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id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Basculer la table des matières" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Basculer la table des matières</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Élément SECIS</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input 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Disponible en 9 langues." > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-9" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">9 langues</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B9%D9%86%D8%B5%D8%B1_SECIS" title="عنصر SECIS – arabe" lang="ar" hreflang="ar" data-title="عنصر SECIS" data-language-autonym="العربية" data-language-local-name="arabe" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Element_SECIS" title="Element SECIS – catalan" lang="ca" hreflang="ca" data-title="Element SECIS" data-language-autonym="Català" data-language-local-name="catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/SECIS-element" title="SECIS-element – danois" lang="da" hreflang="da" data-title="SECIS-element" data-language-autonym="Dansk" data-language-local-name="danois" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/SECIS_element" title="SECIS element – anglais" lang="en" hreflang="en" data-title="SECIS element" data-language-autonym="English" data-language-local-name="anglais" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Elemento_SECIS" title="Elemento SECIS – galicien" lang="gl" hreflang="gl" data-title="Elemento SECIS" data-language-autonym="Galego" data-language-local-name="galicien" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/SECIS-%D1%8D%D0%BB%D0%B5%D0%BC%D0%B5%D0%BD%D1%82" title="SECIS-элемент – russe" lang="ru" hreflang="ru" data-title="SECIS-элемент" data-language-autonym="Русский" data-language-local-name="russe" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/SECIS_element" title="SECIS element – slovaque" lang="sk" hreflang="sk" data-title="SECIS element" data-language-autonym="Slovenčina" data-language-local-name="slovaque" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/SECIS-%D0%B5%D0%BB%D0%B5%D0%BC%D0%B5%D0%BD%D1%82" title="SECIS-елемент – ukrainien" lang="uk" hreflang="uk" data-title="SECIS-елемент" data-language-autonym="Українська" data-language-local-name="ukrainien" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/SECIS%E5%85%83%E4%BB%B6" title="SECIS元件 – chinois" lang="zh" hreflang="zh" data-title="SECIS元件" data-language-autonym="中文" data-language-local-name="chinois" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q3130206#sitelinks-wikipedia" title="Modifier les liens interlangues" class="wbc-editpage">Modifier les liens</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div 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data-event-name="pinnable-header.vector-page-tools.pin">déplacer vers la barre latérale</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">masquer</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="Plus d’options" > <div class="vector-menu-heading"> Actions </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/%C3%89l%C3%A9ment_SECIS"><span>Lire</span></a></li><li id="ca-more-ve-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&veaction=edit" title="Modifier cette page [v]" accesskey="v"><span>Modifier</span></a></li><li id="ca-more-edit" class="collapsible vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&action=edit" title="Modifier le wikicode de cette page [e]" accesskey="e"><span>Modifier le code</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&action=history"><span>Voir l’historique</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> Général </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/Sp%C3%A9cial:Pages_li%C3%A9es/%C3%89l%C3%A9ment_SECIS" title="Liste des pages liées qui pointent sur celle-ci [j]" accesskey="j"><span>Pages liées</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Sp%C3%A9cial:Suivi_des_liens/%C3%89l%C3%A9ment_SECIS" rel="nofollow" title="Liste des modifications récentes des pages appelées par celle-ci [k]" accesskey="k"><span>Suivi des pages liées</span></a></li><li id="t-upload" class="mw-list-item"><a href="/wiki/Aide:Importer_un_fichier" title="Téléverser des fichiers [u]" accesskey="u"><span>Téléverser un fichier</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Sp%C3%A9cial:Pages_sp%C3%A9ciales" title="Liste de toutes les pages spéciales [q]" accesskey="q"><span>Pages spéciales</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&oldid=204310392" title="Adresse permanente de cette version de cette page"><span>Lien permanent</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&action=info" title="Davantage d’informations sur cette page"><span>Informations sur la page</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=Sp%C3%A9cial:Citer&page=%C3%89l%C3%A9ment_SECIS&id=204310392&wpFormIdentifier=titleform" title="Informations sur la manière de citer cette page"><span>Citer cette page</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=Sp%C3%A9cial:UrlShortener&url=https%3A%2F%2Ffr.wikipedia.org%2Fwiki%2F%25C3%2589l%25C3%25A9ment_SECIS"><span>Obtenir l'URL raccourcie</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=Sp%C3%A9cial:QrCode&url=https%3A%2F%2Ffr.wikipedia.org%2Fwiki%2F%25C3%2589l%25C3%25A9ment_SECIS"><span>Télécharger le code QR</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> Imprimer / exporter </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-create_a_book" class="mw-list-item"><a href="/w/index.php?title=Sp%C3%A9cial:Livre&bookcmd=book_creator&referer=%C3%89l%C3%A9ment+SECIS"><span>Créer un livre</span></a></li><li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=Sp%C3%A9cial:DownloadAsPdf&page=%C3%89l%C3%A9ment_SECIS&action=show-download-screen"><span>Télécharger comme PDF</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&printable=yes" title="Version imprimable de cette page [p]" accesskey="p"><span>Version imprimable</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> Dans d’autres projets </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q3130206" title="Lien vers l’élément dans le dépôt de données connecté [g]" accesskey="g"><span>Élément Wikidata</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Outils de la page"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Apparence"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Apparence</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">déplacer vers la barre latérale</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">masquer</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">Un article de Wikipédia, l'encyclopédie libre.</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="fr" dir="ltr"><figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/Fichier:RF00031.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4e/RF00031.jpg/200px-RF00031.jpg" decoding="async" width="200" height="200" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4e/RF00031.jpg/300px-RF00031.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4e/RF00031.jpg/400px-RF00031.jpg 2x" data-file-width="452" data-file-height="452" /></a><figcaption>Variabilité des <a href="/wiki/Base_nucl%C3%A9ique" class="mw-redirect" title="Base nucléique">bases nucléiques</a> sur l'élément SECIS.</figcaption></figure> <p>Un <b>élément SECIS</b>, de l'anglais <span class="lang-en" lang="en"><i><b>se</b>leno<b>c</b>ysteine <b>i</b>nsertion <b>s</b>tructure</i></span>, est un <a href="/wiki/%C3%89l%C3%A9ment_cis-r%C3%A9gulateur" title="Élément cis-régulateur">élément cis-régulateur</a> d'<a href="/wiki/Acide_ribonucl%C3%A9ique_messager" title="Acide ribonucléique messager">ARN messager</a> long d'une soixantaine de <a href="/wiki/Nucl%C3%A9otide" title="Nucléotide">nucléotides</a> caractérisé par sa forme en épingle à cheveux, ou « <a href="/wiki/Tige-boucle" title="Tige-boucle">tige-boucle</a> ». Cette structure tridimensionnelle particulière conduit les <a href="/wiki/Ribosome" title="Ribosome">ribosomes</a> à <a href="/wiki/Traduction_g%C3%A9n%C3%A9tique" title="Traduction génétique">traduire</a> le <a href="/wiki/Codon" title="Codon">codon</a> UGA par un <a href="/wiki/Acide_amin%C3%A9" title="Acide aminé">acide aminé</a>, la <a href="/wiki/S%C3%A9l%C3%A9nocyst%C3%A9ine" title="Sélénocystéine">sélénocystéine</a>, alors qu'il s'agit normalement du <a href="/wiki/Codon-stop" title="Codon-stop">codon-<small>STOP</small></a> <i>Opale</i> dans le <a href="/wiki/Code_g%C3%A9n%C3%A9tique" title="Code génétique">code génétique</a>. Les éléments SECIS sont donc indispensables à la <a href="/wiki/Biosynth%C3%A8se_des_prot%C3%A9ines" title="Biosynthèse des protéines">synthèse</a> des <a href="/wiki/S%C3%A9l%C3%A9noprot%C3%A9ine" title="Sélénoprotéine">sélénoprotéines</a>, essentiellement des <a href="/wiki/Enzyme" title="Enzyme">enzymes</a> de la famille des <a href="/wiki/Oxydo-r%C3%A9ductase" class="mw-redirect" title="Oxydo-réductase">oxydo-réductases</a>. </p><p>Chez les <a href="/wiki/Bact%C3%A9rie" title="Bactérie">bactéries</a>, les éléments SECIS apparaissent peu après les codons UGA dont ils altèrent la traduction. Chez les <a href="/wiki/Archaea" title="Archaea">archées</a> et chez les <a href="/wiki/Eucaryote" class="mw-redirect" title="Eucaryote">eucaryotes</a>, ils se trouvent dans la région 3'-terminale des ARN messagers et peuvent affecter le sens de plusieurs codons UGA. L'un de ces éléments SECIS se trouve dans la région 5'-terminale chez <i><a href="/wiki/Methanococcus" title="Methanococcus">Methanococcus</a></i>, du <a href="/wiki/Phylum" title="Phylum">phylum</a> des <i><a href="/wiki/Euryarchaeota" title="Euryarchaeota">Euryarchaeota</a></i><sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite_crochet">[</span>1<span class="cite_crochet">]</span></a></sup><sup class="reference cite_virgule">,</sup><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite_crochet">[</span>2<span class="cite_crochet">]</span></a></sup>. </p><p>La <a href="/wiki/Structure_secondaire" title="Structure secondaire">structure secondaire</a> particulière des éléments SECIS découle d'une séquence de <a href="/wiki/Base_nucl%C3%A9ique" class="mw-redirect" title="Base nucléique">bases nucléiques</a> présentant certaines caractéristiques distinctives autorisant cette configuration spatiale déterminante, notamment des appariements atypiques entre paires de <a href="/wiki/Base_purique" class="mw-redirect" title="Base purique">bases puriques</a> — <a href="/wiki/Guanine" title="Guanine">guanine</a> et <a href="/wiki/Ad%C3%A9nine" title="Adénine">adénine</a> — qui sont essentiels au bon fonctionnement de l'élément SECIS<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite_crochet">[</span>3<span class="cite_crochet">]</span></a></sup>. </p> <div class="mw-heading mw-heading2"><h2 id="Articles_connexes">Articles connexes</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&veaction=edit&section=1" title="Modifier la section : Articles connexes" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&action=edit&section=1" title="Modifier le code source de la section : Articles connexes"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/%C3%89l%C3%A9ment_PYLIS" title="Élément PYLIS">Élément PYLIS</a>, relatif à la <a href="/wiki/Pyrrolysine" title="Pyrrolysine">pyrrolysine</a>.</li> <li><a href="/wiki/SECISBP2" title="SECISBP2">SECISBP2</a>, une protéine qui se fixe sur le SECIS</li></ul> <div class="mw-heading mw-heading2"><h2 id="Notes_et_références"><span id="Notes_et_r.C3.A9f.C3.A9rences"></span>Notes et références</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&veaction=edit&section=2" title="Modifier la section : Notes et références" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&action=edit&section=2" title="Modifier le code source de la section : Notes et références"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="references-small decimal" style=""><div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink noprint"><a href="#cite_ref-1">↑</a> </span><span class="reference-text"><span class="ouvrage" id="RotherReschWiltingBöck2001"><span class="ouvrage" id="M._RotherA._ReschR._WiltingA._Böck2001">M. <span class="nom_auteur">Rother</span>, A. <span class="nom_auteur">Resch</span>, R. <span class="nom_auteur">Wilting</span> et A. <span class="nom_auteur">Böck</span>, « <cite style="font-style:normal">Selenoprotein synthesis in archaea</cite> », <i>BioFactors (Oxford, England)</i>, <abbr class="abbr" title="volume">vol.</abbr> 14, <abbr class="abbr" title="numéros">n<sup>os</sup></abbr> 1-4,‎ <time>2001</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">75–83</span> <small style="line-height:1em;">(<a href="/wiki/International_Standard_Serial_Number" title="International Standard Serial Number">ISSN</a> <span class="plainlinks noarchive"><a rel="nofollow" class="external text" href="https://portal.issn.org/resource/issn/0951-6433">0951-6433</a></span>, <a href="/wiki/PubMed" title="PubMed">PMID</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/11568443">11568443</a></span>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/11568443">lire en ligne</a>, consulté le <time class="nowrap" datetime="2018-04-12" data-sort-value="2018-04-12">12 avril 2018</time>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Selenoprotein+synthesis+in+archaea&rft.jtitle=BioFactors+%28Oxford%2C+England%29&rft.issue=1-4&rft.aulast=Rother&rft.aufirst=M.&rft.au=Resch%2C+A.&rft.au=Wilting%2C+R.&rft.au=B%C3%B6ck%2C+A.&rft.date=2001&rft.volume=14&rft.pages=75%E2%80%9383&rft.issn=0951-6433&rft_id=info%3Apmid%2F11568443&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink noprint"><a href="#cite_ref-2">↑</a> </span><span class="reference-text"><span class="ouvrage" id="WiltingSchorlingPerssonBöck1997"><span class="ouvrage" id="R._WiltingS._SchorlingB._C._PerssonA._Böck1997">R. <span class="nom_auteur">Wilting</span>, S. <span class="nom_auteur">Schorling</span>, B. C. <span class="nom_auteur">Persson</span> et A. <span class="nom_auteur">Böck</span>, « <cite style="font-style:normal">Selenoprotein synthesis in archaea: identification of an mRNA element of Methanococcus jannaschii probably directing selenocysteine insertion</cite> », <i>Journal of Molecular Biology</i>, <abbr class="abbr" title="volume">vol.</abbr> 266, <abbr class="abbr" title="numéro">n<sup>o</sup></abbr> 4,‎ <time class="nowrap" datetime="1997-03-07" data-sort-value="1997-03-07">7 mars 1997</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">637–641</span> <small style="line-height:1em;">(<a href="/wiki/International_Standard_Serial_Number" title="International Standard Serial Number">ISSN</a> <span class="plainlinks noarchive"><a rel="nofollow" class="external text" href="https://portal.issn.org/resource/issn/0022-2836">0022-2836</a></span>, <a href="/wiki/PubMed" title="PubMed">PMID</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/9102456">9102456</a></span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">DOI</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://dx.doi.org/10.1006/jmbi.1996.0812">10.1006/jmbi.1996.0812</a></span>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/9102456">lire en ligne</a>, consulté le <time class="nowrap" datetime="2018-04-12" data-sort-value="2018-04-12">12 avril 2018</time>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Selenoprotein+synthesis+in+archaea%3A+identification+of+an+mRNA+element+of+Methanococcus+jannaschii+probably+directing+selenocysteine+insertion&rft.jtitle=Journal+of+Molecular+Biology&rft.issue=4&rft.aulast=Wilting&rft.aufirst=R.&rft.au=Schorling%2C+S.&rft.au=Persson%2C+B.+C.&rft.au=B%C3%B6ck%2C+A.&rft.date=1997-03-07&rft.volume=266&rft.pages=637%E2%80%93641&rft.issn=0022-2836&rft_id=info%3Adoi%2F10.1006%2Fjmbi.1996.0812&rft_id=info%3Apmid%2F9102456&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink noprint"><a href="#cite_ref-3">↑</a> </span><span class="reference-text"><span class="ouvrage" id="LambertLescureGautheret2002"><span class="ouvrage" id="André_LambertAlain_LescureDaniel_Gautheret2002">André <span class="nom_auteur">Lambert</span>, Alain <span class="nom_auteur">Lescure</span> et Daniel <span class="nom_auteur">Gautheret</span>, « <cite style="font-style:normal">A survey of metazoan selenocysteine insertion sequences</cite> », <i>Biochimie</i>, <abbr class="abbr" title="volume">vol.</abbr> 84, <abbr class="abbr" title="numéro">n<sup>o</sup></abbr> 9,‎ <time class="nowrap" datetime="2002-09" data-sort-value="2002-09">septembre 2002</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">953–959</span> <small style="line-height:1em;">(<a href="/wiki/International_Standard_Serial_Number" title="International Standard Serial Number">ISSN</a> <span class="plainlinks noarchive"><a rel="nofollow" class="external text" href="https://portal.issn.org/resource/issn/0300-9084">0300-9084</a></span>, <a href="/wiki/PubMed" title="PubMed">PMID</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/12458087">12458087</a></span>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/12458087">lire en ligne</a>, consulté le <time class="nowrap" datetime="2018-04-12" data-sort-value="2018-04-12">12 avril 2018</time>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=A+survey+of+metazoan+selenocysteine+insertion+sequences&rft.jtitle=Biochimie&rft.issue=9&rft.aulast=Lambert&rft.aufirst=Andr%C3%A9&rft.au=Lescure%2C+Alain&rft.au=Gautheret%2C+Daniel&rft.date=2002-09&rft.volume=84&rft.pages=953%E2%80%93959&rft.issn=0300-9084&rft_id=info%3Apmid%2F12458087&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></span></span> </li> </ol></div> </div> <div class="mw-heading mw-heading2"><h2 id="Liens_externes">Liens externes</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&veaction=edit&section=3" title="Modifier la section : Liens externes" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=%C3%89l%C3%A9ment_SECIS&action=edit&section=3" title="Modifier le code source de la section : Liens externes"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <div style="column-width:24em;column-gap:1em;" class="colonnes"> <ul><li><span class="ouvrage" id="R_Walczak,_E_Westhof,_P_Carbon_et_A_Krol1996"><abbr class="abbr indicateur-langue" title="Langue : anglais">(en)</abbr> <span class="nom_auteur">R Walczak, E Westhof, P Carbon et A Krol</span>, « <cite style="font-style:normal" lang="en">A novel RNA structural motif in the selenocysteine insertion element of eukaryotic selenoprotein mRNAs</cite> », <i><span class="lang-en" lang="en">RNA</span></i>, <abbr class="abbr" title="volume">vol.</abbr> 4,‎ <time class="nowrap" datetime="1996-04-02" data-sort-value="1996-04-02">2 avril 1996</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">367-379</span> <small style="line-height:1em;">(<a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1369379/?tool=pmcentrez">lire en ligne</a>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=A+novel+RNA+structural+motif+in+the+selenocysteine+insertion+element+of+eukaryotic+selenoprotein+mRNAs&rft.jtitle=RNA&rft.aulast=R+Walczak%2C+E+Westhof%2C+P+Carbon+et+A+Krol&rft.date=1996-04-02&rft.volume=4&rft.pages=367-379&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></li> <li><span class="ouvrage" id="André_Lambert,_Alain_Lescure_et_Daniel_Gautheret2002"><abbr class="abbr indicateur-langue" title="Langue : anglais">(en)</abbr> <span class="nom_auteur">André Lambert, Alain Lescure et Daniel Gautheret</span>, « <cite style="font-style:normal" lang="en">A survey of metazoan selenocysteine insertion sequences</cite> », <i><span class="lang-en" lang="en">Biochimie</span></i>, <abbr class="abbr" title="volume">vol.</abbr> 84, <abbr class="abbr" title="numéro">n<sup>o</sup></abbr> 9,‎ <time class="nowrap" datetime="2002-09" data-sort-value="2002-09">septembre 2002</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">953-959</span> <small style="line-height:1em;">(<a href="/wiki/PubMed" title="PubMed">PMID</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/12458087">12458087</a></span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">DOI</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://dx.doi.org/10.1016/S0300-9084%2802%2901441-4">10.1016/S0300-9084(02)01441-4</a></span>, <a rel="nofollow" class="external text" href="http://www.sciencedirect.com/science?_ob=ArticleURL&_udi=B6VRJ-46JYF9F-4&_user=10&_coverDate=09%2F30%2F2002&_rdoc=1&_fmt=high&_orig=search&_origin=search&_sort=d&_docanchor=&view=c&_acct=C000050221&_version=1&_urlVersion=0&_userid=10&md5=8bd5266c930970de6cb7911635ba33cf&searchtype=a">lire en ligne</a>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=A+survey+of+metazoan+selenocysteine+insertion+sequences&rft.jtitle=Biochimie&rft.issue=9&rft.aulast=Andr%C3%A9+Lambert%2C+Alain+Lescure+et+Daniel+Gautheret&rft.date=2002-09&rft.volume=84&rft.pages=953-959&rft_id=info%3Adoi%2F10.1016%2FS0300-9084%2802%2901441-4&rft_id=info%3Apmid%2F12458087&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></li> <li><span class="ouvrage" id="Heiko_Mix,_Alexey_V._Lobanov_et_Vadim_N._Gladyshev2006"><abbr class="abbr indicateur-langue" title="Langue : anglais">(en)</abbr> <span class="nom_auteur">Heiko Mix, Alexey V. Lobanov et Vadim N. Gladyshev</span>, « <cite style="font-style:normal" lang="en">SECIS elements in the coding regions of selenoprotein transcripts are functional in higher eukaryotes</cite> », <i><span class="lang-en" lang="en">Nucleic Acids Research</span></i>, <abbr class="abbr" title="volume">vol.</abbr> 35, <abbr class="abbr" title="numéro">n<sup>o</sup></abbr> 2,‎ <time class="nowrap" datetime="2006-12-14" data-sort-value="2006-12-14">14 décembre 2006</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">414-423</span> <small style="line-height:1em;">(<a href="/wiki/PubMed" title="PubMed">PMID</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/17169995">17169995</a></span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">DOI</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://dx.doi.org/10.1093/nar/gkl1060">10.1093/nar/gkl1060</a></span>, <a rel="nofollow" class="external text" href="http://nar.oxfordjournals.org/content/35/2/414">lire en ligne</a>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=SECIS+elements+in+the+coding+regions+of+selenoprotein+transcripts+are+functional+in+higher+eukaryotes&rft.jtitle=Nucleic+Acids+Research&rft.issue=2&rft.aulast=Heiko+Mix%2C+Alexey+V.+Lobanov+et+Vadim+N.+Gladyshev&rft.date=2006-12-14&rft.volume=35&rft.pages=414-423&rft_id=info%3Adoi%2F10.1093%2Fnar%2Fgkl1060&rft_id=info%3Apmid%2F17169995&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></li> <li><span class="ouvrage" id="A._Cassago,_E.M._Rodrigues,_E.L._Prieto,_K.W._Gaston,_J.D._Alfonzo_M.P._Iribar,_M.J._Berry,_A.K._Cruz_et_O.H._Thiemann2006"><abbr class="abbr indicateur-langue" title="Langue : anglais">(en)</abbr> <span class="nom_auteur">A. Cassago, E.M. Rodrigues, E.L. Prieto, K.W. Gaston, J.D. Alfonzo M.P. Iribar, M.J. Berry, A.K. Cruz et O.H. Thiemann</span>, « <cite style="font-style:normal" lang="en">Identification of <i>Leishmania</i> selenoproteins and SECIS element</cite> », <i><span class="lang-en" lang="en">Molecular and Biochemical Parasitology</span></i>, <abbr class="abbr" title="volume">vol.</abbr> 149, <abbr class="abbr" title="numéro">n<sup>o</sup></abbr> 2,‎ <time class="nowrap" datetime="2006-10" data-sort-value="2006-10">octobre 2006</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">128-134</span> <small style="line-height:1em;">(<a href="/wiki/PubMed" title="PubMed">PMID</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/16766053">16766053</a></span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">DOI</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://dx.doi.org/10.1016/j.molbiopara.2006.05.002">10.1016/j.molbiopara.2006.05.002</a></span>, <a rel="nofollow" class="external text" href="http://www.sciencedirect.com/science?_ob=ArticleURL&_udi=B6T29-4K18TKD-1&_user=10&_coverDate=10%2F31%2F2006&_rdoc=1&_fmt=high&_orig=search&_origin=search&_sort=d&_docanchor=&view=c&_acct=C000050221&_version=1&_urlVersion=0&_userid=10&md5=3c4bc380c02509d8e3af8b522a273ffc&searchtype=a">lire en ligne</a>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Identification+of+%27%27Leishmania%27%27+selenoproteins+and+SECIS+element&rft.jtitle=Molecular+and+Biochemical+Parasitology&rft.issue=2&rft.aulast=A.+Cassago%2C+E.M.+Rodrigues%2C+E.L.+Prieto%2C+K.W.+Gaston%2C+J.D.+Alfonzo+M.P.+Iribar%2C+M.J.+Berry%2C+A.K.+Cruz+et+O.H.+Thiemann&rft.date=2006-10&rft.volume=149&rft.pages=128-134&rft_id=info%3Adoi%2F10.1016%2Fj.molbiopara.2006.05.002&rft_id=info%3Apmid%2F16766053&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></li> <li><span class="ouvrage" id="Tobias_Mourier,_Arnab_Pain,_Bart_Barrell_et_Sam_Griffiths-Jones2005"><abbr class="abbr indicateur-langue" title="Langue : anglais">(en)</abbr> <span class="nom_auteur">Tobias Mourier, Arnab Pain, Bart Barrell et Sam Griffiths-Jones</span>, « <cite style="font-style:normal" lang="en">A selenocysteine tRNA and SECIS element in <i>Plasmodium falciparum</i></cite> », <i><span class="lang-en" lang="en">RNA</span></i>, <abbr class="abbr" title="volume">vol.</abbr> 11, <abbr class="abbr" title="numéro">n<sup>o</sup></abbr> 2,‎ <time>2005</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">119-122</span> <small style="line-height:1em;">(<a href="/wiki/PubMed" title="PubMed">PMID</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/15659354">15659354</a></span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">DOI</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://dx.doi.org/10.1261/rna.7185605">10.1261/rna.7185605</a></span>, <a rel="nofollow" class="external text" href="http://rnajournal.cshlp.org/content/11/2/119">lire en ligne</a>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=A+selenocysteine+tRNA+and+SECIS+element+in+%27%27Plasmodium+falciparum%27%27&rft.jtitle=RNA&rft.issue=2&rft.aulast=Tobias+Mourier%2C+Arnab+Pain%2C+Bart+Barrell+et+Sam+Griffiths-Jones&rft.date=2005&rft.volume=11&rft.pages=119-122&rft_id=info%3Adoi%2F10.1261%2Frna.7185605&rft_id=info%3Apmid%2F15659354&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></li> <li><span class="ouvrage" id="Gregory_V._Kryukov,_Sergi_Castellano,_Sergey_V._Novoselov,_Alexey_V._Lobanov,_Omid_Zehtab,_Roderic_Guigó_et_Vadim_N._Gladyshev2003"><abbr class="abbr indicateur-langue" title="Langue : anglais">(en)</abbr> <span class="nom_auteur">Gregory V. Kryukov, Sergi Castellano, Sergey V. Novoselov, Alexey V. Lobanov, Omid Zehtab, Roderic Guigó et Vadim N. Gladyshev</span>, « <cite style="font-style:normal" lang="en">Characterization of Mammalian Selenoproteomes</cite> », <i><span class="lang-en" lang="en">Science</span></i>, <abbr class="abbr" title="volume">vol.</abbr> 300, <abbr class="abbr" title="numéro">n<sup>o</sup></abbr> 5624,‎ <time class="nowrap" datetime="2003-05-30" data-sort-value="2003-05-30">30 mai 2003</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">1439-1443</span> <small style="line-height:1em;">(<a href="/wiki/PubMed" title="PubMed">PMID</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/12775843">12775843</a></span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">DOI</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://dx.doi.org/10.1126/science.1083516">10.1126/science.1083516</a></span>, <a rel="nofollow" class="external text" href="http://www.sciencemag.org/content/300/5624/1439">lire en ligne</a>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Characterization+of+Mammalian+Selenoproteomes&rft.jtitle=Science&rft.issue=5624&rft.aulast=Gregory+V.+Kryukov%2C+Sergi+Castellano%2C+Sergey+V.+Novoselov%2C+Alexey+V.+Lobanov%2C+Omid+Zehtab%2C+Roderic+Guig%C3%B3+et+Vadim+N.+Gladyshev&rft.date=2003-05-30&rft.volume=300&rft.pages=1439-1443&rft_id=info%3Adoi%2F10.1126%2Fscience.1083516&rft_id=info%3Apmid%2F12775843&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></li> <li><span class="ouvrage" id="Gregory_V._Kryukov_et_Vadim_N._Gladyshev2004"><abbr class="abbr indicateur-langue" title="Langue : anglais">(en)</abbr> <span class="nom_auteur">Gregory V. Kryukov et Vadim N. Gladyshev</span>, « <cite style="font-style:normal" lang="en">The prokaryotic selenoproteome</cite> », <i><span class="lang-en" lang="en">EMBO reports</span></i>, <abbr class="abbr" title="volume">vol.</abbr> 5, <abbr class="abbr" title="numéro">n<sup>o</sup></abbr> 5,‎ <time class="nowrap" datetime="2004-04-23" data-sort-value="2004-04-23">23 avril 2004</time>, <abbr class="abbr" title="pages">p.</abbr> <span class="nowrap">538-543</span> <small style="line-height:1em;">(<a href="/wiki/PubMed" title="PubMed">PMID</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/15105824">15105824</a></span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">DOI</a> <span class="plainlinks noarchive nowrap"><a rel="nofollow" class="external text" href="https://dx.doi.org/10.1038/sj.embor.7400126">10.1038/sj.embor.7400126</a></span>, <a rel="nofollow" class="external text" href="http://www.nature.com/embor/journal/v5/n5/full/7400126.html">lire en ligne</a>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=The+prokaryotic+selenoproteome&rft.jtitle=EMBO+reports&rft.issue=5&rft.aulast=Gregory+V.+Kryukov+et+Vadim+N.+Gladyshev&rft.date=2004-04-23&rft.volume=5&rft.pages=538-543&rft_id=info%3Adoi%2F10.1038%2Fsj.embor.7400126&rft_id=info%3Apmid%2F15105824&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></li> <li><span class="ouvrage" id="Alain_Krol2002"><abbr class="abbr indicateur-langue" title="Langue : anglais">(en)</abbr> <span class="nom_auteur">Alain Krol</span>, « <cite style="font-style:normal" lang="en">Evolutionarily different RNA motifs and RNA–protein complexes to achieve selenoprotein synthesis</cite> », <i><span class="lang-en" lang="en">Biochimie</span></i>, <abbr class="abbr" title="volume">vol.</abbr> 84, <abbr class="abbr" 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href="http://www.sciencedirect.com/science?_ob=ArticleURL&_udi=B6VRJ-4680PNG-4&_user=10&_coverDate=08%2F31%2F2002&_rdoc=1&_fmt=high&_orig=search&_origin=search&_sort=d&_docanchor=&view=c&_acct=C000050221&_version=1&_urlVersion=0&_userid=10&md5=fb5cb64c60b272ba2d181668390e639f&searchtype=a">lire en ligne</a>)</small><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Evolutionarily+different+RNA+motifs+and+RNA%E2%80%93protein+complexes+to+achieve+selenoprotein+synthesis&rft.jtitle=Biochimie&rft.issue=8&rft.aulast=Alain+Krol&rft.date=2002-08&rft.volume=84&rft.pages=765-774&rft_id=info%3Adoi%2F10.1016%2FS0300-9084%2802%2901405-0&rft_id=info%3Apmid%2F12457564&rfr_id=info%3Asid%2Ffr.wikipedia.org%3A%C3%89l%C3%A9ment+SECIS"></span></span></li></ul> </div> <ul id="bandeau-portail" class="bandeau-portail"><li><span class="bandeau-portail-element"><span class="bandeau-portail-icone"><span class="noviewer" 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