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</button> <ul id="toc-Taxonomy_and_naming-sublist" class="vector-toc-list"> <li id="toc-Genetics_and_evolution" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Genetics_and_evolution"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.1</span> <span>Genetics and evolution</span> </div> </a> <ul id="toc-Genetics_and_evolution-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Physical_description" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Physical_description"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Physical description</span> </div> </a> <ul id="toc-Physical_description-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Parasites" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Parasites"> <div class="vector-toc-text"> <span 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</button> <ul id="toc-Ecology_and_behavior-sublist" class="vector-toc-list"> <li id="toc-Diet" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Diet"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.1</span> <span>Diet</span> </div> </a> <ul id="toc-Diet-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Reproduction" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Reproduction"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2</span> <span>Reproduction</span> </div> </a> <ul id="toc-Reproduction-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Habitat_and_distribution" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Habitat_and_distribution"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>Habitat and distribution</span> </div> </a> <ul id="toc-Habitat_and_distribution-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Threats_and_conservation" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Threats_and_conservation"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>Threats and conservation</span> </div> </a> <ul id="toc-Threats_and_conservation-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">9</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Nyala</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 45 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-45" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">45 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Njala" title="Njala – Afrikaans" lang="af" hreflang="af" data-title="Njala" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-am mw-list-item"><a href="https://am.wikipedia.org/wiki/%E1%8A%92%E1%8B%AB%E1%88%8B" title="ኒያላ – Amharic" lang="am" hreflang="am" data-title="ኒያላ" data-language-autonym="አማርኛ" data-language-local-name="Amharic" class="interlanguage-link-target"><span>አማርኛ</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%86%D9%8A%D8%A7%D9%84%D8%A7_(%D8%B8%D8%A8%D9%8A)" title="نيالا (ظبي) – Arabic" lang="ar" hreflang="ar" data-title="نيالا (ظبي)" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Nyala" title="Nyala – Azerbaijani" lang="az" hreflang="az" data-title="Nyala" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%9D%D0%B8%D0%B0%D0%BB%D0%B0" title="Ниала – Bulgarian" lang="bg" hreflang="bg" data-title="Ниала" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-br mw-list-item"><a href="https://br.wikipedia.org/wiki/Nyala" title="Nyala – Breton" lang="br" hreflang="br" data-title="Nyala" data-language-autonym="Brezhoneg" data-language-local-name="Breton" class="interlanguage-link-target"><span>Brezhoneg</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Niala" title="Niala – Catalan" lang="ca" hreflang="ca" data-title="Niala" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ceb mw-list-item"><a href="https://ceb.wikipedia.org/wiki/Tragelaphus_angasii" title="Tragelaphus angasii – Cebuano" lang="ceb" hreflang="ceb" data-title="Tragelaphus angasii" data-language-autonym="Cebuano" data-language-local-name="Cebuano" class="interlanguage-link-target"><span>Cebuano</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Nyala_n%C3%AD%C5%BEinn%C3%A1" title="Nyala nížinná – Czech" lang="cs" hreflang="cs" data-title="Nyala nížinná" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Nyala_(Antilope)" title="Nyala (Antilope) – German" lang="de" hreflang="de" data-title="Nyala (Antilope)" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-nv mw-list-item"><a href="https://nv.wikipedia.org/wiki/J%C3%A1d%C3%ADtsoh_di%C5%82hi%C5%82%C3%ADg%C3%AD%C3%AD" title="Jádítsoh diłhiłígíí – Navajo" lang="nv" hreflang="nv" data-title="Jádítsoh diłhiłígíí" data-language-autonym="Diné bizaad" data-language-local-name="Navajo" class="interlanguage-link-target"><span>Diné bizaad</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Tragelaphus_angasii" title="Tragelaphus angasii – Spanish" lang="es" hreflang="es" data-title="Tragelaphus angasii" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Njalo" title="Njalo – Esperanto" lang="eo" hreflang="eo" data-title="Njalo" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Niala" title="Niala – Basque" lang="eu" hreflang="eu" data-title="Niala" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%86%DB%8C%D8%A7%D9%84%D8%A7_(%D8%AC%D8%A7%D9%86%D9%88%D8%B1)" title="نیالا (جانور) – Persian" lang="fa" hreflang="fa" data-title="نیالا (جانور)" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Nyala" title="Nyala – French" lang="fr" hreflang="fr" data-title="Nyala" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%8B%88%EC%95%8C%EB%9D%BC%EC%98%81%EC%96%91" title="니알라영양 – Korean" lang="ko" hreflang="ko" data-title="니알라영양" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%A8%E0%A5%8D%E0%A4%AF%E0%A4%BE%E0%A4%B2%E0%A4%BE" title="न्याला – Hindi" lang="hi" hreflang="hi" data-title="न्याला" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Nyala" title="Nyala – Indonesian" lang="id" hreflang="id" data-title="Nyala" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Tragelaphus_angasii" title="Tragelaphus angasii – Italian" lang="it" hreflang="it" data-title="Tragelaphus angasii" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A7%D7%95%D7%93%D7%95_%D7%A0%D7%99%D7%90%D7%9C%D7%94" title="קודו ניאלה – Hebrew" lang="he" hreflang="he" data-title="קודו ניאלה" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-avk mw-list-item"><a href="https://avk.wikipedia.org/wiki/Putcuvol_(Tragelaphus_angasii)" title="Putcuvol (Tragelaphus angasii) – Kotava" lang="avk" hreflang="avk" data-title="Putcuvol (Tragelaphus angasii)" data-language-autonym="Kotava" data-language-local-name="Kotava" class="interlanguage-link-target"><span>Kotava</span></a></li><li class="interlanguage-link interwiki-lez mw-list-item"><a href="https://lez.wikipedia.org/wiki/%D0%9D%D0%B9%D0%B0%D0%BB%D0%B0" title="Нйала – Lezghian" lang="lez" hreflang="lez" data-title="Нйала" data-language-autonym="Лезги" data-language-local-name="Lezghian" class="interlanguage-link-target"><span>Лезги</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Nijala" title="Nijala – Lithuanian" lang="lt" hreflang="lt" data-title="Nijala" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Nyala" title="Nyala – Hungarian" lang="hu" hreflang="hu" data-title="Nyala" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%86%D9%8A%D8%A7%D9%84%D8%A7_(%D9%86%D9%88%D8%B9_%D9%85%D9%86_%D8%A7%D9%84%D8%AB%D8%AF%D9%8A%D9%8A%D8%A7%D8%AA)" title="نيالا (نوع من الثدييات) – Egyptian Arabic" lang="arz" hreflang="arz" data-title="نيالا (نوع من الثدييات)" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Nyala" title="Nyala – Malay" lang="ms" hreflang="ms" data-title="Nyala" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Nyala_(antilope)" title="Nyala (antilope) – Dutch" lang="nl" hreflang="nl" data-title="Nyala (antilope)" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%8B%E3%82%A2%E3%83%A9" title="ニアラ – Japanese" lang="ja" hreflang="ja" data-title="ニアラ" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Nyala" title="Nyala – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Nyala" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-mhr mw-list-item"><a href="https://mhr.wikipedia.org/wiki/%D0%9D%D1%8C%D1%8F%D0%BB%D0%B0" title="Ньяла – Eastern Mari" lang="mhr" hreflang="mhr" data-title="Ньяла" data-language-autonym="Олык марий" data-language-local-name="Eastern Mari" class="interlanguage-link-target"><span>Олык марий</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D9%86%DB%8C%D8%A7%D9%84%DB%81" title="نیالہ – Western Punjabi" lang="pnb" hreflang="pnb" data-title="نیالہ" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pl badge-Q17437798 badge-goodarticle mw-list-item" title="good article badge"><a href="https://pl.wikipedia.org/wiki/Niala_grzywiasta" title="Niala grzywiasta – Polish" lang="pl" hreflang="pl" data-title="Niala grzywiasta" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Inhala" title="Inhala – Portuguese" lang="pt" hreflang="pt" data-title="Inhala" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9D%D1%8C%D1%8F%D0%BB%D0%B0" title="Ньяла – Russian" lang="ru" hreflang="ru" data-title="Ньяла" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Nyala" title="Nyala – Scots" lang="sco" hreflang="sco" data-title="Nyala" data-language-autonym="Scots" data-language-local-name="Scots" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Nyala" title="Nyala – Simple English" lang="en-simple" hreflang="en-simple" data-title="Nyala" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Njala" title="Njala – Finnish" lang="fi" hreflang="fi" data-title="Njala" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Nyala" title="Nyala – Swedish" lang="sv" hreflang="sv" data-title="Nyala" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%A8%E0%AE%AF%E0%AF%8D%E0%AE%AF%E0%AE%BE%E0%AE%B2%E0%AE%BE_%E0%AE%AE%E0%AE%BE%E0%AE%A9%E0%AF%8D" title="நய்யாலா மான் – Tamil" lang="ta" hreflang="ta" data-title="நய்யாலா மான்" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-udm mw-list-item"><a href="https://udm.wikipedia.org/wiki/%D0%9D%D1%8C%D1%8F%D0%BB%D0%B0" title="Ньяла – Udmurt" lang="udm" hreflang="udm" data-title="Ньяла" data-language-autonym="Удмурт" data-language-local-name="Udmurt" class="interlanguage-link-target"><span>Удмурт</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9D%D1%8C%D1%8F%D0%BB%D0%B0" title="Ньяла – Ukrainian" lang="uk" hreflang="uk" data-title="Ньяла" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Linh_d%C6%B0%C6%A1ng_Nyala" title="Linh dương Nyala – Vietnamese" lang="vi" hreflang="vi" data-title="Linh dương Nyala" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Tragelaphus_angasii" title="Tragelaphus angasii – Waray" lang="war" hreflang="war" data-title="Tragelaphus angasii" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%AE%89%E6%B0%8F%E6%9E%97%E7%BE%9A" title="安氏林羚 – Chinese" lang="zh" hreflang="zh" data-title="安氏林羚" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a 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Click here for more information."><img alt="This is a good article. Click here for more information." src="//upload.wikimedia.org/wikipedia/en/thumb/9/94/Symbol_support_vote.svg/19px-Symbol_support_vote.svg.png" decoding="async" width="19" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/94/Symbol_support_vote.svg/29px-Symbol_support_vote.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/94/Symbol_support_vote.svg/39px-Symbol_support_vote.svg.png 2x" data-file-width="180" data-file-height="185" /></a></span></div></div> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Species of antelope</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">This article is about the antelope. For other uses, see <a href="/wiki/Nyala_(disambiguation)" class="mw-disambig" title="Nyala (disambiguation)">Nyala (disambiguation)</a>.</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Nyala </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Nyala_bull_staring_at_me_(6073691776).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/b/bb/Nyala_bull_staring_at_me_%286073691776%29.jpg/220px-Nyala_bull_staring_at_me_%286073691776%29.jpg" decoding="async" width="220" height="176" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/bb/Nyala_bull_staring_at_me_%286073691776%29.jpg/330px-Nyala_bull_staring_at_me_%286073691776%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/bb/Nyala_bull_staring_at_me_%286073691776%29.jpg/440px-Nyala_bull_staring_at_me_%286073691776%29.jpg 2x" data-file-width="960" data-file-height="768" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Male </td></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Nyala-Female-Kruger-National-Park.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/49/Nyala-Female-Kruger-National-Park.jpg/220px-Nyala-Female-Kruger-National-Park.jpg" decoding="async" width="220" height="150" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/49/Nyala-Female-Kruger-National-Park.jpg/330px-Nyala-Female-Kruger-National-Park.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/49/Nyala-Female-Kruger-National-Park.jpg/440px-Nyala-Female-Kruger-National-Park.jpg 2x" data-file-width="5356" data-file-height="3648" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Female </td></tr> <tr style="color:inherit; text-align: center; background-color: rgb(235,235,210)"> <th colspan="2"><div style="text-align: center"><a href="/wiki/Conservation_status" title="Conservation status">Conservation status</a></div> </th></tr> <tr> <td colspan="2"><div style="text-align: center"><span class="mw-default-size skin-invert-image" typeof="mw:File/Frameless"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Status_iucn3.1_LC.svg/220px-Status_iucn3.1_LC.svg.png" decoding="async" width="220" height="59" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Status_iucn3.1_LC.svg/330px-Status_iucn3.1_LC.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Status_iucn3.1_LC.svg/440px-Status_iucn3.1_LC.svg.png 2x" data-file-width="512" data-file-height="137" /></span></span><br /><a href="/wiki/Least_Concern" class="mw-redirect" title="Least Concern">Least Concern</a> <small> (<a href="/wiki/IUCN_Red_List" title="IUCN Red List">IUCN 3.1</a>)<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></small></div> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Tragelaphus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Mammal" title="Mammal">Mammalia</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Artiodactyl" title="Artiodactyl">Artiodactyla</a> </td></tr> <tr> <td>Family: </td> <td><a href="/wiki/Bovidae" title="Bovidae">Bovidae</a> </td></tr> <tr> <td>Subfamily: </td> <td><a href="/wiki/Bovinae" title="Bovinae">Bovinae</a> </td></tr> <tr> <td>Genus: </td> <td><a href="/wiki/Tragelaphus" title="Tragelaphus"><i>Tragelaphus</i></a> </td></tr> <tr> <td>Species: </td> <td><div style="display:inline" class="species"><i><b>T. angasii</b></i></div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Binomial_nomenclature" title="Binomial nomenclature">Binomial name</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><b><span class="binomial"><span style="font-weight:normal;"></span><i>Tragelaphus angasii</i></span></b><br /><div style="font-size: 85%;">(<a href="/wiki/George_French_Angas" title="George French Angas">Angas</a>, 1849)</div> </td></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Tragelaphus_angasii_distribution.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/96/Tragelaphus_angasii_distribution.svg/220px-Tragelaphus_angasii_distribution.svg.png" decoding="async" width="220" height="189" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/96/Tragelaphus_angasii_distribution.svg/330px-Tragelaphus_angasii_distribution.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/96/Tragelaphus_angasii_distribution.svg/440px-Tragelaphus_angasii_distribution.svg.png 2x" data-file-width="501" data-file-height="430" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Geographic range </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Synonym_(taxonomy)" title="Synonym (taxonomy)">Synonyms</a><sup id="cite_ref-huffman_2-0" class="reference"><a href="#cite_note-huffman-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><i>Nyala angasii</i></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p>The <b>lowland nyala</b> or simply <b>nyala</b> (<i><b>Tragelaphus angasii</b></i>)<sup id="cite_ref-burton_3-0" class="reference"><a href="#cite_note-burton-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> is a spiral-horned <a href="/wiki/Artiodactyl" title="Artiodactyl">artiodactyl</a> <a href="/wiki/Antelope" title="Antelope">antelope</a> native to <a href="/wiki/Southern_Africa" title="Southern Africa">Southern Africa</a>. The species is part of the family <a href="/wiki/Bovidae" title="Bovidae">Bovidae</a> and the genus <i><a href="/wiki/Tragelaphus" title="Tragelaphus">Tragelaphus</a></i> (formerly placed in the genus <i>Nyala</i>). It was first <a href="/wiki/Species_description" title="Species description">described</a> in 1849 by <a href="/wiki/George_French_Angas" title="George French Angas">George French Angas</a> and exhibits the highest <a href="/wiki/Sexual_dimorphism" title="Sexual dimorphism">sexual dimorphism</a> among the spiral-horned antelopes. It is not to be confused with the endangered <a href="/wiki/Mountain_nyala" title="Mountain nyala">mountain nyala</a> living in the Bale region of Ethiopia. </p><p>The nyala's range encompasses much of <a href="/wiki/Southern_Africa" title="Southern Africa">Southern Africa</a>. As its population is relatively stable, it has been listed as of <a href="/wiki/Least_concern" class="mw-redirect" title="Least concern">least concern</a> by the <a href="/wiki/International_Union_for_Conservation_of_Nature" title="International Union for Conservation of Nature">International Union for Conservation of Nature</a>. It generally browses during the day in warm weather and during the night in the rainy season. As a herbivore, the nyala feeds upon foliage, fruits and grasses, and requires sufficient fresh water. A shy animal, it prefers water holes rather than open spaces. The nyala does not show signs of territoriality, and individuals' areas can overlap. They are very cautious creatures. </p><p>They live in single-sex or mixed family groups of up to ten individuals, but old males live alone. They inhabit thickets within dense and dry savanna woodlands. The principal threats to the species are illegal poaching and habitat loss resulting from human settlement and livestock grazing. However, the large-horned, impressive males are highly prized as <a href="/wiki/Game_(food)" class="mw-redirect" title="Game (food)">game animals</a> by sport hunters. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Taxonomy_and_naming">Taxonomy and naming</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=1" title="Edit section: Taxonomy and naming"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <table style="border: 1px solid #ccc; vertical-align: middle; width: 248px; margin:0.2em 1em 0.2em 0; float:left; clear:left;"> <tbody><tr> <td style="padding: 5px"> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1258728058">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="font-size:90%;line-height:100%;width:300px;"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Giant_eland" title="Giant eland">Giant eland</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Common_eland" title="Common eland">Common eland</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Greater_kudu" title="Greater kudu">Greater kudu</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Mountain_nyala" title="Mountain nyala">Mountain nyala</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Bongo_(antelope)" title="Bongo (antelope)">Bongo</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Sitatunga" title="Sitatunga">Sitatunga</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Cape_bushbuck" title="Cape bushbuck">Cape bushbuck</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Harnessed_bushbuck" title="Harnessed bushbuck">Harnessed bushbuck</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><b>Lowland nyala</b> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Lesser_kudu" title="Lesser kudu">Lesser kudu</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td style="text-align: left; font-size: 88%; border: 0; padding: 0;">Phylogenetic relationships of the mountain nyala from combined analysis of all molecular data (Willows-Munro et.al. 2005) </td></tr></tbody></table> <p>The nyala was first <a href="/wiki/Species_description" title="Species description">described</a> by <a href="/wiki/George_French_Angas" title="George French Angas">George French Angas</a>, an English naturalist, in 1849. The <a href="/wiki/Scientific_name" class="mw-redirect" title="Scientific name">scientific name</a> of nyala is <i>Tragelaphus angasii</i>. The name <i>angasii</i> is attributed to Angas, who said that <a href="/wiki/John_Edward_Gray" title="John Edward Gray">John Edward Gray</a> had named this species after Angas' father, <a href="/wiki/George_Fife_Angas" title="George Fife Angas">George Fife Angas</a> of South Australia. According to Article 50.1.1 of the <a href="/wiki/International_Code_of_Zoological_Nomenclature" title="International Code of Zoological Nomenclature">International Code of Zoological Nomenclature</a> and <a href="/wiki/International_Commission_on_Zoological_Nomenclature" title="International Commission on Zoological Nomenclature">International Commission on Zoological Nomenclature</a>, though, this is insufficient to state Gray as the author.<sup id="cite_ref-MSW3_4-0" class="reference"><a href="#cite_note-MSW3-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> The name "nyala" is the <a href="/wiki/Tsonga_language" title="Tsonga language">Tsonga</a> name for this antelope, which is likely the source of the English, along with <a href="/wiki/Zulu_language" title="Zulu language">Zulu</a> <span title="Zulu-language text"><i lang="zu">inyala</i></span>.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> Its first known use was in 1899. The word has a <a href="/wiki/Bantu_languages" title="Bantu languages">Bantu</a> origin, similar to the <a href="/wiki/Venda_language" title="Venda language">Venda</a> word <span title="Venda-language text"><i lang="ve">dzì-nyálà</i></span> (nyala buck).<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>The nyala is the second <a href="/wiki/Taxon" title="Taxon">taxon</a> to branch off from the tragelaphine family tree just after the <a href="/wiki/Lesser_kudu" title="Lesser kudu">lesser kudu</a>. As the nyala line has remained separate for a considerable time (over 5 million years), some authorities have placed it in its own monotypic genus <i>Nyala</i>. <i>Nyala</i> was proposed in 1912 by American zoologist <a href="/wiki/Edmund_Heller" title="Edmund Heller">Edmund Heller</a>, who also proposed <i>Ammelaphus</i> for the lesser kudu,<sup id="cite_ref-heller_7-0" class="reference"><a href="#cite_note-heller-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> but it was not widely recognized. It was re-erected as a valid genus in 2011 under the classification of <a href="/wiki/Peter_Grubb_(zoologist)" title="Peter Grubb (zoologist)">Peter Grubb</a> and <a href="/wiki/Colin_Groves" title="Colin Groves">Colin Groves</a>,<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> but has not been embraced by taxonomic authorities such as the Mammal Diversity Database.<sup id="cite_ref-burton_3-1" class="reference"><a href="#cite_note-burton-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> Among its closest extant relatives are the forest-inhabiting <a href="/wiki/Bongo_(antelope)" title="Bongo (antelope)">bongo</a>, the <a href="/wiki/Cape_bushbuck" title="Cape bushbuck">bushbuck</a>, the large <a href="/wiki/Common_eland" title="Common eland">common</a> and <a href="/wiki/Giant_eland" title="Giant eland">giant elands</a>, the <a href="/wiki/Greater_kudu" title="Greater kudu">greater</a> and <a href="/wiki/Lesser_kudu" title="Lesser kudu">lesser kudus</a>, the <a href="/wiki/Mountain_nyala" title="Mountain nyala">mountain nyala</a> (of <a href="/wiki/Ethiopia" title="Ethiopia">Ethiopia</a>) and the swamp-dwelling <a href="/wiki/Sitatunga" title="Sitatunga">sitatunga</a>, all of which share similar characteristics, such as hornless females, vertical side-stripes and other unique white markings, and the spiraling horns and distinct "beard" or <a href="/wiki/Dewlap" title="Dewlap">dewlap</a> of males, often with a raised trail of fur extending down the back and underside.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (May 2024)">citation needed</span></a></i>]</sup> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:ZOO_Hellabrunn,_M%C3%BCnchen_-_kudui.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f9/ZOO_Hellabrunn%2C_M%C3%BCnchen_-_kudui.jpg/220px-ZOO_Hellabrunn%2C_M%C3%BCnchen_-_kudui.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f9/ZOO_Hellabrunn%2C_M%C3%BCnchen_-_kudui.jpg/330px-ZOO_Hellabrunn%2C_M%C3%BCnchen_-_kudui.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f9/ZOO_Hellabrunn%2C_M%C3%BCnchen_-_kudui.jpg/440px-ZOO_Hellabrunn%2C_M%C3%BCnchen_-_kudui.jpg 2x" data-file-width="2592" data-file-height="1944" /></a><figcaption>Nyala family in captivity</figcaption></figure> <p>In 2005, Sandi Willows-Munro (of the <a href="/wiki/University_of_KwaZulu-Natal" title="University of KwaZulu-Natal">University of KwaZulu-Natal</a>) and colleagues carried out a <a href="/wiki/Mitochondrial_DNA" title="Mitochondrial DNA">mitochondrial DNA</a> analysis of the nine <i>Tragelaphus</i> species. Mitochondrial DNA and <a href="/wiki/NDNA" class="mw-redirect" title="NDNA">nuclear DNA</a> data were compared. The results showed the tribe Tragelaphini to be <a href="/wiki/Monophyletic" class="mw-redirect" title="Monophyletic">monophyletic</a>, with the <a href="/wiki/Lesser_kudu" title="Lesser kudu">lesser kudu</a> (<i>T.  imberbis</i>) <a href="/wiki/Basal_(phylogenetics)" title="Basal (phylogenetics)">basal</a> in the phylogeny, followed by the nyala.<sup id="cite_ref-cladogram_10-0" class="reference"><a href="#cite_note-cladogram-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-groves2014_11-0" class="reference"><a href="#cite_note-groves2014-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> On the basis of mitochondrial data, studies have estimated that the lesser kudu separated from its <a href="/wiki/Sister_clade" class="mw-redirect" title="Sister clade">sister clade</a> around 13.7 million years ago. However, nuclear DNA data shows lesser kudu and nyala forming a clade, which collectively separated from the sister clade 13.8 million years ago.<sup id="cite_ref-ropiquet_12-0" class="reference"><a href="#cite_note-ropiquet-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-hassanin_13-0" class="reference"><a href="#cite_note-hassanin-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Genetics_and_evolution">Genetics and evolution</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=2" title="Edit section: Genetics and evolution"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The nyala has 55 male <a href="/wiki/Chromosome" title="Chromosome">chromosomes</a> and 56 female chromosomes.<sup id="cite_ref-Rubes_14-0" class="reference"><a href="#cite_note-Rubes-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> The <a href="/wiki/Y_chromosome" title="Y chromosome">Y chromosome</a> has been <a href="/wiki/Chromosomal_translocation" title="Chromosomal translocation">translocated</a> onto the 14th chromosome, as in other <a href="/wiki/Tragelaphus" title="Tragelaphus">tragelaphids</a>, but no <a href="/wiki/Chromosomal_inversion" title="Chromosomal inversion">inversion</a> of the Y chromosome occurs.<sup id="cite_ref-MSW3_4-1" class="reference"><a href="#cite_note-MSW3-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Cranial studies have shown that the <a href="/wiki/Mountain_nyala" title="Mountain nyala">mountain nyala</a> and nyala, though sharing a common name, are actually distant relatives.<sup id="cite_ref-antelope_15-0" class="reference"><a href="#cite_note-antelope-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p><p>Fossil evidence suggests that the nyala has been a separate species since the end of the <a href="/wiki/Miocene" title="Miocene">Miocene</a> (5.8 million years ago). Genetic evidence suggests that the proto-nyala had some early hybridization with the proto-lesser kudu, but the two have remained separate long after this crossing.<sup id="cite_ref-MSW3_4-2" class="reference"><a href="#cite_note-MSW3-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Physical_description">Physical description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=3" title="Edit section: Physical description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Nyalas_male.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/04/Nyalas_male.jpg/220px-Nyalas_male.jpg" decoding="async" width="220" height="167" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/04/Nyalas_male.jpg/330px-Nyalas_male.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/04/Nyalas_male.jpg/440px-Nyalas_male.jpg 2x" data-file-width="1011" data-file-height="768" /></a><figcaption>Male in the Kruger National Park</figcaption></figure> <p>The nyala is a spiral-horned and middle-sized antelope, between a <a href="/wiki/Bushbuck" class="mw-redirect" title="Bushbuck">bushbuck</a> and a <a href="/wiki/Kudu" title="Kudu">kudu</a>.<sup id="cite_ref-unwin_16-0" class="reference"><a href="#cite_note-unwin-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> It is considered the most <a href="/wiki/Sexually_dimorphic" class="mw-redirect" title="Sexually dimorphic">sexually dimorphic</a> antelope.<sup id="cite_ref-huffman_2-1" class="reference"><a href="#cite_note-huffman-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The nyala is typically between 135–195 cm (53–77 in) in head-and-body length.<sup id="cite_ref-huffman_2-2" class="reference"><a href="#cite_note-huffman-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The male stands up to 110 cm (43 in), the female is up to 90 cm (3.0 ft) tall. Males weigh 98–125 kg (216–276 lb), while females weigh 55–68 kg (121–150 lb). Life expectancy of the nyala is about 19 years.<sup id="cite_ref-adw_17-0" class="reference"><a href="#cite_note-adw-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p><p>The coat is rusty or <a href="/wiki/Rufous" title="Rufous">rufous</a> brown in females and juveniles. It grows a dark brown or slate grey in adult males, often with a bluish tinge.<sup id="cite_ref-huffman_2-3" class="reference"><a href="#cite_note-huffman-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Females and young males have ten or more white vertical stripes on their sides. Other markings are visible on the face, throat, flanks and thighs. Stripes are very reduced or absent in older males. Both males and females have a white <a href="/wiki/Chevron_(anatomy)" class="mw-redirect" title="Chevron (anatomy)">chevron</a> between their eyes, and a 40–55 cm (16–22 in) long bushy tail white underside. Both sexes have a dorsal crest of hair running right from the back of the head to the end of the tail. Males have another line of hair along the midline of their chest and belly.<sup id="cite_ref-adw_17-1" class="reference"><a href="#cite_note-adw-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-estes_18-0" class="reference"><a href="#cite_note-estes-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p><p>Only the males have horns. Horns are 60–83 cm (24–33 in) long and yellow-tipped. There are one or two twists.<sup id="cite_ref-huffman_2-4" class="reference"><a href="#cite_note-huffman-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The <a href="/wiki/Spoor_(animal)" title="Spoor (animal)">spoor</a> is similar to that of the bushbuck, but larger. It is 5–6 cm (2.0–2.4 in) long. The feces resemble round to spherical pellets.<sup id="cite_ref-walker_19-0" class="reference"><a href="#cite_note-walker-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> The nyala has hairy glands on its feet, which leave their scent wherever it walks.<sup id="cite_ref-burton_3-2" class="reference"><a href="#cite_note-burton-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p>The condition of the nyala often varies between the sexes. According to a study, this can be attributed to the differences in their body sizes. It was noted that during nutritional stress, old adults died in more numbers, of which most were males.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> During an attempt of blood sampling in the nyala, it was found that <a href="/wiki/Vitamin_E" title="Vitamin E">Vitamin E</a> levels varied during stress.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Parasites">Parasites</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=4" title="Edit section: Parasites"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>A study of the <a href="/wiki/Helminth" class="mw-redirect" title="Helminth">helminths</a> from 77 nyalas from four game reserves in <a href="/wiki/Natal_Province" class="mw-redirect" title="Natal Province">Natal</a> revealed the presence of ten <a href="/wiki/Nematode" title="Nematode">nematode</a> species and four nematode genera, a <a href="/wiki/Trematoda" title="Trematoda">trematode</a> species and paramphistomes (members of superfamily <i><a href="/wiki/Paramphistomoidea" class="mw-redirect" title="Paramphistomoidea">Paramphistomoidea</a></i>), and two <a href="/wiki/Cestode" class="mw-redirect" title="Cestode">cestode</a> genera. The research discovered new parasites that the nyala was host of - namely a <i><a href="/w/index.php?title=Cooperia_rotundispiculum&action=edit&redlink=1" class="new" title="Cooperia rotundispiculum (page does not exist)">Cooperia rotundispiculum</a></i> race, <i><a href="/w/index.php?title=Gaigeria_pachyscelis&action=edit&redlink=1" class="new" title="Gaigeria pachyscelis (page does not exist)">Gaigeria pachyscelis</a></i>, a <i><a href="/wiki/Gongylonema" title="Gongylonema">Gongylonema</a></i> species, <i><a href="/w/index.php?title=Haemonchus_vegliai&action=edit&redlink=1" class="new" title="Haemonchus vegliai (page does not exist)">Haemonchus vegliai</a></i>, <i><a href="/w/index.php?title=Impalaia_tuberculata&action=edit&redlink=1" class="new" title="Impalaia tuberculata (page does not exist)">Impalaia tuberculata</a></i>, an <i><a href="/wiki/Oesophagostomum" title="Oesophagostomum">Oesophagostomum</a></i> species, a <i><a href="/wiki/Setaria" title="Setaria">Setaria</a></i> species, <i><a href="/w/index.php?title=Trichostrongylus_deflexus&action=edit&redlink=1" class="new" title="Trichostrongylus deflexus (page does not exist)">Trichostrongylus deflexus</a></i>, <i><a href="/w/index.php?title=Trichostrongylus_falculatus&action=edit&redlink=1" class="new" title="Trichostrongylus falculatus (page does not exist)">Trichostrongylus falculatus</a></i>, the larval stage of a <i><a href="/wiki/Taenia_(flatworm)" title="Taenia (flatworm)">Taenia</a></i> species, a <i><a href="/wiki/Thysaniezia" title="Thysaniezia">Thysaniezia</a></i> species and <i><a href="/wiki/Schistosoma_mattheei" class="mw-redirect" title="Schistosoma mattheei">Schistosoma mattheei</a></i>. <i><a href="/w/index.php?title=Ostertagia_harrisi&action=edit&redlink=1" class="new" title="Ostertagia harrisi (page does not exist)">Ostertagia harrisi</a></i> and <i>C. rotundispiculum</i> were the most dominant nematodes in the antelope.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> </p><p>Another study of 97 blood samples of South African nyalas revealed the presence of <a href="/wiki/Tick" title="Tick">tick</a>-borne hemoparasites (blood parasites). The methods used were <a href="/wiki/Polymerase_chain_reaction" title="Polymerase chain reaction">polymerase chain reaction</a> (PCR) and reverse line blot (RLB) hybridization. The dominant parasites were <i><a href="/wiki/Theileria" title="Theileria">Theileria</a></i> species, <i><a href="/w/index.php?title=Theileria_buffeli&action=edit&redlink=1" class="new" title="Theileria buffeli (page does not exist)">T. buffeli</a></i>, <i><a href="/w/index.php?title=Theileria_bicornis&action=edit&redlink=1" class="new" title="Theileria bicornis (page does not exist)">T. bicornis</a></i>, <i><a href="/wiki/Ehrlichia" title="Ehrlichia">Ehrlichia</a></i> species, <i><a href="/wiki/Anaplasma_marginale" class="mw-redirect" title="Anaplasma marginale">Anaplasma marginale</a></i> and <i><a href="/wiki/Anaplasma_bovis" title="Anaplasma bovis">A. bovis</a></i>.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> Ten tick species, two <a href="/wiki/Louse" title="Louse">louse</a> species and a louse fly species were recovered in a study of 73 nyalas at Umfolozi, Mkuzi and Ndumu Game Reserves in northeastern KwaZulu-Natal in 1983 and 1984 and an additional six individuals in 1994. It was found that nyalas were hosts to all stages of development in <i><a href="/w/index.php?title=Boophilus_decoloratus&action=edit&redlink=1" class="new" title="Boophilus decoloratus (page does not exist)">Boophilus decoloratus</a></i>, <i><a href="/wiki/Rhipicephalus_appendiculatus" title="Rhipicephalus appendiculatus">Rhipicephalus appendiculatus</a></i> and <i><a href="/w/index.php?title=Rhipicephalus_muehlensi&action=edit&redlink=1" class="new" title="Rhipicephalus muehlensi (page does not exist)">R. muehlensi</a></i> and the immature stages of <i><a href="/wiki/Amblyomma_hebraeum" title="Amblyomma hebraeum">Amblyomma hebraeum</a></i> and <i><a href="/w/index.php?title=Rhipicephalus_maculatus&action=edit&redlink=1" class="new" title="Rhipicephalus maculatus (page does not exist)">Rhipicephalus maculatus</a></i>. Adult males served hosts to more ticks and lice than adult females did.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> Also, a <a href="/wiki/Trypanosoma" title="Trypanosoma">trypanosome</a> was isolated from a nyala, wild-caught in Mozambique, which was diagnosed and found as akin to <i><a href="/wiki/Trypanosoma_vivax" title="Trypanosoma vivax">Trypanosoma vivax</a></i>, based on biological, morphological and molecular data.<sup id="cite_ref-rodrigues_25-0" class="reference"><a href="#cite_note-rodrigues-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Diseases">Diseases</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=5" title="Edit section: Diseases"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure typeof="mw:File/Thumb"><a href="/wiki/File:Nyala_heart_(Tragelaphus_angasii).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/20/Nyala_heart_%28Tragelaphus_angasii%29.jpg/200px-Nyala_heart_%28Tragelaphus_angasii%29.jpg" decoding="async" width="200" height="299" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/20/Nyala_heart_%28Tragelaphus_angasii%29.jpg/300px-Nyala_heart_%28Tragelaphus_angasii%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/20/Nyala_heart_%28Tragelaphus_angasii%29.jpg/400px-Nyala_heart_%28Tragelaphus_angasii%29.jpg 2x" data-file-width="2307" data-file-height="3446" /></a><figcaption>Nyala heart</figcaption></figure> <p>The nyala can also suffer from <a href="/wiki/Myopathy" title="Myopathy">myopathy</a>. In between January 1973 and June 1981, 21 nyalas succumbed to the disease. The main symptoms were stiffness, inability to rise, and failure to suckle in newborns. <a href="/wiki/Necrosis" title="Necrosis">Necrosis</a> (that is, the premature death of cells in a living tissue) and <a href="/wiki/Ossification" title="Ossification">mineralization</a> were found in the skeletal muscle after a histological analysis. In the juveniles there was acute necrosis of the <a href="/wiki/Cardiac_muscle" title="Cardiac muscle">cardiac muscle</a>. In adults, there was interstitial <a href="/wiki/Fibrosis" title="Fibrosis">fibrosis</a> of the cardiac muscle, along with <a href="/wiki/Arteriosclerosis" title="Arteriosclerosis">arteriosclerosis</a>.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup> </p><p>In a report published in 1994 entitled "Epidemiological observations on spongiform encephalopathies in captive wild animals in the British Isles", it was noted that <a href="/wiki/Spongiform_encephalopathy" class="mw-redirect" title="Spongiform encephalopathy">spongiform encephalopathy</a> had been diagnosed in one nyala captive in a zoo.<sup id="cite_ref-bse_27-0" class="reference"><a href="#cite_note-bse-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup> The nyala was formerly affected by the disease <a href="/wiki/Rinderpest" title="Rinderpest">rinderpest</a>, although the viral disease is considered eradicated now.<sup id="cite_ref-iucn_28-0" class="reference"><a href="#cite_note-iucn-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Ecology_and_behavior">Ecology and behavior</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=6" title="Edit section: Ecology and behavior"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Nursing_nyala.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2b/Nursing_nyala.jpg/220px-Nursing_nyala.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2b/Nursing_nyala.jpg/330px-Nursing_nyala.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2b/Nursing_nyala.jpg/440px-Nursing_nyala.jpg 2x" data-file-width="3872" data-file-height="2592" /></a><figcaption>Nursing juvenile</figcaption></figure> <p>The nyala is active mainly in the early morning and late afternoon. It browses during the day if temperatures are 20–30 °C (68–86 °F) and during the night in rainy season.<sup id="cite_ref-tello_29-0" class="reference"><a href="#cite_note-tello-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> These antelopes rest in thick bushes during the hot hours of the day.<sup id="cite_ref-huffman_2-5" class="reference"><a href="#cite_note-huffman-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The nyala is very shy and cautious in nature, and often remains hidden rather than coming out in the open. Most sightings of the nyala in the wild are at <a href="/wiki/Depression_(geology)" title="Depression (geology)">water holes</a>.<sup id="cite_ref-huffman_2-6" class="reference"><a href="#cite_note-huffman-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> But in protected areas they become less shy and often come out in view of tourists.<sup id="cite_ref-burton_3-3" class="reference"><a href="#cite_note-burton-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p>Nyala groups are according to sex or mixed. Herds usually browse and drink water together.<sup id="cite_ref-unwin_16-1" class="reference"><a href="#cite_note-unwin-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> Each group consists of two to ten individuals. A study in <a href="/wiki/Zinave_National_Park" title="Zinave National Park">Zinave National Park</a> at Mozambique showed that 67% of the observations were of groups of one to three nyalas, and the rest of the herds consisted of up to 30 nyalas. Herds often broke up and re-formed.<sup id="cite_ref-tello_29-1" class="reference"><a href="#cite_note-tello-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> Generally adult males remain alone. Females often remain near their mothers when they have their offspring, so the relationships in female herds may be closer than those of males.<sup id="cite_ref-adw_17-2" class="reference"><a href="#cite_note-adw-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup> </p><p>Alert and wary in nature, the nyala use a sharp, high, dog-like bark to warn others in a group of danger. This feature is mainly used by females.<sup id="cite_ref-adw_17-3" class="reference"><a href="#cite_note-adw-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> They also react to the alarm calls of <a href="/wiki/Impala" title="Impala">impala</a>, <a href="/wiki/Baboon" title="Baboon">baboon</a> and kudu. The impala has been found to react to the calls of the nyala as well.<sup id="cite_ref-adw_17-4" class="reference"><a href="#cite_note-adw-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> The main predators of nyala are <a href="/wiki/Lion" title="Lion">lion</a>, <a href="/wiki/Leopard" title="Leopard">leopard</a>, <a href="/wiki/Cheetah" title="Cheetah">cheetah</a>, <a href="/wiki/Spotted_hyena" title="Spotted hyena">spotted hyena</a>, <a href="/wiki/African_wild_dog" title="African wild dog">African wild dog</a> and <a href="/wiki/Nile_crocodile" title="Nile crocodile">nile crocodile</a> while baboons and <a href="/wiki/Bird_of_prey" title="Bird of prey">raptorial birds</a> are predators of juveniles.<sup id="cite_ref-tello_29-2" class="reference"><a href="#cite_note-tello-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-planet_31-0" class="reference"><a href="#cite_note-planet-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Diet">Diet</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=7" title="Edit section: Diet"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>As a <a href="/wiki/Herbivore" title="Herbivore">herbivore</a>, the nyala's diet consists of foliage, fruits, flowers and twigs. During the rainy season they feed upon the fresh grass. They need a regular intake of water, and thus choose places with a water source nearby. However, they are adapted to live in areas with only a seasonal availability of water.<sup id="cite_ref-adw_17-5" class="reference"><a href="#cite_note-adw-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> A study in <a href="/wiki/Zululand_District_Municipality" title="Zululand District Municipality">Zululand</a> showed that the nyala fed mainly in the early morning and the late afternoon.<sup id="cite_ref-nowak_32-0" class="reference"><a href="#cite_note-nowak-32"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup> They feed at night during the rainy season.<sup id="cite_ref-planet_31-1" class="reference"><a href="#cite_note-planet-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup> </p><p>A study in <a href="/wiki/Mkhuze_Game_Reserve" class="mw-redirect" title="Mkhuze Game Reserve">Mkhuze Game Reserve</a> and <a href="/w/index.php?title=Ndumu_Game_Reserve&action=edit&redlink=1" class="new" title="Ndumu Game Reserve (page does not exist)">Ndumu Game Reserve</a> in Natal focused on the dietary habits of the impala and the nyala showed that the amount of <a href="/wiki/Dicotyledon" title="Dicotyledon">dicotyledons</a> in their diets varied seasonally. In the dry season, the nyala's dicotyledon diet content was 83.2% and the impala's 52%. In this season, the diet grew richer in fiber and dietary proteins were less. The reverse occurred in the rainy season. As the rainy season arrived, both species took to a diet of mainly <a href="/wiki/Monocotyledon" title="Monocotyledon">monocotyledons</a>, and the impala consumed more of them. The diet contained more proteins than fiber.<sup id="cite_ref-rooyen_33-0" class="reference"><a href="#cite_note-rooyen-33"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup> </p><p>Another study was done to find whether the sexual dimorphism in the nyala influenced its foraging habits. Vegetation surveys were conducted with the end of each feeding bout. It was found that females spent equal periods of time foraging in all the three habitats, but males preferred sand forest more. More differences were noted, as males ate woody species at a greater average height whereas females fed from the low <a href="/wiki/Herbaceous_layer" class="mw-redirect" title="Herbaceous layer">herbaceous layer</a>. It was concluded that the differences resulted from varying nutritional and energetic demands according to their diverse body sizes and differing reproductive strategies.<sup id="cite_ref-kirby_34-0" class="reference"><a href="#cite_note-kirby-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Reproduction">Reproduction</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=8" title="Edit section: Reproduction"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Fighting_Nyalas.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e9/Fighting_Nyalas.JPG/220px-Fighting_Nyalas.JPG" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e9/Fighting_Nyalas.JPG/330px-Fighting_Nyalas.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e9/Fighting_Nyalas.JPG/440px-Fighting_Nyalas.JPG 2x" data-file-width="3648" data-file-height="2736" /></a><figcaption>Males fighting over dominance</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Nyala_(Tragelaphus_angasii)_young_male_2w.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Nyala_%28Tragelaphus_angasii%29_young_male_2w.jpg/220px-Nyala_%28Tragelaphus_angasii%29_young_male_2w.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Nyala_%28Tragelaphus_angasii%29_young_male_2w.jpg/330px-Nyala_%28Tragelaphus_angasii%29_young_male_2w.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Nyala_%28Tragelaphus_angasii%29_young_male_2w.jpg/440px-Nyala_%28Tragelaphus_angasii%29_young_male_2w.jpg 2x" data-file-width="5472" data-file-height="3648" /></a><figcaption>Male, two weeks old<br /><a href="/wiki/Tswalu_Kalahari_Reserve" title="Tswalu Kalahari Reserve">Tswalu Kalahari Reserve</a>, <a href="/wiki/South_Africa" title="South Africa">South Africa</a></figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Nyala_(Tragelaphus_angasii)_young_male_2m_head.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/21/Nyala_%28Tragelaphus_angasii%29_young_male_2m_head.jpg/220px-Nyala_%28Tragelaphus_angasii%29_young_male_2m_head.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/21/Nyala_%28Tragelaphus_angasii%29_young_male_2m_head.jpg/330px-Nyala_%28Tragelaphus_angasii%29_young_male_2m_head.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/21/Nyala_%28Tragelaphus_angasii%29_young_male_2m_head.jpg/440px-Nyala_%28Tragelaphus_angasii%29_young_male_2m_head.jpg 2x" data-file-width="5472" data-file-height="3648" /></a><figcaption>Male, two months old<br />feeding</figcaption></figure> <p>The nyala breeds throughout the year, but <a href="/wiki/Mating" title="Mating">mating</a> peaks in spring and autumn. The reason for this is still unknown, but attributed to the <a href="/wiki/Photoperiodism" title="Photoperiodism">photoperiod</a> and the feeding habits of the animal.<sup id="cite_ref-anderson2_35-0" class="reference"><a href="#cite_note-anderson2-35"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup> Females reach <a href="/wiki/Sexual_maturity" title="Sexual maturity">sexual maturity</a> at 11 to 12 months of age and males at 18 months (though they are socially immature until five years old),<sup id="cite_ref-huffman_2-7" class="reference"><a href="#cite_note-huffman-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> though they begin to show active <a href="/wiki/Spermatogenesis" title="Spermatogenesis">spermatogenesis</a> at 14 months.<sup id="cite_ref-anderson_36-0" class="reference"><a href="#cite_note-anderson-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </p><p>Before ovulation, the <a href="/wiki/Graafian_follicles" class="mw-redirect" title="Graafian follicles">Graafian follicles</a> reach a length of at least 6.7 cm (2.6 in).<sup id="cite_ref-anderson_36-1" class="reference"><a href="#cite_note-anderson-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> A female's estrous cycle is about 19 days long. Males will attempt to mate with the female for two days of the cycle, but she allows it for only six hours per cycle. When the male enters a females' herd during mating, he makes a display by raising his white dorsal crest, lowering his horns and moving stiffly. As in many other animals, the males fight over <a href="/wiki/Dominance_(ethology)" class="mw-redirect" title="Dominance (ethology)">dominance</a> during mating.<sup id="cite_ref-adw_17-6" class="reference"><a href="#cite_note-adw-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p><p>The kidney fat indices (KFIs) of impalas and nyalas have been studied to understand the influence of social class and reproduction on them. To determine the KFI, the kidney is removed and weighed with the fat and once again excluding the fat. The resultant difference is the amount of fat on the kidney. The more the fat, the healthier the animal.<sup id="cite_ref-kfi_37-0" class="reference"><a href="#cite_note-kfi-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup> In <a href="/wiki/Rut_(mammalian_reproduction)" title="Rut (mammalian reproduction)">rut</a>, male nyalas had lower KFIs, which did not vary much with the season. Pregnant females of both nyala and impala had higher KFIs than non-pregnant ones.<sup id="cite_ref-kfi2_38-0" class="reference"><a href="#cite_note-kfi2-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup> </p><p>There is a significant increase in <a href="/wiki/Corpus_luteum" title="Corpus luteum">corpus luteum</a> in the last third of gestation.<sup id="cite_ref-anderson_36-2" class="reference"><a href="#cite_note-anderson-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> Gestation is of seven months. A single calf is born, weighing 5 kg (11 lb). Birth takes place generally away from the sight of predators, in places such as a thicket. The calf remains hidden for up to 18 days, and the mother nurses it at regular intervals. The calf remains with its mother until the birth of the next calf, during which males in rut drive it away from the mother.<sup id="cite_ref-adw_17-7" class="reference"><a href="#cite_note-adw-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Habitat_and_distribution">Habitat and distribution</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=9" title="Edit section: Habitat and distribution"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Nyalas.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a1/Nyalas.jpg/220px-Nyalas.jpg" decoding="async" width="220" height="97" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a1/Nyalas.jpg/330px-Nyalas.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a1/Nyalas.jpg/440px-Nyalas.jpg 2x" data-file-width="1924" data-file-height="844" /></a><figcaption>Nyalas choose habitat with fresh water sources nearby.</figcaption></figure> <p>The nyala inhabits dense lowland woodlands and thickets, mainly in southern Malawi, Mozambique, Zimbabwe, and eastern South Africa. It chooses places with good quality grasslands as well as provision of fresh water.<sup id="cite_ref-adw_17-8" class="reference"><a href="#cite_note-adw-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> It also inhabits lush green river country.<sup id="cite_ref-burton_3-4" class="reference"><a href="#cite_note-burton-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> The nyala's natural range stretches across southeast Africa from the Lower Shire Valley in Malawi through Mozambique and Zimbabwe to eastern South Africa and Eswatini.<sup id="cite_ref-iucn_28-1" class="reference"><a href="#cite_note-iucn-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p><p>The geographic distribution of the nyala may be based on the genetic variation. According to a study of nyala in South Africa, Mozambique, Malawi and Zimbabwe, there was a marked difference in the <a href="/wiki/Gene_frequencies" class="mw-redirect" title="Gene frequencies">gene frequencies</a> at three <a href="/wiki/Microsatellite" title="Microsatellite">microsatellite</a> <a href="/wiki/Locus_(genetics)" title="Locus (genetics)">loci</a>. Mitochondrial DNA analysis revealed the presence of a unique <a href="/wiki/Haplotype" title="Haplotype">haplotype</a> in individuals from each location. Thus, the geographic variation in the nyala may be due to a distribution pattern based on habitat specificity.<sup id="cite_ref-grobler_39-0" class="reference"><a href="#cite_note-grobler-39"><span class="cite-bracket">[</span>39<span class="cite-bracket">]</span></a></sup> </p><p>Today nyala are found in South African protected areas in the <a href="/wiki/Ndumo_Game_Reserve" title="Ndumo Game Reserve">Ndumo Game Reserve</a>, uMkuze Game Reserve and <a href="/wiki/Hluhluwe-Umfolozi_Game_Reserve" class="mw-redirect" title="Hluhluwe-Umfolozi Game Reserve">Hluhluwe-Umfolozi Game Reserve</a>, all in <a href="/wiki/KwaZulu-Natal" title="KwaZulu-Natal">KwaZulu-Natal</a>, as well as in <a href="/wiki/Kruger_National_Park" title="Kruger National Park">Kruger National Park</a>. As of 1999, 10–15% of nyala occurred on private land. Efforts are being made to retain the populations of nyala in <a href="/wiki/Gorongosa_National_Park" title="Gorongosa National Park">Gorongosa National Park</a> and <a href="/wiki/Banhine_National_Park" title="Banhine National Park">Banhine National Park</a> in Mozambique.<sup id="cite_ref-iucn_28-2" class="reference"><a href="#cite_note-iucn-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> Nyala also thrive in <a href="/wiki/Lengwe_National_Park" title="Lengwe National Park">Lengwe National Park</a> in Malawi. </p><p>Nyala have never been observed showing <a href="/wiki/Territory_(animal)" title="Territory (animal)">territoriality</a>. Territories of either sex overlap extensively.<sup id="cite_ref-tello_29-3" class="reference"><a href="#cite_note-tello-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> The home ranges of males are approximately equal to that of females, about 10 square kilometres (3.9 sq mi) in area.<sup id="cite_ref-tello_29-4" class="reference"><a href="#cite_note-tello-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:South_African_Safari_Wildlife1.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/76/South_African_Safari_Wildlife1.jpg/220px-South_African_Safari_Wildlife1.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/76/South_African_Safari_Wildlife1.jpg/330px-South_African_Safari_Wildlife1.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/76/South_African_Safari_Wildlife1.jpg/440px-South_African_Safari_Wildlife1.jpg 2x" data-file-width="1080" data-file-height="720" /></a><figcaption>Young nyala bull in the Kruger National Park</figcaption></figure> <div class="mw-heading mw-heading2"><h2 id="Threats_and_conservation">Threats and conservation</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=10" title="Edit section: Threats and conservation"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The major threats to the population of the nyala are poaching, habitat loss, agriculture and cattle grazing.<sup id="cite_ref-iucn_28-3" class="reference"><a href="#cite_note-iucn-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> Rinderpest outbreaks have also contributed in population loss.<sup id="cite_ref-iucn_28-4" class="reference"><a href="#cite_note-iucn-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> This species is currently of <a href="/wiki/Least_Concern" class="mw-redirect" title="Least Concern">Least Concern</a>, and the population is considered stable by both the IUCN and <a href="/wiki/CITES" title="CITES">CITES</a>.<sup id="cite_ref-antelope_15-1" class="reference"><a href="#cite_note-antelope-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> As of 1999, the total population of the nyala was around 32,000 individuals. More recent estimates show that South Africa has at least 30,000 nyalas, with 25,000 in KwaZulu-Natal. There are now more than 1,000 on protected areas and ranches in Eswatini. In Mozambique there are not more than 3,000, in Zimbabwe over 1,000, and numbers in Malawi have fallen from 3,000 to about 1,500. Namibia has the smallest population, at about 250.<sup id="cite_ref-iucn_28-5" class="reference"><a href="#cite_note-iucn-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p><p>Today over 80% of the total population is protected in national parks and sanctuaries, mostly in South African protected areas. In South Africa there is a high demand for adult males as game trophies.<sup id="cite_ref-iucn_28-6" class="reference"><a href="#cite_note-iucn-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=11" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width reflist-columns-2"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text">IUCN SSC Antelope Specialist Group. 2016. <i>Tragelaphus angasii</i> (errata version published in 2017). The IUCN Red List of Threatened Species 2016: e.T22052A115165681. <a rel="nofollow" class="external free" href="https://dx.doi.org/10.2305/IUCN.UK.2016-3.RLTS.T22052A50196443.en">https://dx.doi.org/10.2305/IUCN.UK.2016-3.RLTS.T22052A50196443.en</a>. Accessed on 01 May 2022.</span> </li> <li id="cite_note-huffman-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-huffman_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-huffman_2-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-huffman_2-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-huffman_2-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-huffman_2-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-huffman_2-5"><sup><i><b>f</b></i></sup></a> <a href="#cite_ref-huffman_2-6"><sup><i><b>g</b></i></sup></a> <a href="#cite_ref-huffman_2-7"><sup><i><b>h</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFHuffman" class="citation web cs1">Huffman, B. <a rel="nofollow" class="external text" href="http://www.ultimateungulate.com/artiodactyla/Tragelaphus_angasii.html">"Nyala"</a>. Ultimate Ungulate.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Nyala&rft.pub=Ultimate+Ungulate&rft.aulast=Huffman&rft.aufirst=B.&rft_id=http%3A%2F%2Fwww.ultimateungulate.com%2Fartiodactyla%2FTragelaphus_angasii.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANyala" class="Z3988"></span></span> </li> <li id="cite_note-burton-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-burton_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-burton_3-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-burton_3-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-burton_3-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-burton_3-4"><sup><i><b>e</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBurtonBurton,_R.2002" class="citation book cs1">Burton, M.; Burton, R. (2002). <a rel="nofollow" class="external text" href="https://archive.org/details/internationalwil03burt0">"Nyala"</a>. <i>International wildlife encyclopedia</i> (3rd ed.). New York: Marshall Cavendish. pp. 1765–6. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-7614-7269-8" title="Special:BookSources/978-0-7614-7269-8"><bdi>978-0-7614-7269-8</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Nyala&rft.btitle=International+wildlife+encyclopedia&rft.place=New+York&rft.pages=1765-6&rft.edition=3rd&rft.pub=Marshall+Cavendish&rft.date=2002&rft.isbn=978-0-7614-7269-8&rft.aulast=Burton&rft.aufirst=M.&rft.au=Burton%2C+R.&rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Finternationalwil03burt0&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANyala" class="Z3988"></span></span> </li> <li id="cite_note-MSW3-4"><span class="mw-cite-backlink">^ <a href="#cite_ref-MSW3_4-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-MSW3_4-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-MSW3_4-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGrubb2005" class="citation book cs1"><a href="/wiki/Peter_Grubb_(zoologist)" title="Peter Grubb (zoologist)">Grubb, P.</a> (2005). <a rel="nofollow" class="external text" href="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=14200727">"Order Artiodactyla"</a>. In <a href="/wiki/Don_E._Wilson" title="Don E. Wilson">Wilson, D.E.</a>; Reeder, D.M (eds.). <a rel="nofollow" class="external text" href="http://www.google.com/books?id=JgAMbNSt8ikC&pg=PA126"><i>Mammal Species of the World: A Taxonomic and Geographic Reference</i></a> (3rd ed.). 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New York: American Museum of Natural History: 323. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<a rel="nofollow" class="external text" href="https://hdl.handle.net/2246%2F613">2246/613</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Bulletin+of+the+American+Museum+of+Natural+History&rft.atitle=The+natural+history+of+nyala%2C+Tragelaphus+angasi+%28Mammalia%2C+Bovidae%29+in+Mozambique&rft.volume=155&rft.pages=323&rft.date=1975&rft_id=info%3Ahdl%2F2246%2F613&rft.au=Tello%2C+Jose+L.P.L.&rft.au=Van+Gelder%2C+Richard+G.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANyala" class="Z3988"></span></span> </li> <li id="cite_note-30"><span class="mw-cite-backlink"><b><a href="#cite_ref-30">^</a></b></span> <span class="reference-text">Alden, P. C.; Estes, R. D.; Schlitter, D.; McBride, B. (1995) <i>National Audubon Society Field Guide to African Wildlife</i>. Chanticleer Press.</span> </li> <li id="cite_note-planet-31"><span class="mw-cite-backlink">^ <a href="#cite_ref-planet_31-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-planet_31-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFitzpatrickArmstrong,_K.2006" class="citation book cs1">Fitzpatrick, M.; Armstrong, K. (2006). <i>South Africa, Lesotho & Swaziland</i>. 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"Reproduction in the Nyala (<i>Tragelaphus angasi</i>) (Mammalia: Ungulata)". <i>Journal of Zoology</i>. <b>204</b> (1): 129–42. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fj.1469-7998.1984.tb02366.x">10.1111/j.1469-7998.1984.tb02366.x</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Zoology&rft.atitle=Reproduction+in+the+Nyala+%28Tragelaphus+angasi%29+%28Mammalia%3A+Ungulata%29&rft.volume=204&rft.issue=1&rft.pages=129-42&rft.date=2009-08-20&rft_id=info%3Adoi%2F10.1111%2Fj.1469-7998.1984.tb02366.x&rft.aulast=Anderson&rft.aufirst=J.+L.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANyala" class="Z3988"></span></span> </li> <li id="cite_note-kfi-37"><span class="mw-cite-backlink"><b><a href="#cite_ref-kfi_37-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKarns2008" class="citation web cs1">Karns, G. 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"Variation in body condition of impala and nyala in relation to social status and reproduction". <i>South African Journal of Wildlife Research</i>. <b>23</b> (2): 36–8. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0379-4369">0379-4369</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=South+African+Journal+of+Wildlife+Research&rft.atitle=Variation+in+body+condition+of+impala+and+nyala+in+relation+to+social+status+and+reproduction&rft.volume=23&rft.issue=2&rft.pages=36-8&rft.date=1993&rft.issn=0379-4369&rft.au=A.F.+V.+Rooyen&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANyala" class="Z3988"></span></span> </li> <li id="cite_note-grobler-39"><span class="mw-cite-backlink"><b><a href="#cite_ref-grobler_39-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGroblerPretorius,_D.M.Botha,_K.Kotze,_A.2005" class="citation journal cs1 cs1-prop-foreign-lang-source">Grobler, J.P.; Pretorius, D.M.; Botha, K.; Kotze, A.; Hallerman, E.M.; Jansen Van Vuuren, B. 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"An exploratory analysis of geographic genetic variation in southern African nyala (<i>Tragelaphus angasii</i>)". <i>Mammalian Biology - Zeitschrift für Säugetierkunde</i> (in English and German). <b>70</b> (5): 291–9. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2005MamBi..70..291G">2005MamBi..70..291G</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.mambio.2005.01.001">10.1016/j.mambio.2005.01.001</a>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/10019.1%2F117692">10019.1/117692</a></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Mammalian+Biology+-+Zeitschrift+f%C3%BCr+S%C3%A4ugetierkunde&rft.atitle=An+exploratory+analysis+of+geographic+genetic+variation+in+southern+African+nyala+%28Tragelaphus+angasii%29&rft.volume=70&rft.issue=5&rft.pages=291-9&rft.date=2005-09-01&rft_id=info%3Ahdl%2F10019.1%2F117692&rft_id=info%3Adoi%2F10.1016%2Fj.mambio.2005.01.001&rft_id=info%3Abibcode%2F2005MamBi..70..291G&rft.aulast=Grobler&rft.aufirst=J.P.&rft.au=Pretorius%2C+D.M.&rft.au=Botha%2C+K.&rft.au=Kotze%2C+A.&rft.au=Hallerman%2C+E.M.&rft.au=Jansen+Van+Vuuren%2C+B.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANyala" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nyala&action=edit&section=12" title="Edit section: External 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.navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Extant_Artiodactyla_species" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" 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mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Ruminantia" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Ruminant" title="Ruminant">Ruminantia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Antilocapridae" title="Antilocapridae">Antilocapridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Antilocapra" scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Pronghorn" title="Pronghorn">Antilocapra</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pronghorn" title="Pronghorn">Pronghorn (<i>A. americana</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Giraffidae" title="Giraffidae">Giraffidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Okapi" title="Okapi">Okapia</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Okapi" title="Okapi">Okapi (<i>O. johnstoni</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Giraffe" title="Giraffe">Giraffa</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Northern_giraffe" title="Northern giraffe">Northern giraffe (<i>G. camelopardalis</i>)</a></li> <li><a href="/wiki/Southern_giraffe" title="Southern giraffe">Southern giraffe (<i>G. giraffa</i>)</a></li> <li><a href="/wiki/Reticulated_giraffe" title="Reticulated giraffe">Reticulated giraffe (<i>G. reticulata</i>)</a></li> <li><a href="/wiki/Masai_giraffe" title="Masai giraffe">Masai giraffe (<i>G. tippelskirchi</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Moschidae" title="Moschidae">Moschidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Moschus" scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Musk_deer" title="Musk deer">Moschus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anhui_musk_deer" title="Anhui musk deer">Anhui musk deer (<i>M. anhuiensis</i>)</a></li> <li><a href="/wiki/Dwarf_musk_deer" title="Dwarf musk deer">Dwarf musk deer (<i>M. berezovskii</i>)</a></li> <li><a href="/wiki/Alpine_musk_deer" title="Alpine musk deer">Alpine musk deer (<i>M. chrysogaster</i>)</a></li> <li><a href="/wiki/Kashmir_musk_deer" title="Kashmir musk deer">Kashmir musk deer (<i>M. cupreus</i>)</a></li> <li><a href="/wiki/Black_musk_deer" title="Black musk deer">Black musk deer (<i>M. fuscus</i>)</a></li> <li><a href="/wiki/White-bellied_musk_deer" title="White-bellied musk deer">Himalayan musk deer (<i>M. leucogaster</i>)</a></li> <li><a href="/wiki/Siberian_musk_deer" title="Siberian musk deer">Siberian musk deer (<i>M. moschiferus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Chevrotain" title="Chevrotain">Tragulidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Water_chevrotain" title="Water chevrotain">Hyemoschus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Water_chevrotain" title="Water chevrotain">Water chevrotain (<i>H. aquaticus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Moschiola" title="Moschiola">Moschiola</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Indian_spotted_chevrotain" title="Indian spotted chevrotain">Indian spotted chevrotain (<i>M. indica</i>)</a></li> <li><a href="/wiki/Yellow-striped_chevrotain" title="Yellow-striped chevrotain">Yellow-striped chevrotain (<i>M. kathygre</i>)</a></li> <li><a href="/wiki/Sri_Lankan_spotted_chevrotain" title="Sri Lankan spotted chevrotain">Sri Lankan spotted chevrotain (<i>M. meminna</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Tragulus" title="Tragulus">Tragulus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Java_mouse-deer" title="Java mouse-deer">Java mouse-deer (<i>T. javanicus</i>)</a></li> <li><a href="/wiki/Lesser_mouse-deer" title="Lesser mouse-deer">Lesser mouse-deer (<i>T. kanchil</i>)</a></li> <li><a href="/wiki/Greater_mouse-deer" title="Greater mouse-deer">Greater mouse-deer (<i>T. napu</i>)</a></li> <li><a href="/wiki/Philippine_mouse-deer" title="Philippine mouse-deer">Philippine mouse-deer (<i>T. nigricans</i>)</a></li> <li><a href="/wiki/Vietnam_mouse-deer" title="Vietnam mouse-deer">Vietnam mouse-deer (<i>T. versicolor</i>)</a></li> <li><a href="/wiki/Williamson%27s_mouse-deer" title="Williamson's mouse-deer">Williamson's mouse-deer (<i>T. williamsoni</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Deer" title="Deer">Cervidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em">Large family listed below</div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Bovidae" title="Bovidae">Bovidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em">Large family listed below</div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Family_Cervidae" style="font-size:114%;margin:0 4em"><span class="nobold">Family <a href="/wiki/Deer" title="Deer">Cervidae</a></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Cervinae" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cervinae" title="Cervinae">Cervinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Muntjac" title="Muntjac">Muntiacus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bornean_yellow_muntjac" title="Bornean yellow muntjac">Bornean yellow muntjac (<i>M. atherodes</i>)</a></li> <li><a href="/wiki/Hairy-fronted_muntjac" title="Hairy-fronted muntjac">Hairy-fronted muntjac (<i>M. crinifrons</i>)</a></li> <li><a href="/wiki/Fea%27s_muntjac" title="Fea's muntjac">Fea's muntjac (<i>M. feae</i>)</a></li> <li><a href="/wiki/Gongshan_muntjac" title="Gongshan muntjac">Gongshan muntjac (<i>M. gongshanensis</i>)</a></li> <li><a href="/wiki/Sumatran_muntjac" title="Sumatran muntjac">Sumatran muntjac (<i>M. montanus</i>)</a></li> <li><a href="/wiki/Southern_red_muntjac" title="Southern red muntjac">Southern red muntjac (<i>M. muntjak</i>)</a></li> <li><a href="/wiki/Pu_Hoat_muntjac" title="Pu Hoat muntjac">Pu Hoat muntjac (<i>M. puhoatensis</i>)</a></li> <li><a href="/wiki/Leaf_muntjac" title="Leaf muntjac">Leaf muntjac (<i>M. putaoensis</i>)</a></li> <li><a href="/wiki/Reeves%27s_muntjac" title="Reeves's muntjac">Reeves's muntjac (<i>M. reevesi</i>)</a></li> <li><a href="/wiki/Roosevelt%27s_muntjac" title="Roosevelt's muntjac">Roosevelt's muntjac (<i>M. rooseveltorum</i>)</a></li> <li><a href="/wiki/Truong_Son_muntjac" title="Truong Son muntjac">Truong Son muntjac (<i>M. truongsonensis</i>)</a></li> <li><a href="/wiki/Northern_red_muntjac" title="Northern red muntjac">Northern red muntjac (<i>M. vaginalis</i>)</a></li> <li><a href="/wiki/Giant_muntjac" title="Giant muntjac">Giant muntjac (<i>M. vuquangensis</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Tufted_deer" title="Tufted deer">Elaphodus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Tufted_deer" title="Tufted deer">Tufted deer (<i>E. cephalophus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Dama_(genus)" class="mw-redirect" title="Dama (genus)">Dama</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/European_fallow_deer" title="European fallow deer">European fallow deer (<i>D. dama</i>)</a></li> <li><a href="/wiki/Persian_fallow_deer" title="Persian fallow deer">Persian fallow deer (<i>D. mesopotamica</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Axis_(genus)" title="Axis (genus)">Axis</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Chital" title="Chital">Chital (<i>A. axis</i>)</a></li> <li><a href="/wiki/Calamian_deer" title="Calamian deer">Calamian deer (<i>A. calamianensis</i>)</a></li> <li><a href="/wiki/Bawean_deer" title="Bawean deer">Bawean deer (<i>A. kuhlii</i>)</a></li> <li><a href="/wiki/Indian_hog_deer" title="Indian hog deer">Hog deer (<i>A. porcinus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Rucervus" title="Rucervus">Rucervus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Barasingha" title="Barasingha">Barasingha (<i>R. duvaucelii</i>)</a></li> <li><a href="/wiki/Eld%27s_deer" title="Eld's deer">Eld's deer (<i>R. eldii</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Elaphurus" title="Elaphurus">Elaphurus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/P%C3%A8re_David%27s_deer" title="Père David's deer">Père David's deer (<i>E. davidianus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Rusa_(genus)" title="Rusa (genus)">Rusa</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Visayan_spotted_deer" title="Visayan spotted deer">Visayan spotted deer (<i>R. alfredi</i>)</a></li> <li><a href="/wiki/Philippine_deer" title="Philippine deer">Philippine sambar (<i>R. mariannus</i>)</a></li> <li><a href="/wiki/Javan_rusa" title="Javan rusa">Rusa deer (<i>R. timorensis</i>)</a></li> <li><a href="/wiki/Sambar_deer" title="Sambar deer">Sambar (<i>R. unicolor</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Cervus" title="Cervus">Cervus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Thorold%27s_deer" title="Thorold's deer">Thorold's deer (<i>C. albirostris</i>)</a></li> <li><a href="/wiki/Red_deer" title="Red deer">Red deer (<i>C. elaphus</i>)</a></li> <li><a href="/wiki/Elk" title="Elk">Elk (<i>C. canadensis</i>)</a></li> <li><a href="/wiki/Central_Asian_red_deer" title="Central Asian red deer">Central Asian red deer (<i>C. hanglu</i>)</a></li> <li><a href="/wiki/Sika_deer" title="Sika deer">Sika deer (<i>C. nippon</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Capreolinae" title="Capreolinae">Capreolinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Moose" title="Moose">Alces</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Moose" title="Moose">Moose (<i>A. alces</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Water_deer" title="Water deer">Hydropotes</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Water_deer" title="Water deer">Water deer (<i>H. inermis</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Capreolus" title="Capreolus">Capreolus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Roe_deer" title="Roe deer">European roe deer (<i>C. capreolus</i>)</a></li> <li><a href="/wiki/Siberian_roe_deer" title="Siberian roe deer">Siberian roe deer (<i>C. pygargus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Reindeer" title="Reindeer">Rangifer</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Reindeer" title="Reindeer">Reindeer (<i>R. tarandus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Hippocamelus" title="Hippocamelus">Hippocamelus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Taruca" title="Taruca">Taruca (<i>H. antisensis</i>)</a></li> <li><a href="/wiki/South_Andean_deer" title="South Andean deer">South Andean deer (<i>H. bisulcus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Brocket_deer" title="Brocket deer">Mazama</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Red_brocket" title="Red brocket">Red brocket (<i>M. americana</i>)</a></li> <li><a href="/wiki/Small_red_brocket" title="Small red brocket">Small red brocket (<i>M. bororo</i>)</a></li> <li><a href="/wiki/Merida_brocket" class="mw-redirect" title="Merida brocket">Merida brocket (<i>M. bricenii</i>)</a></li> <li><a href="/wiki/Dwarf_brocket" title="Dwarf brocket">Dwarf brocket (<i>M. chunyi</i>)</a></li> <li><a href="/wiki/Gray_brocket" title="Gray brocket">Gray brocket (<i>M. gouazoubira</i>)</a></li> <li><a href="/wiki/Pygmy_brocket" title="Pygmy brocket">Pygmy brocket (<i>M. nana</i>)</a></li> <li><a href="/wiki/Amazonian_brown_brocket" title="Amazonian brown brocket">Amazonian brown brocket (<i>M. nemorivaga</i>)</a></li> <li><a href="/wiki/Little_red_brocket" title="Little red brocket">Little red brocket (<i>M. rufina</i>)</a></li> <li><a href="/wiki/Central_American_red_brocket" title="Central American red brocket">Central American red brocket (<i>M. temama</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Pampas_deer" title="Pampas deer">Ozotoceros</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pampas_deer" title="Pampas deer">Pampas deer (<i>O. bezoarticus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Marsh_deer" title="Marsh deer">Blastocerus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Marsh_deer" title="Marsh deer">Marsh deer (<i>B. dichotomus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Pudu" title="Pudu">Pudu</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Northern_pudu" title="Northern pudu">Northern pudu (<i>P. mephistophiles</i>)</a>?</li> <li><a href="/wiki/Southern_pudu" title="Southern pudu">Southern pudu (<i>P. pudu</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Pudella" class="mw-redirect" title="Pudella">Pudella</a></i>?</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pudella_carlae" title="Pudella carlae">Peruvian Yungas pudu (<i>P. carlae</i>)</a></li> <li><a href="/wiki/Northern_pudu" title="Northern pudu">Northern pudu (<i>P. mephistophiles</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Odocoileus" title="Odocoileus">Odocoileus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mule_deer" title="Mule deer">Mule deer (<i>O. hemionus</i>)</a></li> <li><a href="/wiki/Yucatan_brown_brocket" title="Yucatan brown brocket">Yucatan brown brocket (<i>O. pandora</i>)</a></li> <li><a href="/wiki/White-tailed_deer" title="White-tailed deer">White-tailed deer (<i>O. virginianus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Family_Bovidae" style="font-size:114%;margin:0 4em"><span class="nobold">Family <a href="/wiki/Bovidae" title="Bovidae">Bovidae</a></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grazing_antelope" title="Grazing antelope">Hippotraginae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Hippotragus" title="Hippotragus">Hippotragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Roan_antelope" title="Roan antelope">Roan antelope (<i>H. equinus</i>)</a></li> <li><a href="/wiki/Sable_antelope" title="Sable antelope">Sable antelope (<i>H. niger</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Oryx" title="Oryx">Oryx</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/East_African_oryx" title="East African oryx">East African oryx (<i>O. beisa</i>)</a></li> <li><a href="/wiki/Scimitar_oryx" title="Scimitar oryx">Scimitar oryx (<i>O. dammah</i>)</a></li> <li><a href="/wiki/Gemsbok" title="Gemsbok">Gemsbok (<i>O. gazella</i>)</a></li> <li><a href="/wiki/Arabian_oryx" title="Arabian oryx">Arabian oryx (<i>O. leucoryx</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Addax" title="Addax">Addax</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Addax" title="Addax">Addax (<i>A. nasomaculatus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Reduncinae" title="Reduncinae">Reduncinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Kobus_(antelope)" title="Kobus (antelope)">Kobus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Waterbuck" title="Waterbuck">Waterbuck (<i>K. ellipsiprymnus</i>)</a></li> <li><a href="/wiki/Kob" title="Kob">Kob (<i>K. kob</i>)</a></li> <li><a href="/wiki/Lechwe" title="Lechwe">Lechwe (<i>K. leche</i>)</a></li> <li><a href="/wiki/Nile_lechwe" title="Nile lechwe">Nile lechwe (<i>K. megaceros</i>)</a></li> <li><a href="/wiki/Puku" title="Puku">Puku (<i>K. vardonii</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Reedbuck" title="Reedbuck">Redunca</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Southern_reedbuck" title="Southern reedbuck">Southern reedbuck (<i>R. arundinum</i>)</a></li> <li><a href="/wiki/Mountain_reedbuck" title="Mountain reedbuck">Mountain reedbuck (<i>R. fulvorufula</i>)</a></li> <li><a href="/wiki/Bohor_reedbuck" title="Bohor reedbuck">Bohor reedbuck (<i>R. redunca</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Impala" title="Impala">Aepycerotinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Aepyceros" scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Impala" title="Impala">Aepyceros</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Impala" title="Impala">Impala (<i>A. melampus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grey_rhebok" title="Grey rhebok">Peleinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pelea" scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Grey_rhebok" title="Grey rhebok">Pelea</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Grey_rhebok" title="Grey rhebok">Grey rhebok (<i>P. capreolus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Alcelaphinae" title="Alcelaphinae">Alcelaphinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Hirola" title="Hirola">Beatragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hirola" title="Hirola">Hirola (<i>B. hunteri</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Damaliscus" title="Damaliscus">Damaliscus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Damaliscus_lunatus" title="Damaliscus lunatus">Common tsessebe (<i>D. lunatus</i>)</a></li> <li><a href="/wiki/Bontebok" title="Bontebok">Bontebok (<i>D. pygargus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Hartebeest" title="Hartebeest">Alcelaphus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hartebeest" title="Hartebeest">Hartebeest (<i>A. buselaphus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Wildebeest" title="Wildebeest">Connochaetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Black_wildebeest" title="Black wildebeest">Black wildebeest (<i>C. gnou</i>)</a></li> <li><a href="/wiki/Blue_wildebeest" title="Blue wildebeest">Blue wildebeest (<i>C. taurinus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tibetan_antelope" title="Tibetan antelope">Pantholopinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pantholops" scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Tibetan_antelope" title="Tibetan antelope">Pantholops</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Tibetan_antelope" title="Tibetan antelope">Tibetan antelope (<i>P. hodgsonii</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Caprinae" title="Caprinae">Caprinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em">Large subfamily listed below</div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Bovinae" title="Bovinae">Bovinae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em">Large subfamily listed below</div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Antilopinae" title="Antilopinae">Antilopinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em">Large subfamily listed below</div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Family_Bovidae_(subfamily_Caprinae)" style="font-size:114%;margin:0 4em"><span class="nobold">Family <a href="/wiki/Bovidae" title="Bovidae">Bovidae</a> (subfamily <a href="/wiki/Caprinae" title="Caprinae">Caprinae</a>)</span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Barbary_sheep" title="Barbary sheep">Ammotragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Barbary_sheep" title="Barbary sheep">Barbary sheep (<i>A. lervia</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Arabian_tahr" title="Arabian tahr">Arabitragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arabian_tahr" title="Arabian tahr">Arabian tahr (<i>A. jayakari</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Takin" title="Takin">Budorcas</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Takin" title="Takin">Takin (<i>B. taxicolor</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Capra_(genus)" title="Capra (genus)">Capra</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Wild_goat" title="Wild goat">Wild goat (<i>C. aegagrus</i>)</a></li> <li><a href="/wiki/West_Caucasian_tur" title="West Caucasian tur">West Caucasian tur (<i>C. caucasia</i>)</a></li> <li><a href="/wiki/East_Caucasian_tur" title="East Caucasian tur">East Caucasian tur (<i>C. cylindricornis</i>)</a></li> <li><a href="/wiki/Markhor" title="Markhor">Markhor (<i>C. falconeri</i>)</a></li> <li><a href="/wiki/Goat" title="Goat">Domestic goat (<i>C. hircus</i>)</a></li> <li><a href="/wiki/Alpine_ibex" title="Alpine ibex">Alpine ibex (<i>C. ibex</i>)</a></li> <li><a href="/wiki/Nubian_ibex" title="Nubian ibex">Nubian ibex (<i>C. nubiana</i>)</a></li> <li><a href="/wiki/Iberian_ibex" title="Iberian ibex">Iberian ibex (<i>C. pyrenaica</i>)</a></li> <li><a href="/wiki/Siberian_ibex" title="Siberian ibex">Siberian ibex (<i>C. sibirica</i>)</a></li> <li><a href="/wiki/Walia_ibex" title="Walia ibex">Walia ibex (<i>C. walie</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Serow" title="Serow">Capricornis</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Japanese_serow" title="Japanese serow">Japanese serow (<i>C. crispus</i>)</a></li> <li><a href="/wiki/Red_serow" title="Red serow">Red serow (<i>C. rubidus</i>)</a></li> <li><a href="/wiki/Mainland_serow" title="Mainland serow">Mainland serow (<i>C. sumatraensis</i>)</a></li> <li><a href="/wiki/Taiwan_serow" title="Taiwan serow">Taiwan serow (<i>C. swinhoei</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Tahr" title="Tahr">Hemitragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Himalayan_tahr" title="Himalayan tahr">Himalayan tahr (<i>H. jemlahicus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Goral" title="Goral">Naemorhedus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Red_goral" title="Red goral">Red goral (<i>N. baileyi</i>)</a></li> <li><a href="/wiki/Long-tailed_goral" title="Long-tailed goral">Long-tailed goral (<i>N. caudatus</i>)</a></li> <li><a href="/wiki/Himalayan_goral" title="Himalayan goral">Himalayan goral (<i>N. goral</i>)</a></li> <li><a href="/wiki/Chinese_goral" title="Chinese goral">Chinese goral (<i>N. griseus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Mountain_goat" title="Mountain goat">Oreamnos</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mountain_goat" title="Mountain goat">Mountain goat (<i>O. americanus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Muskox" title="Muskox">Ovibos</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Muskox" title="Muskox">Muskox (<i>O. moschatus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Nilgiri_tahr" title="Nilgiri tahr">Nilgiritragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nilgiri_tahr" title="Nilgiri tahr">Nilgiri tahr (<i>N. hylocrius</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Ovis" title="Ovis">Ovis</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Argali" title="Argali">Argali (<i>O. ammon</i>)</a></li> <li><a href="/wiki/Sheep" title="Sheep">Domestic sheep (<i>O. aries</i>)</a></li> <li><a href="/wiki/Bighorn_sheep" title="Bighorn sheep">Bighorn sheep (<i>O. canadensis</i>)</a></li> <li><a href="/wiki/Dall_sheep" title="Dall sheep">Dall sheep (<i>O. dalli</i>)</a></li> <li><a href="/wiki/Mouflon" title="Mouflon">Mouflon (<i>O. gmelini</i>)</a></li> <li><a href="/wiki/Snow_sheep" title="Snow sheep">Snow sheep (<i>O. nivicola</i>)</a></li> <li><a href="/wiki/Urial" title="Urial">Urial (<i>O. vignei</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Pseudois" class="mw-redirect" title="Pseudois">Pseudois</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bharal" title="Bharal">Bharal (<i>P. nayaur</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><i><a href="/wiki/Rupicapra" title="Rupicapra">Rupicapra</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pyrenean_chamois" title="Pyrenean chamois">Pyrenean chamois (<i>R. pyrenaica</i>)</a></li> <li><a href="/wiki/Chamois" title="Chamois">Chamois (<i>R. rupicapra</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Family_Bovidae_(subfamily_Bovinae)" style="font-size:114%;margin:0 4em"><span class="nobold">Family <a href="/wiki/Bovidae" title="Bovidae">Bovidae</a> (subfamily <a href="/wiki/Bovinae" title="Bovinae">Bovinae</a>)</span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Boselaphini" title="Boselaphini">Boselaphini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Four-horned_antelope" title="Four-horned antelope">Tetracerus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Four-horned_antelope" title="Four-horned antelope">Four-horned antelope (<i>T. quadricornis</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Nilgai" title="Nilgai">Boselaphus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nilgai" title="Nilgai">Nilgai (<i>B. tragocamelus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Bovini" title="Bovini">Bovini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Bubalus" title="Bubalus">Bubalus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Wild_water_buffalo" title="Wild water buffalo">Wild water buffalo (<i>B. arnee</i>)</a></li> <li><a href="/wiki/Water_buffalo" title="Water buffalo">Domestic water buffalo (<i>B. bubalis</i>)</a></li> <li><a href="/wiki/Lowland_anoa" title="Lowland anoa">Lowland anoa (<i>B. depressicornis</i>)</a></li> <li><a href="/wiki/Tamaraw" title="Tamaraw">Tamaraw (<i>B. mindorensis</i>)</a></li> <li><a href="/wiki/Mountain_anoa" title="Mountain anoa">Mountain anoa (<i>B. quarlesi</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Bos" title="Bos">Bos</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/American_bison" title="American bison">American bison (<i>B. bison</i>)</a></li> <li><a href="/wiki/European_bison" title="European bison">European bison (<i>B. bonasus</i>)</a></li> <li><a href="/wiki/Bali_cattle" title="Bali cattle">Bali cattle (<i>B. domesticus</i>)</a></li> <li><a href="/wiki/Gayal" title="Gayal">Gayal (<i>B. frontalis</i>)</a></li> <li><a href="/wiki/Gaur" title="Gaur">Gaur (<i>B. gaurus</i>)</a></li> <li><a href="/wiki/Yak" title="Yak">Domestic yak (<i>B. grunniens</i>)</a></li> <li><a href="/wiki/Zebu" title="Zebu">Zebu (<i>B. indicus</i>)</a></li> <li><a href="/wiki/Banteng" title="Banteng">Banteng (<i>B. javanicus</i>)</a></li> <li><a href="/wiki/Wild_yak" title="Wild yak">Wild yak (<i>B. mutus</i>)</a></li> <li><a href="/wiki/Cattle" title="Cattle">Cattle (<i>B. taurus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Saola" title="Saola">Pseudoryx</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Saola" title="Saola">Saola (<i>P. nghetinhensis</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/African_buffalo" title="African buffalo">Syncerus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/African_buffalo" title="African buffalo">African buffalo (<i>S. caffer</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tragelaphini" title="Tragelaphini">Tragelaphini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Tragelaphus" title="Tragelaphus">Tragelaphus</a></i><br /><small><span style="color:#696969"><span class="nobold">(including <a href="/wiki/Kudu" title="Kudu">kudus</a>)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Nyala (<i>T. angasii</i>)</a></li> <li><a href="/wiki/Mountain_nyala" title="Mountain nyala">Mountain nyala (<i>T. buxtoni</i>)</a></li> <li><a href="/wiki/Bongo_(antelope)" title="Bongo (antelope)">Bongo (<i>T. eurycerus</i>)</a></li> <li><a href="/wiki/Lesser_kudu" title="Lesser kudu">Lesser kudu (<i>T. imberbis</i>)</a></li> <li><a href="/wiki/Harnessed_bushbuck" title="Harnessed bushbuck">Harnessed bushbuck (<i>T. scriptus</i>)</a></li> <li><a href="/wiki/Sitatunga" title="Sitatunga">Sitatunga (<i>T. spekeii</i>)</a></li> <li><a href="/wiki/Greater_kudu" title="Greater kudu">Greater kudu (<i>T. strepsiceros</i>)</a></li> <li><a href="/wiki/Cape_bushbuck" title="Cape bushbuck">Cape bushbuck (<i>T. sylvaticus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Taurotragus" title="Taurotragus">Taurotragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Giant_eland" title="Giant eland">Giant eland (<i>T. derbianus</i>)</a></li> <li><a href="/wiki/Common_eland" title="Common eland">Common eland (<i>T. oryx</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Family_Bovidae_(subfamily_Antilopinae)" style="font-size:114%;margin:0 4em"><span class="nobold">Family <a href="/wiki/Bovidae" title="Bovidae">Bovidae</a> (subfamily <a href="/wiki/Antilopinae" title="Antilopinae">Antilopinae</a>)</span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Antilopini" title="Antilopini">Antilopini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Dibatag" title="Dibatag">Ammodorcas</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dibatag" title="Dibatag">Dibatag (<i>A. clarkei</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Springbok" title="Springbok">Antidorcas</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Springbok" title="Springbok">Springbok (<i>A. marsupialis</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Blackbuck" title="Blackbuck">Antilope</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Blackbuck" title="Blackbuck">Blackbuck (<i>A. cervicapra</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Eudorcas" title="Eudorcas">Eudorcas</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mongalla_gazelle" title="Mongalla gazelle">Mongalla gazelle (<i>E. albonotata</i>)</a></li> <li><a href="/wiki/Red-fronted_gazelle" title="Red-fronted gazelle">Red-fronted gazelle (<i>E. rufifrons</i>)</a></li> <li><a href="/wiki/Thomson%27s_gazelle" title="Thomson's gazelle">Thomson's gazelle (<i>E. thomsonii</i>)</a></li> <li><a href="/wiki/Heuglin%27s_gazelle" title="Heuglin's gazelle">Heuglin's gazelle (<i>E. tilonura</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Gazelle" title="Gazelle">Gazella</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Chinkara" title="Chinkara">Chinkara (<i>G. bennettii</i>)</a></li> <li><a href="/wiki/Cuvier%27s_gazelle" title="Cuvier's gazelle">Cuvier's gazelle (<i>G. cuvieri</i>)</a></li> <li><a href="/wiki/Dorcas_gazelle" title="Dorcas gazelle">Dorcas gazelle (<i>G. dorcas</i>)</a></li> <li><a href="/wiki/Erlanger%27s_gazelle" title="Erlanger's gazelle">Erlanger's gazelle (<i>G. erlangeri</i>)</a></li> <li><a href="/wiki/Mountain_gazelle" title="Mountain gazelle">Mountain gazelle (<i>G. gazella</i>)</a></li> <li><a href="/wiki/Rhim_gazelle" title="Rhim gazelle">Rhim gazelle (<i>G. leptoceros</i>)</a></li> <li><a href="/wiki/Speke%27s_gazelle" title="Speke's gazelle">Speke's gazelle (<i>G. spekei</i>)</a></li> <li><a href="/wiki/Goitered_gazelle" title="Goitered gazelle">Goitered gazelle (<i>G. subgutturosa</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Gerenuk" title="Gerenuk">Litocranius</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Gerenuk" title="Gerenuk">Gerenuk (<i>L. walleri</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Nanger" title="Nanger">Nanger</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dama_gazelle" title="Dama gazelle">Dama gazelle (<i>N. dama</i>)</a></li> <li><a href="/wiki/Grant%27s_gazelle" title="Grant's gazelle">Grant's gazelle (<i>N. granti</i>)</a></li> <li><a href="/wiki/Bright%27s_gazelle" class="mw-redirect" title="Bright's gazelle">Bright's gazelle (<i>N. notatus</i>)</a></li> <li><a href="/wiki/Peter%27s_gazelle" class="mw-redirect" title="Peter's gazelle">Peter's gazelle (<i>N. petersii</i>)</a></li> <li><a href="/wiki/Soemmerring%27s_gazelle" title="Soemmerring's gazelle">Soemmerring's gazelle (<i>N. soemmerringii</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Procapra" title="Procapra">Procapra</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mongolian_gazelle" title="Mongolian gazelle">Mongolian gazelle (<i>P. gutturosa</i>)</a></li> <li><a href="/wiki/Goa_(antelope)" title="Goa (antelope)">Goa (<i>P. picticaudata</i>)</a></li> <li><a href="/wiki/Przewalski%27s_gazelle" title="Przewalski's gazelle">Przewalski's gazelle (<i>P. przewalskii</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Saigini</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Saiga" scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Saiga_antelope" title="Saiga antelope">Saiga</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Saiga_antelope" title="Saiga antelope">Saiga antelope (<i>S. tatarica</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Neotragini" title="Neotragini">Neotragini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Beira_(antelope)" title="Beira (antelope)">Dorcatragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Beira_(antelope)" title="Beira (antelope)">Beira (<i>D. megalotis</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Dik-dik" title="Dik-dik">Madoqua</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/G%C3%BCnther%27s_dik-dik" title="Günther's dik-dik">Günther's dik-dik (<i>M. guentheri</i>)</a></li> <li><a href="/wiki/Kirk%27s_dik-dik" title="Kirk's dik-dik">Kirk's dik-dik (<i>M. kirkii</i>)</a></li> <li><a href="/wiki/Silver_dik-dik" title="Silver dik-dik">Silver dik-dik (<i>M. piacentinii</i>)</a></li> <li><a href="/wiki/Salt%27s_dik-dik" title="Salt's dik-dik">Salt's dik-dik (<i>M. saltiana</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Neotragus" title="Neotragus">Neotragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Royal_antelope" title="Royal antelope">Royal antelope (<i>N. pygmaeus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Nesotragus" title="Nesotragus">Nesotragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bates%27s_pygmy_antelope" title="Bates's pygmy antelope">Bates' pygmy antelope (<i>N. batesi</i>)</a></li> <li><a href="/wiki/Suni" title="Suni">Suni (<i>N. moschatus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Klipspringer" title="Klipspringer">Oreotragus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Klipspringer" title="Klipspringer">Klipspringer (<i>O. oreotragus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Oribi" title="Oribi">Ourebia</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Oribi" title="Oribi">Oribi (<i>O. ourebi</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Raphicerus" title="Raphicerus">Raphicerus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Steenbok" title="Steenbok">Steenbok (<i>R. campestris</i>)</a></li> <li><a href="/wiki/Cape_grysbok" title="Cape grysbok">Cape grysbok (<i>R. melanotis</i>)</a></li> <li><a href="/wiki/Sharpe%27s_grysbok" title="Sharpe's grysbok">Sharpe's grysbok (<i>R. sharpei</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Duiker" title="Duiker">Cephalophini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Duiker" title="Duiker">Cephalophus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aders%27s_duiker" title="Aders's duiker">Aders's duiker (<i>C. adersi</i>)</a></li> <li><a href="/wiki/Brooke%27s_duiker" title="Brooke's duiker">Brooke's duiker (<i>C. brookei</i>)</a></li> <li><a href="/wiki/Peters%27s_duiker" title="Peters's duiker">Peters' duiker (<i>C. callipygus</i>)</a></li> <li><a href="/wiki/White-legged_duiker" title="White-legged duiker">White-legged duiker (<i>C. crusalbum</i>)</a></li> <li><a href="/wiki/Bay_duiker" title="Bay duiker">Bay duiker (<i>C. dorsalis</i>)</a></li> <li><a href="/wiki/Harvey%27s_duiker" title="Harvey's duiker">Harvey's duiker (<i>C. harveyi</i>)</a></li> <li><a href="/wiki/Jentink%27s_duiker" title="Jentink's duiker">Jentink's duiker (<i>C. jentinki</i>)</a></li> <li><a href="/wiki/White-bellied_duiker" title="White-bellied duiker">White-bellied duiker (<i>C. leucogaster</i>)</a></li> <li><a href="/wiki/Red_forest_duiker" title="Red forest duiker">Red forest duiker (<i>C. natalensis</i>)</a></li> <li><a href="/wiki/Black_duiker" title="Black duiker">Black duiker (<i>C. niger</i>)</a></li> <li><a href="/wiki/Black-fronted_duiker" title="Black-fronted duiker">Black-fronted duiker (<i>C. nigrifrons</i>)</a></li> <li><a href="/wiki/Ogilby%27s_duiker" title="Ogilby's duiker">Ogilby's duiker (<i>C. ogilbyi</i>)</a></li> <li><a href="/wiki/Ruwenzori_duiker" title="Ruwenzori duiker">Ruwenzori duiker (<i>C. rubidis</i>)</a></li> <li><a href="/wiki/Red-flanked_duiker" title="Red-flanked duiker">Red-flanked duiker (<i>C. rufilatus</i>)</a></li> <li><a href="/wiki/Yellow-backed_duiker" title="Yellow-backed duiker">Yellow-backed duiker (<i>C. silvicultor</i>)</a></li> <li><a href="/wiki/Abbott%27s_duiker" title="Abbott's duiker">Abbott's duiker (<i>C. spadix</i>)</a></li> <li><a href="/wiki/Weyns%27s_duiker" title="Weyns's duiker">Weyns's duiker (<i>C. weynsi</i>)</a></li> <li><a href="/wiki/Zebra_duiker" title="Zebra duiker">Zebra duiker (<i>C. zebra</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Duiker" title="Duiker">Philantomba</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Blue_duiker" title="Blue duiker">Blue duiker (<i>P. monticola</i>)</a></li> <li><a href="/wiki/Maxwell%27s_duiker" title="Maxwell's duiker">Maxwell's duiker (<i>P. maxwellii</i>)</a></li> <li><a href="/wiki/Walter%27s_duiker" title="Walter's duiker">Walter's duiker (<i>P. walteri</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Common_duiker" title="Common duiker">Sylvicapra</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Common_duiker" title="Common duiker">Common duiker (<i>S. grimmia</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Suina" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Suina" title="Suina">Suina</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Suidae" title="Suidae">Suidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9.5em"><i><a href="/wiki/Babirusa" title="Babirusa">Babyrousa</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Buru_babirusa" title="Buru babirusa">Buru babirusa (<i>B. babyrussa</i>)</a></li> <li><a href="/wiki/North_Sulawesi_babirusa" title="North Sulawesi babirusa">North Sulawesi babirusa (<i>B. celebensis</i>)</a></li> <li><a href="/wiki/Togian_babirusa" title="Togian babirusa">Togian babirusa (<i>B. togeanensis</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9.5em"><i><a href="/wiki/Giant_forest_hog" title="Giant forest hog">Hylochoerus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Giant_forest_hog" title="Giant forest hog">Giant forest hog (<i>H. meinertzhageni</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9.5em"><i><a href="/wiki/Phacochoerus" class="mw-redirect" title="Phacochoerus">Phacochoerus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Desert_warthog" title="Desert warthog">Desert warthog (<i>P. aethiopicus</i>)</a></li> <li><a href="/wiki/Common_warthog" title="Common warthog">Common warthog (<i>P. africanus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9.5em"><i><a href="/wiki/Pygmy_hog" title="Pygmy hog">Porcula</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pygmy_hog" title="Pygmy hog">Pygmy hog (<i>P. salvania</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9.5em"><i><a href="/wiki/Potamochoerus" title="Potamochoerus">Potamochoerus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bushpig" title="Bushpig">Bushpig (<i>P. larvatus</i>)</a></li> <li><a href="/wiki/Red_river_hog" title="Red river hog">Red river hog (<i>P. porcus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9.5em"><i><a href="/wiki/Sus_(genus)" title="Sus (genus)">Sus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Palawan_bearded_pig" title="Palawan bearded pig">Palawan bearded pig (<i>S. ahoenobarbus</i>)</a></li> <li><a href="/wiki/Bornean_bearded_pig" title="Bornean bearded pig">Bornean bearded pig (<i>S. barbatus</i>)</a></li> <li><a href="/wiki/Visayan_warty_pig" title="Visayan warty pig">Visayan warty pig (<i>S. cebifrons</i>)</a></li> <li><a href="/wiki/Celebes_warty_pig" title="Celebes warty pig">Celebes warty pig (<i>S. celebensis</i>)</a></li> <li><a href="/wiki/Pig" title="Pig">Domestic pig (<i>S. domesticus</i>)</a></li> <li><a href="/wiki/Flores_warty_pig" title="Flores warty pig">Flores warty pig (<i>S. heureni</i>)</a></li> <li><a href="/wiki/Oliver%27s_warty_pig" title="Oliver's warty pig">Oliver's warty pig (<i>S. oliveri</i>)</a></li> <li><a href="/wiki/Philippine_warty_pig" title="Philippine warty pig">Philippine warty pig (<i>S. philippensis</i>)</a></li> <li><a href="/wiki/Wild_boar" title="Wild boar">Wild boar (<i>S. scrofa</i>)</a></li> <li><a href="/wiki/Timor_warty_pig" title="Timor warty pig">Timor warty pig (<i>S. timoriensis</i>)</a></li> <li><a href="/wiki/Javan_warty_pig" title="Javan warty pig">Javan warty pig (<i>S. verrucosus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Peccary" title="Peccary">Tayassuidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9.5em"><i><a href="/wiki/White-lipped_peccary" title="White-lipped peccary">Tayassu</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/White-lipped_peccary" title="White-lipped peccary">White-lipped peccary (<i>T. pecari</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9.5em"><i><a href="/wiki/Chacoan_peccary" title="Chacoan peccary">Catagonus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Chacoan_peccary" title="Chacoan peccary">Chacoan peccary (<i>C. wagneri</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9.5em"><i><a href="/wiki/Collared_peccary" title="Collared peccary">Dicotyles</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Collared_peccary" title="Collared peccary">Collared peccary (<i>D. tajacu</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Tylopoda" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Tylopoda" title="Tylopoda">Tylopoda</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Camelidae" scope="row" class="navbox-group" style="width:6.5em"><a href="/wiki/Camelidae" title="Camelidae">Camelidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:6.5em"><i><a href="/wiki/Lama_(genus)" title="Lama (genus)">Lama</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Llama" title="Llama">Llama (<i>L. glama</i>)</a></li> <li><a href="/wiki/Guanaco" title="Guanaco">Guanaco (<i>L. guanicoe</i>)</a></li> <li><a href="/wiki/Alpaca" title="Alpaca">Alpaca (<i>L. pacos</i>)</a></li> <li><a href="/wiki/Vicu%C3%B1a" title="Vicuña">Vicuña (<i>L. vicugna</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:6.5em"><i><a href="/wiki/Camel" title="Camel">Camelus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bactrian_camel" title="Bactrian camel">Domestic Bactrian camel (<i>C. bactrianus</i>)</a></li> <li><a href="/wiki/Dromedary" title="Dromedary">Dromedary/Arabian camel (<i>C. dromedarius</i>)</a></li> <li><a href="/wiki/Wild_Bactrian_camel" title="Wild Bactrian camel">Wild Bactrian camel (<i>C. ferus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Whippomorpha" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Whippomorpha" title="Whippomorpha">Whippomorpha</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8.5em"><a href="/wiki/Hippopotamidae" title="Hippopotamidae">Hippopotamidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Hippopotamus_(genus)" title="Hippopotamus (genus)">Hippopotamus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hippopotamus" title="Hippopotamus">Hippopotamus (<i>H. amphibius</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><i><a href="/wiki/Pygmy_hippopotamus" title="Pygmy hippopotamus">Choeropsis</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pygmy_hippopotamus" title="Pygmy hippopotamus">Pygmy hippopotamus (<i>C. liberiensis</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.5em"><a href="/wiki/Cetacea" title="Cetacea">Cetacea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li>see <a href="/wiki/Template:Cetacea" title="Template:Cetacea">Cetacea</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Tragelaphus angasii</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q239032" class="extiw" title="wikidata:Q239032">Q239032</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Tragelaphus_angasii" class="extiw" title="wikispecies:Tragelaphus angasii">Tragelaphus angasii</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Animal_Diversity_Web" title="Animal Diversity Web">ADW</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://animaldiversity.org/accounts/Tragelaphus_angasii/">Tragelaphus_angasii</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Barcode_of_Life_Data_System" title="Barcode of Life Data System">BOLD</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.boldsystems.org/index.php/TaxBrowser_TaxonPage?taxid=462472">462472</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/57VQN">57VQN</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/1038786">1038786</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/5220181">5220181</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/42342">42342</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=11272279">11272279</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=625128">625128</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/IUCN_Red_List" title="IUCN Red List">IUCN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://apiv3.iucnredlist.org/api/v3/taxonredirect/22052">22052</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/American_Society_of_Mammalogists#Mammal_Diversity_Database" title="American Society of Mammalogists">MDD</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.mammaldiversity.org/taxon/1006275">1006275</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Mammal_Species_of_the_World" title="Mammal Species of the World">MSW</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?s=y&id=14200727">14200727</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=66437">66437</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Observation.org" title="Observation.org">Observation.org</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://observation.org/species/84551/">84551</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=40135">40135</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=149711">149711</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-label="Navbox" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a 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