, <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> · 1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_Macleod_(medicus)" title="John Macleod (medicus)">Macleod</a> · 1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> · 1926: <a href="/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger">Fibiger</a> · 1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> · 1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> · 1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> · 1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> · 1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> · 1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Adrian</a> · 1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> · 1934: <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a>, <a href="/wiki/George_Minot" title="George Minot">Minot</a>, <a href="/wiki/William_Murphy" title="William Murphy">Murphy</a> · 1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> · 1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> · 1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> · 1938: <a href="/wiki/Corneel_Heymans" title="Corneel Heymans">Heymans</a> · 1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> · 1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> · 1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> · 1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> · 1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> · 1947: <span class="nowrap"><a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a></span>, <span class="nowrap"><a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a></span>, <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> · 1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> · 1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> · 1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> · 1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> · 1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> · 1953: <a href="/wiki/Hans_Krebs_(arts)" title="Hans Krebs (arts)">Krebs</a>,<a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> · 1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> · 1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> · 1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> · 1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> · 1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> · 1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> · 1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> · 1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> · 1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> · 1963: <a href="/wiki/John_Eccles" title="John Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> · 1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a>, <a href="/wiki/Feodor_Felix_Konrad_Lynen" title="Feodor Felix Konrad Lynen">Lynen</a> · 1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Michael_Lwoff" title="André Michael Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> · 1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> · 1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> · 1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> · 1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a>, <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a>, <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> · 1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_Svante_von_Euler" title="Ulf Svante von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> · 1971: <a href="/wiki/Earl_Wilbur_Sutherland_jr." title="Earl Wilbur Sutherland jr.">Sutherland</a> · 1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a>, <a href="/wiki/Rodney_Porter" title="Rodney Porter">Porter</a> · 1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> · 1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">De Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> · 1975: <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Temin</a> · 1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> · 1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> · 1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a href="/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith">Smith</a> · 1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> · 1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> · 1981: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> · 1982: <a href="/wiki/Sune_Karl_Bergstr%C3%B6m" title="Sune Karl Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> · 1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> · 1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a href="/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> · 1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Goldstein</a> · 1986: <a href="/wiki/Stanley_Cohen_(biochemicus)" title="Stanley Cohen (biochemicus)">Cohen</a>, <span class="nowrap"><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a></span> · 1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> · 1988: <a href="/wiki/James_Whyte_Black" title="James Whyte Black">Black</a>, <a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Elion</a>, <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> · 1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> · 1990: <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Murray</a>, <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">Thomas</a> · 1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> · 1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a>, <a href="/wiki/Edwin_Gerhard_Krebs" title="Edwin Gerhard Krebs">Krebs</a> · 1993: <a href="/wiki/Richard_John_Roberts" title="Richard John Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> · 1994: <a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> · 1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <span class="nowrap"><a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a></span>, <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> · 1996: <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a>, <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> · 1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> · 1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a>, <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> · 1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> · 2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> · 2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> · 2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a class="mw-selflink selflink">Sulston</a> · 2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> · 2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> · 2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> · 2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> · 2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> · 2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Zur Hausen</a>, <span class="nowrap"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré–Sinoussi</a>,</span> <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> · 2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a>, <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> · 2010: <a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Edwards</a> · 2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> · 2012: <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> · 2013: <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> · 2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">O'Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">Moser</a> · 2015: <a href="/wiki/William_C._Campbell_(wetenschapper)" title="William C. Campbell (wetenschapper)">Campbell</a>, <a href="/wiki/Satoshi_Omura" title="Satoshi Omura">Omura</a>, <a href="/wiki/Youyou_Tu" title="Youyou Tu">Tu</a> · 2016: <a href="/wiki/Yoshinori_Osumi" title="Yoshinori Osumi">Osumi</a> · 2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> · 2018: <a href="/wiki/James_Allison" title="James Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> · 2019: <a href="/wiki/William_Kaelin_jr." title="William Kaelin jr.">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> · 2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> · 2021: <a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> · 2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> · 2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> · 2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div></div> <!-- NewPP limit report Parsed by mw‐web.eqiad.main‐68db4b6789‐rsxdm Cached time: 20241126112201 Cache expiry: 2592000 Reduced expiry: false Complications: [] CPU time usage: 0.180 seconds Real time usage: 0.279 seconds Preprocessor visited node count: 1762/1000000 Post‐expand include size: 37992/2097152 bytes Template argument size: 13448/2097152 bytes Highest expansion depth: 14/100 Expensive parser function count: 1/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 5858/5000000 bytes Lua time usage: 0.028/10.000 seconds Lua memory usage: 959507/52428800 bytes Number of Wikibase entities loaded: 1/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 206.511 1 -total 51.60% 106.551 1 Sjabloon:Infobox_Nobelprijswinnaar 49.61% 102.455 1 Sjabloon:Infobox_generiek 28.69% 59.244 1 Sjabloon:Wikidata_afbeelding 18.35% 37.886 1 Sjabloon:Appendix 15.76% 32.553 1 Sjabloon:Bibliografische_informatie 12.11% 25.007 1 Sjabloon:References 9.05% 18.686 1 Sjabloon:PAGENAMEBASE 6.58% 13.582 2 Sjabloon:En 6.26% 12.926 1 Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_de_Fysiologie_of_Geneeskunde --> <!-- Saved in parser cache with key nlwiki:pcache:idhash:1492613-0!dateformat=default and timestamp 20241126112201 and revision id 68338837. 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