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<li id="toc-Cerebellum" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Cerebellum"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.2</span> <span>Cerebellum</span> </div> </a> <ul id="toc-Cerebellum-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Amygdala" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Amygdala"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.3</span> <span>Amygdala</span> </div> </a> <ul id="toc-Amygdala-sublist" class="vector-toc-list"> <li id="toc-Memory_of_fear_conditioning" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Memory_of_fear_conditioning"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.3.1</span> <span>Memory of fear conditioning</span> </div> </a> <ul id="toc-Memory_of_fear_conditioning-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Memory_consolidation" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Memory_consolidation"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.3.2</span> <span>Memory consolidation</span> </div> </a> <ul id="toc-Memory_consolidation-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Basal_ganglia_and_motor_memory" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Basal_ganglia_and_motor_memory"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.4</span> <span>Basal ganglia and motor memory</span> </div> </a> <ul id="toc-Basal_ganglia_and_motor_memory-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Cortical_structures" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Cortical_structures"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Cortical structures</span> </div> </a> <button 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vector-toc-level-2"> <a class="vector-toc-link" href="#Parietal_lobe"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.3</span> <span>Parietal lobe</span> </div> </a> <ul id="toc-Parietal_lobe-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Occipital_lobe" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Occipital_lobe"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.4</span> <span>Occipital lobe</span> </div> </a> <ul id="toc-Occipital_lobe-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Damage_to_the_cortex" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Damage_to_the_cortex"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.5</span> <span>Damage to the cortex</span> </div> </a> <ul id="toc-Damage_to_the_cortex-sublist" class="vector-toc-list"> <li id="toc-Frontotemporal_lobar_degeneration_and_memory" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Frontotemporal_lobar_degeneration_and_memory"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.5.1</span> <span>Frontotemporal lobar degeneration and memory</span> </div> </a> <ul id="toc-Frontotemporal_lobar_degeneration_and_memory-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Parkinson's_disease_and_memory" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Parkinson's_disease_and_memory"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.5.2</span> <span>Parkinson's disease and memory</span> </div> </a> <ul id="toc-Parkinson's_disease_and_memory-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>References</span> </div> </a> <ul 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mw-heading2"><h2 id="Subcortical_structures">Subcortical structures</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=1" title="Edit section: Subcortical structures"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Hippocampus">Hippocampus</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=2" title="Edit section: Hippocampus"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:NIA_human_brain_drawing.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a6/NIA_human_brain_drawing.jpg/300px-NIA_human_brain_drawing.jpg" decoding="async" width="300" height="342" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a6/NIA_human_brain_drawing.jpg/450px-NIA_human_brain_drawing.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/a/a6/NIA_human_brain_drawing.jpg 2x" data-file-width="559" data-file-height="637" /></a><figcaption>The <a href="/wiki/Hippocampus" title="Hippocampus">hippocampus</a></figcaption></figure> <p>The <a href="/wiki/Hippocampus" title="Hippocampus">hippocampus</a> is a structure in the brain that has been associated with various <a href="/wiki/Memory" title="Memory">memory</a> functions. It is part of the <a href="/wiki/Limbic_system" title="Limbic system">limbic system</a>, and lies next to the medial temporal lobe. It is made up of two structures, the <b>Ammon's Horn</b>, and the <a href="/wiki/Dentate_gyrus" title="Dentate gyrus">Dentate gyrus</a>, each containing different types of <a href="/wiki/Cell_(biology)" title="Cell (biology)">cells</a>.<sup id="cite_ref-Kolb_1-0" class="reference"><a href="#cite_note-Kolb-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Cognitive_maps">Cognitive maps</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=3" title="Edit section: Cognitive maps"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>There is evidence that the hippocampus contains <a href="/wiki/Cognitive_map" title="Cognitive map">cognitive maps</a> in humans. In one study, single-cell recordings were taken from electrodes implanted in a rat's hippocampus, and it was found that certain <a href="/wiki/Neurons" class="mw-redirect" title="Neurons">neurons</a> responded strongly only when the rat was in certain locations. These cells are called <a href="/wiki/Place_cells" class="mw-redirect" title="Place cells">place cells</a>, and collections of these cells can be considered to be mental maps. Individual place cells do not only respond to one unique area only however, the patterns of activation of these cells overlap to form layered mental maps within the hippocampus. A good analogy is the example of the same television or computer screen pixels being used to light up any trillions of possible combinations to produce images, just as the place cells can be used in any multiple possible combinations to represent mental maps. The hippocampus' right side is more oriented towards responding to spatial aspects, whereas the left side is associated with other context information. Also, there is evidence that experience in building extensive mental maps, such as driving a city taxi for a long time (since this requires considerable memorization of routes), can increase the volume of one's hippocampus.<sup id="cite_ref-Ward_2-0" class="reference"><a href="#cite_note-Ward-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Encoding">Encoding</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=4" title="Edit section: Encoding"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Damage to the hippocampus and surrounding area can cause <a href="/wiki/Anterograde_amnesia" title="Anterograde amnesia">anterograde amnesia</a>, the inability to form new memories.<sup id="cite_ref-Mahut_3-0" class="reference"><a href="#cite_note-Mahut-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> This implies that the hippocampus is important not only for storing cognitive maps, but for <a href="/wiki/Encoding_(memory)" title="Encoding (memory)">encoding memories</a>. </p><p>The hippocampus is also involved in <a href="/wiki/Memory_consolidation" title="Memory consolidation">memory consolidation</a>, the slow process by which memories are converted from <a href="/wiki/Short_term_memory" class="mw-redirect" title="Short term memory">short</a> to <a href="/wiki/Long_term_memory" class="mw-redirect" title="Long term memory">long term memory</a>. This is supported by studies in which lesions are applied to rat hippocampi at different times after learning.<sup id="cite_ref-Ward_2-1" class="reference"><a href="#cite_note-Ward-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The process of consolidation may take up to a couple years. </p><p>It has also been found that it is possible to form new <a href="/wiki/Semantic_memories" class="mw-redirect" title="Semantic memories">semantic memories</a> without the hippocampus, but not <a href="/wiki/Episodic_memories" class="mw-redirect" title="Episodic memories">episodic memories</a>, which means that explicit descriptions of actual events (episodic) cannot be learned, but some meaning and knowledge is gained from experiences (semantic).<sup id="cite_ref-Ward_2-2" class="reference"><a href="#cite_note-Ward-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Cerebellum">Cerebellum</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=5" title="Edit section: Cerebellum"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Brain_bulbar_region.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7d/Brain_bulbar_region.svg/200px-Brain_bulbar_region.svg.png" decoding="async" width="200" height="203" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7d/Brain_bulbar_region.svg/300px-Brain_bulbar_region.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7d/Brain_bulbar_region.svg/400px-Brain_bulbar_region.svg.png 2x" data-file-width="295" data-file-height="299" /></a><figcaption>The <a href="/wiki/Cerebellum" title="Cerebellum">cerebellum</a></figcaption></figure> <p>The <a href="/wiki/Cerebellum" title="Cerebellum">cerebellum</a> ("little brain") is a structure located at the rear of the brain, near the <a href="/wiki/Spinal_cord" title="Spinal cord">spinal cord</a>. It looks like a miniature version of the <a href="/wiki/Cerebral_cortex" title="Cerebral cortex">cerebral cortex</a>, in that it has a wavy, or convoluted surface.<sup id="cite_ref-Mahut_3-1" class="reference"><a href="#cite_note-Mahut-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p>Unlike the hippocampus which is involved in the encoding of complex memories, the cerebellum plays a role in the learning of <a href="/wiki/Procedural_memory" title="Procedural memory">procedural memory</a>, and motor learning, such as skills requiring co-ordination and fine motor control.<sup id="cite_ref-Mishkin_4-0" class="reference"><a href="#cite_note-Mishkin-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> An example of a skill requiring procedural memory would be playing a musical instrument, or driving a car or riding a bike. Individuals with <a href="/wiki/Transient_global_amnesia" title="Transient global amnesia">transient global amnesia</a> that have difficulty forming new memories and/or remembering old events may sometimes retain the ability to perform complex musical pieces, suggesting that procedural memory is completely dissociated from conscious memory, also known as <a href="/wiki/Explicit_memory" title="Explicit memory">explicit memory</a>. </p><p>This separation makes sense if the cerebellum, which is far removed from the hippocampus, is responsible for procedural learning. The cerebellum is more generally involved in motor learning, and damage to it can result in problems with movement, specifically it is considered to co-ordinate timing and accuracy of movements, and to make long-term changes (learning) to improve these skills.<sup id="cite_ref-Kolb_1-1" class="reference"><a href="#cite_note-Kolb-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Amygdala">Amygdala</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=6" title="Edit section: Amygdala"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Amgydala.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/08/Amgydala.jpg/200px-Amgydala.jpg" decoding="async" width="200" height="230" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/08/Amgydala.jpg/300px-Amgydala.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/08/Amgydala.jpg/400px-Amgydala.jpg 2x" data-file-width="1181" data-file-height="1357" /></a><figcaption>The <a href="/wiki/Amygdala" title="Amygdala">amygdala</a>.</figcaption></figure> <p>Located above the hippocampus in the medial temporal lobes are two amygdalae (singular "amygdala"). The amygdalae are associated with both emotional learning and memory, as it responds strongly to emotional stimuli, especially fear. These neurons assist in encoding emotional memories and enhancing them. This process results in emotional events being more deeply and accurately encoded into memory. Lesions to the amygdalae in monkeys have been shown to impair motivation, as well as the processing of emotions.<sup id="cite_ref-Robbins_5-0" class="reference"><a href="#cite_note-Robbins-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Memory_of_fear_conditioning">Memory of fear conditioning</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=7" title="Edit section: Memory of fear conditioning"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><a href="/wiki/Pavlovian_conditioning" class="mw-redirect" title="Pavlovian conditioning">Pavlovian conditioning</a> tests have shown the active role of the amygdala in fear conditioning in rats. Research involving lesions to the basolateral nucleus have shown a strong association with memories involving fear. The central nucleus is linked with the behavioral responses that are dependent on the basolateral's reaction to fear.<sup id="cite_ref-Rabinak_6-0" class="reference"><a href="#cite_note-Rabinak-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> The central nucleus of the amygdala is also linked to emotions and behaviors motivated by food and sex.<sup id="cite_ref-McGaugh_7-0" class="reference"><a href="#cite_note-McGaugh-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Memory_consolidation">Memory consolidation</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=8" title="Edit section: Memory consolidation"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Emotional experiences and events are somewhat fragile and take a while to be completely set into memory. This slow process, referred to as consolidation, allows emotions to influence the way the memory is stored.<sup id="cite_ref-McGaugh_7-1" class="reference"><a href="#cite_note-McGaugh-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p>The amygdala is involved in <a href="/wiki/Memory_consolidation" title="Memory consolidation">memory consolidation</a>, which is the process of transferring information that is currently in working memory into ones long-term memory. This process is also known as memory modulation.<sup id="cite_ref-McGaugh_7-2" class="reference"><a href="#cite_note-McGaugh-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> The amygdala works to encode recent emotional information into memory. Memory research has shown that the greater ones emotional arousal level at the time of the event, the greater the chance that the event will be remembered.<sup id="cite_ref-McGaugh_7-3" class="reference"><a href="#cite_note-McGaugh-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> This may be due to the amygdala enhancing the emotional aspect of the information during encoding, causing the memory to be processed at a deeper level and therefore, more likely to withstand forgetting. </p> <div class="mw-heading mw-heading3"><h3 id="Basal_ganglia_and_motor_memory">Basal ganglia and motor memory</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=9" title="Edit section: Basal ganglia and motor memory"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Basal_ganglia_and_related_structures_(2).svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/85/Basal_ganglia_and_related_structures_%282%29.svg/300px-Basal_ganglia_and_related_structures_%282%29.svg.png" decoding="async" width="300" height="176" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/85/Basal_ganglia_and_related_structures_%282%29.svg/450px-Basal_ganglia_and_related_structures_%282%29.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/85/Basal_ganglia_and_related_structures_%282%29.svg/600px-Basal_ganglia_and_related_structures_%282%29.svg.png 2x" data-file-width="861" data-file-height="504" /></a><figcaption><a href="/wiki/Basal_ganglia" title="Basal ganglia">Basal ganglia</a> (red) and related structures (blue)</figcaption></figure> <p>The <a href="/wiki/Basal_ganglia" title="Basal ganglia">basal ganglia</a> are a group of <a href="/wiki/Nucleus_(neuroanatomy)" title="Nucleus (neuroanatomy)">nuclei</a> which are located in the medial temporal lobe, above the <a href="/wiki/Thalamus" title="Thalamus">thalamus</a> and connected to the cerebral cortex. Specifically, the basal ganglia includes the <a href="/wiki/Subthalamic_nucleus" title="Subthalamic nucleus">subthalamic nucleus</a>, <a href="/wiki/Substantia_nigra" title="Substantia nigra">substantia nigra</a>, the <a href="/wiki/Globus_pallidus" title="Globus pallidus">globus pallidus</a>, the <a href="/wiki/Ventral_striatum" class="mw-redirect" title="Ventral striatum">ventral striatum</a> and the <a href="/wiki/Dorsal_striatum" class="mw-redirect" title="Dorsal striatum">dorsal striatum</a>, which consists of the <a href="/wiki/Putamen" title="Putamen">putamen</a> and the <a href="/wiki/Caudate_nucleus" title="Caudate nucleus">caudate nucleus</a>.<sup id="cite_ref-Packard_8-0" class="reference"><a href="#cite_note-Packard-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> The basic functions of these nuclei deal with cognition, learning, and motor control and activities. The basal ganglia are also associated with learning, memory, and unconscious memory processes, such as motor skills and <a href="/wiki/Implicit_memory" title="Implicit memory">implicit memory</a>.<sup id="cite_ref-Mishkin_4-1" class="reference"><a href="#cite_note-Mishkin-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Particularly, one division within the ventral striatum, the nucleus accumbens core, is involved in the consolidation, retrieval and reconsolidation of drug memory.<sup id="cite_ref-drug_memory_9-0" class="reference"><a href="#cite_note-drug_memory-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p><p>The caudate nucleus is thought to assist in learning and memory of associations taught during <a href="/wiki/Operant_conditioning" title="Operant conditioning">operant conditioning</a>. Specifically, research has shown that this part of the basal ganglia plays a role in acquiring stimulus-response habits, as well as in solving sequence tasks.<sup id="cite_ref-Packard_8-1" class="reference"><a href="#cite_note-Packard-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p>Damage to the basal ganglia has been linked to dysfunctional learning of motor and perceptual-motor skills. Most disorders that are associated with damage to these areas of the brain involve some type of motor dysfunction, as well as trouble with mental switching between tasks in <a href="/wiki/Working_memory" title="Working memory">working memory</a>. Such symptoms are often present in those who suffer from <a href="/wiki/Dystonia" title="Dystonia">dystonia</a>, <a href="/wiki/Athymhormic_syndrome" title="Athymhormic syndrome">athymhormic syndrome</a>, <a href="/wiki/Fahr%27s_syndrome" class="mw-redirect" title="Fahr's syndrome">Fahr's syndrome</a>, <a href="/wiki/Huntington%27s_disease" title="Huntington's disease">Huntington's disease</a> or <a href="/wiki/Parkinson%27s_disease" title="Parkinson's disease">Parkinson's disease</a>. Huntington's and Parkinson's disease involve both motor deficits and cognitive impairment.<sup id="cite_ref-Packard_8-2" class="reference"><a href="#cite_note-Packard-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Cortical_structures">Cortical structures</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=10" title="Edit section: Cortical structures"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Lobes_of_the_brain_NL.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0e/Lobes_of_the_brain_NL.svg/220px-Lobes_of_the_brain_NL.svg.png" decoding="async" width="220" height="157" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0e/Lobes_of_the_brain_NL.svg/330px-Lobes_of_the_brain_NL.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0e/Lobes_of_the_brain_NL.svg/440px-Lobes_of_the_brain_NL.svg.png 2x" data-file-width="1024" data-file-height="731" /></a><figcaption>The cortical structures</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Frontal_lobe">Frontal lobe</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=11" title="Edit section: Frontal lobe"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The frontal lobes are located at the front of each <a href="/wiki/Cerebral_hemisphere" title="Cerebral hemisphere">cerebral hemisphere</a> and positioned <a href="/wiki/Anterior" class="mw-redirect" title="Anterior">anterior</a> to the <a href="/wiki/Parietal_lobes" class="mw-redirect" title="Parietal lobes">parietal lobes</a>. It is separated from the parietal lobe by the <a href="/wiki/Primary_motor_cortex" title="Primary motor cortex">primary motor cortex</a>, which controls voluntary movements of specific body parts associated with the <a href="/wiki/Precentral_gyrus" title="Precentral gyrus">precentral gyrus</a>.<sup id="cite_ref-Kuypers_10-0" class="reference"><a href="#cite_note-Kuypers-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> The cortex here serves our ability to plan the day, organize work, type a letter, pay attention to details and control the movements of your arms and legs. It also contributed to your personality and behaviour. </p><p>When considering the frontal lobes in regards to <a href="/wiki/Memory" title="Memory">memory</a>, we see that it is very important in the <a href="https://en.wiktionary.org/wiki/coordination" class="extiw" title="wikt:coordination">coordination</a> of information. Therefore, the frontal lobes are important in <a href="/wiki/Working_memory" title="Working memory">working memory</a>. For example, when you are thinking about how to get to a mall you have never been to before, you combine various bits of knowledge you already have: the layout of the city the mall is in, information from a map, knowledge of traffic patterns in that area and conversations with your friends about the location of the mall. By actively using all of this information, you can determine the best route for you to take. This action involves the controlled use of information in working memory, coordinated by the frontal lobes. </p><p>The frontal lobes help a person select out memories that are most relevant on a given occasion. It can coordinate various types of information into a coherent <a href="/wiki/Memory_trace" class="mw-redirect" title="Memory trace">memory trace</a>.<sup id="cite_ref-Frankland_11-0" class="reference"><a href="#cite_note-Frankland-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> For example, the knowledge of the information itself, as well as knowing where information came from must be put together into a single memory representation; this is called source monitoring.<sup id="cite_ref-Johnson_12-0" class="reference"><a href="#cite_note-Johnson-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> Sometimes we experience situations where information becomes separated, such as when we recall something, but cannot remember where we remember it from; this is referred to as a <a href="/wiki/Source_monitoring_error" class="mw-redirect" title="Source monitoring error">source monitoring error</a>.<sup id="cite_ref-Johnson_12-1" class="reference"><a href="#cite_note-Johnson-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p><p>The frontal lobes are also involved in the ability to remember what we need to do in the future; this is called <a href="/wiki/Prospective_memory" title="Prospective memory">prospective memory</a>.<sup id="cite_ref-Winograd_13-0" class="reference"><a href="#cite_note-Winograd-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Temporal_lobe">Temporal lobe</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=12" title="Edit section: Temporal lobe"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The temporal lobes are a region of the <a href="/wiki/Cerebral_cortex" title="Cerebral cortex">cerebral cortex</a> that is located beneath the <a href="/wiki/Sylvian_fissure" class="mw-redirect" title="Sylvian fissure">Sylvian fissure</a> on both the left and right hemispheres of the brain.<sup id="cite_ref-Squire_14-0" class="reference"><a href="#cite_note-Squire-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> Lobes in this cortex are more closely associated with memory and in particular <a href="/wiki/Autobiographical_memory" title="Autobiographical memory">autobiographical memory</a>.<sup id="cite_ref-Conway_15-0" class="reference"><a href="#cite_note-Conway-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p><p>The temporal lobes are also concerned with recognition memory. This is the capacity to identify an item as one that was recently encountered.<sup id="cite_ref-Rugg_16-0" class="reference"><a href="#cite_note-Rugg-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> Recognition memory is widely viewed as consisting of two components, a <a href="https://en.wiktionary.org/wiki/familiar" class="extiw" title="wikt:familiar">familiarity</a> component (i.e. Do I know this person waving at me?) and a recollective component (i.e. That is my friend Julia, from evolutionary psychology class). </p><p>Damage to the temporal lobe can affect an individual in a litany of ways ranging from: disturbance of auditory sensation and perception, disturbance of selective attention of auditory and visual input, disorders of visual perception, impaired organization and categorization of verbal material, disturbance of language comprehension, and altered personality.<sup id="cite_ref-OlderKolb_17-0" class="reference"><a href="#cite_note-OlderKolb-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p><p>In regard to memory, temporal lobe damage can impair <a href="/wiki/Long-term_memory" title="Long-term memory">long-term memory</a>.<sup id="cite_ref-OlderKolb_17-1" class="reference"><a href="#cite_note-OlderKolb-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> Thus, general semantic knowledge or more personal episodic memories of one's childhood could be affected. </p> <div class="mw-heading mw-heading3"><h3 id="Parietal_lobe">Parietal lobe</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=13" title="Edit section: Parietal lobe"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The parietal lobe is located directly behind the <a href="/wiki/Central_sulcus" title="Central sulcus">central sulcus</a>, superior to the occipital lobe and posterior to the frontal lobe, visually at the top of the back of the head.<sup id="cite_ref-Blackmore_18-0" class="reference"><a href="#cite_note-Blackmore-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> The make up of the parietal lobe is defined by four anatomical boundaries in the brain, providing a division of all the four lobes.<sup id="cite_ref-Blackmore_18-1" class="reference"><a href="#cite_note-Blackmore-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p><p>The parietal lobe has many functions and duties in the brain and its main functioning can be divided down into two main areas: (1) sensation and perception (2) constructing a spatial coordinate system to represent the world around us.<sup id="cite_ref-Kandel_19-0" class="reference"><a href="#cite_note-Kandel-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> The parietal lobe helps us to mediate attention when necessary and provides spatial awareness and navigational skills. Also, it integrates all of our sensory information (touch, sight, pain etc.) to form a single perception.<sup id="cite_ref-Kandel_19-1" class="reference"><a href="#cite_note-Kandel-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> Parietal lobe gives the ability to focus our attention on different stimuli at the same time, PET scans show high activity in the parietal lobe when participates being studied were asked to focus their attention at two separate areas of attention.<sup id="cite_ref-Kandel_19-2" class="reference"><a href="#cite_note-Kandel-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> Parietal lobe also assists with verbal short term memory and damage to the supramarginal gyrus cause short term memory loss.<sup id="cite_ref-Cowan_20-0" class="reference"><a href="#cite_note-Cowan-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> </p><p>Damage to the parietal lobe results in the syndrome ‘neglect' which is when patients treat part of their body or objects in their visual field as though it never existed. Damage to the left side of the parietal lobe can result in what is called <a href="/wiki/Gerstmann_syndrome" title="Gerstmann syndrome">Gerstmann syndrome</a>.<sup id="cite_ref-Warrington_21-0" class="reference"><a href="#cite_note-Warrington-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> It includes right-left confusion, difficulty with writing (<a href="/wiki/Agraphia" title="Agraphia">agraphia</a>) and difficulty with mathematics (<a href="/wiki/Acalculia" title="Acalculia">acalculia</a>). It can also produce disorders of language (<a href="/wiki/Aphasia" title="Aphasia">aphasia</a>) and the inability to perceive objects.<sup id="cite_ref-Warrington_21-1" class="reference"><a href="#cite_note-Warrington-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> Damage to the right parietal lobe can result in neglecting part of the body or space (contralateral neglect), which can impair many self-care skills such as dressing and washing. Right side damage can also cause difficulty in making things (constructional <a href="/wiki/Apraxia" title="Apraxia">apraxia</a>), denial of deficits (<a href="/wiki/Anosognosia" title="Anosognosia">anosognosia</a>) and drawing ability.<sup id="cite_ref-Warrington_21-2" class="reference"><a href="#cite_note-Warrington-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> Neglect syndrome tends to be more prevalent on the right side of the parietal lobe, because the right mediates attention to both the left and right fields.<sup id="cite_ref-Warrington_21-3" class="reference"><a href="#cite_note-Warrington-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> Damage in the somatic sensory cortex results in loss of perception of bodily sensations, namely sense of touch. </p> <div class="mw-heading mw-heading3"><h3 id="Occipital_lobe">Occipital lobe</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=14" title="Edit section: Occipital lobe"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The occipital lobe is the smallest of all four lobes in the human cerebral cortex and located in the rearmost part of the skull and considered to be part of the <a href="/wiki/Forebrain" title="Forebrain">forebrain</a>.<sup id="cite_ref-Westmoreland_22-0" class="reference"><a href="#cite_note-Westmoreland-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> The occipital lobe sits directly above the cerebellum and is situated <a href="/wiki/Posterior_(anatomy)" class="mw-redirect" title="Posterior (anatomy)">posterior</a> to the <a href="/wiki/Parieto-occipital_sulcus" title="Parieto-occipital sulcus">Parieto-occipital sulcus</a>, or parieto-occipital sulcus.<sup id="cite_ref-Westmoreland_22-1" class="reference"><a href="#cite_note-Westmoreland-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> This lobe is known as the centre of the <a href="/wiki/Visual_perception" title="Visual perception">visual perception</a> system, the main function of the occipital lobe is that of vision. </p><p>Retinal sensors send signals through the optic tract to the <a href="/wiki/Lateral_geniculate_nucleus" title="Lateral geniculate nucleus">Lateral geniculate nucleus</a>. Once the Lateral Geniculate Nucleus receives the information it is sent down the <a href="/wiki/Primary_visual_cortex" class="mw-redirect" title="Primary visual cortex">primary visual cortex</a> where it is organized and sent down one of two possible path ways; <a href="/wiki/Dorsum_(biology)" class="mw-redirect" title="Dorsum (biology)">dorsal</a> or <a href="/wiki/Ventral_stream" class="mw-redirect" title="Ventral stream">ventral stream</a>.<sup id="cite_ref-Goodale_23-0" class="reference"><a href="#cite_note-Goodale-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> The ventral stream is responsible for object representation and recognition and is also commonly known as the "what" stream. The dorsal stream is responsible for guiding our actions and recognizing where objects are in space, commonly known as the "where" or "how" stream. Once in the information is organized and sent through the pathways it continues to the other areas of the brain responsible for visual processing.<sup id="cite_ref-Goodale_23-1" class="reference"><a href="#cite_note-Goodale-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p><p>The most important function of the Occipital lobe is vision. Due to the positioning of this lobe at the back of the head it is not susceptible to much injury but any significant damage to the brain can cause a variety of damage to our visual perception system. Common problems in the occipital lobe are field defects and <a href="/wiki/Scotoma" title="Scotoma">scotomas</a>, movement and colour discrimination, <a href="/wiki/Hallucinations" class="mw-redirect" title="Hallucinations">hallucinations</a>, <a href="/wiki/Illusion" title="Illusion">illusions</a>, inability to recognize words and inability to recognize movement.<sup id="cite_ref-Kandel_19-3" class="reference"><a href="#cite_note-Kandel-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> A study was done in which patients suffered from a tumour on the occipital lobe and the results shows that the most frequent consequence was <a href="/wiki/Contralateral" class="mw-redirect" title="Contralateral">contralateral</a> damage to the visual field. When damage occurs in the occipital lobe it is most common to see the effects on the opposite side of the brain. Since the brain regions are so specialized in their functioning, damages done to specific areas of the brain can cause specific type of damage. Damage to the left side of the brain can lead to language discrepancies, i.e. difficulty in properly identifying letters, numbers and words, inability to incorporate visual stimuli to comprehend multiple ways an object can be found.<sup id="cite_ref-Kandel_19-4" class="reference"><a href="#cite_note-Kandel-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> Right side damage causes non-verbal problems, i.e. identifying geometric shapes, perception of figures and faces.<sup id="cite_ref-Kandel_19-5" class="reference"><a href="#cite_note-Kandel-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> In almost all regions of the brain left side damage leads to general language problems whereas right side damage leads to general perception and problem solving skills. </p> <div class="mw-heading mw-heading3"><h3 id="Damage_to_the_cortex">Damage to the cortex</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=15" title="Edit section: Damage to the cortex"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Many studies of different disease and disorders that have symptoms of memory loss have provided reinforcing evidence to the study of the anatomy of the brain and which parts are more utilized in memory. </p> <div class="mw-heading mw-heading4"><h4 id="Frontotemporal_lobar_degeneration_and_memory">Frontotemporal lobar degeneration and memory</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=16" title="Edit section: Frontotemporal lobar degeneration and memory"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><a href="/wiki/Frontotemporal_lobar_degeneration" title="Frontotemporal lobar degeneration">Frontotemporal lobar degeneration</a> (FTLD) is a common form of <a href="/wiki/Dementia" title="Dementia">dementia</a> due to the degeneration of the frontal and temporal lobes. Studies have found significant decreases in the essential needs for proper functioning in these lobes. The <a href="/wiki/Autobiographical_memory" title="Autobiographical memory">autobiographical domain in memory</a> is largely affected by this disease. In one study, FTLD patients were interviewed and asked to describe a significant event from five different periods of their lives. Using the interview and different methods of imaging, the experimenters hoped to find links between patterns of brain volume loss and performance in the interview.<sup id="cite_ref-McKinnon_24-0" class="reference"><a href="#cite_note-McKinnon-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> </p><p>Through image processing, patterns of significant reduced <a href="/wiki/Parenchymal" class="mw-redirect" title="Parenchymal">parenchymal</a> volumes that encompass the frontal and temporal lobes were found. Through comparison to a control group of patients it was found that parenchymal volumes increased during episodic recall, and decreased during semantic recall. The experimenters discussed that lifespan autobiographical episodic recall was largely damaged in FTLD patients and semantic autobiographical memory seemed to be spared.<sup id="cite_ref-McKinnon_24-1" class="reference"><a href="#cite_note-McKinnon-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Parkinson's_disease_and_memory"><span id="Parkinson.27s_disease_and_memory"></span>Parkinson's disease and memory</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=17" title="Edit section: Parkinson's disease and memory"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><a href="/wiki/Parkinson%27s_disease" title="Parkinson's disease">Parkinson's disease</a> involves both damage to the basal ganglia and certain memory dysfunctions, suggesting that the basal ganglia are involved in specific types of memory. Those who have this disease have problems with both their working memory and spatial memory.<sup id="cite_ref-Montomery_25-0" class="reference"><a href="#cite_note-Montomery-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> </p><p>Most people can instantly and easily use visual-spatial memory to remember locations and pictures, but a person with Parkinson's disease would find this difficult. He or she would also have trouble encoding this visual and spatial information into long-term memory.<sup id="cite_ref-Montomery_25-1" class="reference"><a href="#cite_note-Montomery-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> This suggests that the basal ganglia work in both encoding and recalling spatial information. </p><p>People with Parkinson's disease display working memory impairment during sequence tasks and tasks involving events in time. They also have difficulty in knowing how to use their memory, such as when to change strategies or maintain a train of thought.<sup id="cite_ref-Montomery_25-2" class="reference"><a href="#cite_note-Montomery-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Neuroanatomy_of_memory&action=edit&section=18" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-Kolb-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-Kolb_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Kolb_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFKolbWhishaw_I2008" class="citation book cs1">Kolb, B; Whishaw I (2008). <i>Fundamentals of Human Neuropsychology, 6th ed</i>. New York: Worth Publishers. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-7167-9586-5" title="Special:BookSources/978-0-7167-9586-5"><bdi>978-0-7167-9586-5</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Fundamentals+of+Human+Neuropsychology%2C+6th+ed&rft.place=New+York&rft.pub=Worth+Publishers&rft.date=2008&rft.isbn=978-0-7167-9586-5&rft.aulast=Kolb&rft.aufirst=B&rft.au=Whishaw+I&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANeuroanatomy+of+memory" class="Z3988"></span></span> </li> <li id="cite_note-Ward-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-Ward_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Ward_2-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Ward_2-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWard2009" class="citation book cs1">Ward, J (2009). <i>The Student's Guide to Cognitive Neuroscience</i>. 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class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Encoding_(memory)" title="Encoding (memory)">Encoding</a></li> <li><a href="/wiki/Storage_(memory)" title="Storage (memory)">Storage</a></li> <li><a href="/wiki/Recall_(memory)" title="Recall (memory)">Recall</a></li></ul> <ul><li><a href="/wiki/Attention" title="Attention">Attention</a></li> <li><a href="/wiki/Memory_consolidation" title="Memory consolidation">Consolidation</a></li> <li><a class="mw-selflink selflink">Neuroanatomy</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Types</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0;background:transparent;"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;font-weight:normal;"><a href="/wiki/Sensory_memory" title="Sensory memory">Sensory</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Echoic_memory" title="Echoic memory">Echoic</a></li> <li><a href="/wiki/Eidetic_memory" title="Eidetic memory">Eidetic</a></li> <li><a href="/wiki/Eyewitness_memory" title="Eyewitness memory">Eyewitness</a></li> <li><a href="/wiki/Haptic_memory" title="Haptic memory">Haptic</a></li> <li><a href="/wiki/Iconic_memory" title="Iconic memory">Iconic</a></li> <li><a href="/wiki/Motor_learning" title="Motor learning">Motor learning</a></li> <li><a href="/wiki/Visual_memory" title="Visual memory">Visual</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;font-weight:normal;"><a href="/wiki/Short-term_memory" title="Short-term memory">Short-term</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>"<a href="/wiki/The_Magical_Number_Seven,_Plus_or_Minus_Two" title="The Magical Number Seven, Plus or Minus Two">The Magical Number Seven, Plus or Minus Two</a>"</li> <li><a href="/wiki/Working_memory" title="Working memory">Working memory</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;font-weight:normal;"><a href="/wiki/Long-term_memory" title="Long-term memory">Long-term</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Active_recall" class="mw-redirect" title="Active recall">Active recall</a></li> <li><a href="/wiki/Autobiographical_memory" title="Autobiographical memory">Autobiographical</a></li> <li><a href="/wiki/Explicit_memory" title="Explicit memory">Explicit</a> <ul><li><a href="/wiki/Explicit_memory" title="Explicit memory">Declarative</a></li> <li><a href="/wiki/Episodic_memory" title="Episodic memory">Episodic</a></li> <li><a href="/wiki/Semantic_memory" title="Semantic memory">Semantic</a></li></ul></li> <li><a href="/wiki/Flashbulb_memory" title="Flashbulb memory">Flashbulb</a></li> <li><a href="/wiki/Hyperthymesia" title="Hyperthymesia">Hyperthymesia</a></li> <li><a href="/wiki/Implicit_memory" title="Implicit memory">Implicit</a></li> <li><a href="/wiki/Meaningful_learning" title="Meaningful learning">Meaningful learning</a></li> <li><a href="/wiki/Personal-event_memory" title="Personal-event memory">Personal-event</a></li> <li><a href="/wiki/Procedural_memory" title="Procedural memory">Procedural</a></li> <li><a href="/wiki/Rote_learning" title="Rote learning">Rote learning</a></li> <li><a href="/wiki/Selective_retention" title="Selective retention">Selective retention</a></li> <li><a href="/wiki/Tip_of_the_tongue" title="Tip of the tongue">Tip of the tongue</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Forgetting" title="Forgetting">Forgetting</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Amnesia" title="Amnesia">Amnesia</a> <ul><li><a href="/wiki/Anterograde_amnesia" title="Anterograde amnesia">anterograde</a></li> <li><a href="/wiki/Childhood_amnesia" title="Childhood amnesia">childhood</a></li> <li><a href="/wiki/Posthypnotic_amnesia" title="Posthypnotic amnesia">post-hypnotic</a></li> <li><a href="/wiki/Post-traumatic_amnesia" title="Post-traumatic amnesia">post-traumatic</a></li> <li><a href="/wiki/Dissociative_amnesia" title="Dissociative amnesia">dissociative (psychogenic)</a></li> <li><a href="/wiki/Retrograde_amnesia" title="Retrograde amnesia">retrograde</a></li> <li><a href="/wiki/Selective_amnesia" title="Selective amnesia">selective</a></li> <li><a href="/wiki/Transient_global_amnesia" title="Transient global amnesia">transient global</a></li></ul></li> <li><a href="/wiki/Decay_theory" title="Decay theory">Decay theory</a></li> <li><a href="/wiki/Forgetting_curve" title="Forgetting curve">Forgetting curve</a></li> <li><a href="/wiki/Interference_theory" title="Interference theory">Interference theory</a></li> <li><a href="/wiki/Memory_erasure" title="Memory erasure">Memory erasure</a></li> <li><a href="/wiki/Memory_inhibition" title="Memory inhibition">Memory inhibition</a></li> <li><a href="/wiki/Motivated_forgetting" title="Motivated forgetting">Motivated forgetting</a></li> <li><a href="/wiki/Repressed_memory" title="Repressed memory">Repressed memory</a></li> <li><a href="/wiki/Retrieval-induced_forgetting" title="Retrieval-induced forgetting">Retrieval-induced forgetting</a></li> <li><a href="/wiki/Weapon_focus" title="Weapon focus">Weapon focus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Memory_error" title="Memory error">Memory errors</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Confabulation" title="Confabulation">Confabulation</a></li> <li><a href="/wiki/Cryptomnesia" title="Cryptomnesia">Cryptomnesia</a></li> <li><a href="/wiki/Hindsight_bias" title="Hindsight bias">Hindsight bias</a></li> <li><a href="/wiki/Imagination_inflation" title="Imagination inflation">Imagination inflation</a></li> <li><a href="/wiki/List_of_memory_biases" class="mw-redirect" title="List of memory biases">Memory biases</a></li> <li><a href="/wiki/Memory_conformity" title="Memory conformity">Memory conformity</a></li> <li><a href="/wiki/Misattribution_of_memory" title="Misattribution of memory">Misattribution of memory</a></li> <li><a href="/wiki/Misinformation_effect" title="Misinformation effect">Misinformation effect</a></li> <li><a href="/wiki/Source-monitoring_error" title="Source-monitoring error">Source-monitoring error</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/False_memory" title="False memory">False memory</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Deese%E2%80%93Roediger%E2%80%93McDermott_paradigm" title="Deese–Roediger–McDermott paradigm">Deese–Roediger–McDermott paradigm</a></li> <li><a href="/wiki/False_memory_syndrome" title="False memory syndrome">False memory syndrome</a></li> <li><a href="/wiki/Memory_implantation" title="Memory implantation">Memory implantation</a> <ul><li><a href="/wiki/Lost_in_the_mall_technique" title="Lost in the mall technique">Lost in the mall technique</a></li></ul></li> <li><a href="/wiki/Recovered-memory_therapy" title="Recovered-memory therapy">Recovered-memory therapy</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Methods_used_to_study_memory" title="Methods used to study memory">Research methods</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Exceptional_memory" title="Exceptional memory">Exceptional memory</a></li> <li><a href="/wiki/Indirect_tests_of_memory" title="Indirect tests of memory">Indirect tests of memory</a></li> <li><a href="/wiki/Memory_disorder" title="Memory disorder">Memory disorder</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">In groups</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Collective_memory" title="Collective memory">Collective memory</a> <ul><li><a href="/wiki/Politics_of_memory" title="Politics of memory">Politics of memory</a></li></ul></li> <li><a href="/wiki/Cultural_memory" title="Cultural memory">Cultural memory</a></li> <li><a href="/wiki/Memory_and_social_interactions" title="Memory and social interactions">Memory and social interactions</a></li> <li><a href="/wiki/Memory_conformity" title="Memory conformity">Memory conformity</a></li> <li><a href="/wiki/Transactive_memory" title="Transactive memory">Transactive memory</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other topics</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Memory_and_aging" title="Memory and aging">Aging</a></li> <li><a href="/wiki/Art_of_memory" title="Art of memory">Art of memory</a> <ul><li><a href="/wiki/Chunking_(psychology)" title="Chunking (psychology)">chunking</a></li> <li><a href="/wiki/Mnemonic" title="Mnemonic">mnemonic</a></li></ul></li> <li><a href="/wiki/Effects_of_alcohol_on_memory" title="Effects of alcohol on memory">Effects of alcohol</a></li> <li><a href="/wiki/Neurobiological_effects_of_physical_exercise#Memory" title="Neurobiological effects of physical exercise">Effects of exercise</a></li> <li><a href="/wiki/Emotion_and_memory" title="Emotion and memory">Emotion</a></li> <li><a href="/wiki/Memory_improvement" title="Memory improvement">Memory improvement</a></li> <li><a href="/wiki/Nutrition_and_memory" class="mw-redirect" title="Nutrition and memory">Nutrition</a></li> <li><a href="/wiki/Sleep_and_memory" title="Sleep and memory">Sleep</a></li> <li><a href="/wiki/Memory_and_trauma" title="Memory and trauma">Trauma</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">In society</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Memory_sport" title="Memory sport">Memory sport</a> <ul><li><a href="/wiki/World_Memory_Championships" title="World Memory Championships">World Memory Championships</a></li></ul></li> <li><a href="/wiki/Shas_Pollak" title="Shas Pollak">Shas Pollak</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Related</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Absent-mindedness" title="Absent-mindedness">Absent-mindedness</a></li> <li><a href="/wiki/Atkinson%E2%80%93Shiffrin_memory_model" title="Atkinson–Shiffrin memory model">Atkinson–Shiffrin memory model</a></li> <li><a href="/wiki/Context-dependent_memory" title="Context-dependent memory">Context-dependent</a> and <a href="/wiki/State-dependent_memory" title="State-dependent memory">state-dependent</a> memory</li> <li><a href="/wiki/Childhood_memory" title="Childhood memory">Childhood memory</a></li> <li><a href="/wiki/Exosomatic_memory" title="Exosomatic memory">Exosomatic memory</a></li> <li><a href="/wiki/Free_recall" title="Free recall">Free recall</a></li> <li><a href="/wiki/Intermediate-term_memory" title="Intermediate-term memory">Intermediate-term memory</a></li> <li><a href="/wiki/Involuntary_memory" title="Involuntary memory">Involuntary memory</a> <ul><li><a href="/wiki/Flashback_(psychology)" title="Flashback (psychology)">flashbacks</a></li></ul></li> <li><a href="/wiki/Levels_of_Processing_model" title="Levels of Processing model">Levels of Processing model</a></li> <li><a href="/wiki/Metamemory" title="Metamemory">Metamemory</a></li> <li><a href="/wiki/Muscle_memory" title="Muscle memory">Muscle memory</a></li> <li><a href="/wiki/Priming_(psychology)" title="Priming (psychology)">Priming</a> <ul><li><a href="/wiki/Intertrial_priming" title="Intertrial priming">intertrial</a></li></ul></li> <li><a href="/wiki/Prospective_memory" title="Prospective memory">Prospective</a> and <a href="/wiki/Retrospective_memory" title="Retrospective memory">retrospective</a> memory</li> <li><i><a href="/wiki/The_Seven_Sins_of_Memory" title="The Seven Sins of Memory">The Seven Sins of Memory</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">People</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;font-weight:normal;">Researchers</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Richard_C._Atkinson" title="Richard C. Atkinson">Richard C. Atkinson</a></li> <li><a href="/wiki/Robert_A._Bjork" title="Robert A. Bjork">Robert A. Bjork</a></li> <li><a href="/wiki/Stephen_J._Ceci" title="Stephen J. Ceci">Stephen J. Ceci</a></li> <li><a href="/wiki/Susan_Clancy" title="Susan Clancy">Susan Clancy</a></li> <li><a href="/wiki/Hermann_Ebbinghaus" title="Hermann Ebbinghaus">Hermann Ebbinghaus</a></li> <li><a href="/wiki/Sigmund_Freud" title="Sigmund Freud">Sigmund Freud</a></li> <li><a href="/wiki/Patricia_Goldman-Rakic" title="Patricia Goldman-Rakic">Patricia Goldman-Rakic</a></li> <li><a href="/wiki/Ivan_Izquierdo" title="Ivan Izquierdo">Ivan Izquierdo</a></li> <li><a href="/wiki/Marcia_K._Johnson" title="Marcia K. Johnson">Marcia K. Johnson</a></li> <li><a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></li> <li><a href="/wiki/Elizabeth_Loftus" title="Elizabeth Loftus">Elizabeth Loftus</a></li> <li><a href="/wiki/Geoffrey_Loftus" title="Geoffrey Loftus">Geoffrey Loftus</a></li> <li><a href="/wiki/James_McGaugh" title="James McGaugh">James McGaugh</a></li> <li><a href="/wiki/Eleanor_Maguire" title="Eleanor Maguire">Eleanor Maguire</a></li> <li><a href="/wiki/George_Armitage_Miller" title="George Armitage Miller">George Armitage Miller</a></li> <li><a href="/wiki/Brenda_Milner" title="Brenda Milner">Brenda Milner</a></li> <li><a href="/wiki/Lynn_Nadel" title="Lynn Nadel">Lynn Nadel</a></li> <li><a href="/wiki/Henry_L._Roediger_III" title="Henry L. Roediger III">Henry L. Roediger III</a></li> <li><a href="/wiki/Daniel_Schacter" title="Daniel Schacter">Daniel Schacter</a></li> <li><a href="/wiki/Richard_Shiffrin" title="Richard Shiffrin">Richard Shiffrin</a></li> <li><a href="/wiki/Arthur_P._Shimamura" title="Arthur P. Shimamura">Arthur P. Shimamura</a></li> <li><a href="/wiki/Larry_Squire" title="Larry Squire">Larry Squire</a></li> <li><a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></li> <li><a href="/wiki/Anne_Treisman" title="Anne Treisman">Anne Treisman</a></li> <li><a href="/wiki/Endel_Tulving" title="Endel Tulving">Endel Tulving</a></li> <li><a href="/wiki/Robert_Stickgold" title="Robert Stickgold">Robert Stickgold</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;font-weight:normal;">Patients</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Henry_Molaison" title="Henry Molaison">HM</a></li> <li><a href="/wiki/Kent_Cochrane" title="Kent Cochrane">KC</a></li> <li><a href="/wiki/Patient_N.A." title="Patient N.A.">NA</a></li> <li><a href="/wiki/Clive_Wearing" title="Clive Wearing">Clive Wearing</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;font-weight:normal;">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Jonathan_Hancock" title="Jonathan Hancock">Jonathan Hancock</a></li> <li><a href="/wiki/Paul_R._McHugh" title="Paul R. McHugh">Paul R. McHugh</a></li> <li><a href="/wiki/Dominic_O%27Brien" title="Dominic O'Brien">Dominic O'Brien</a></li> <li><a href="/wiki/Ben_Pridmore" title="Ben Pridmore">Ben Pridmore</a></li> <li><a href="/wiki/Cosmos_Rossellius" title="Cosmos Rossellius">Cosmos Rossellius</a></li> <li><a href="/wiki/Andriy_Slyusarchuk" title="Andriy Slyusarchuk">Andriy Slyusarchuk</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><span class="nowrap"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Psi2.svg/28px-Psi2.svg.png" decoding="async" width="28" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Psi2.svg/42px-Psi2.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Psi2.svg/56px-Psi2.svg.png 2x" data-file-width="100" data-file-height="100" /></span></span> </span><a href="/wiki/Portal:Psychology" title="Portal:Psychology">Psychology portal</a></li> <li><span class="nowrap"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Socrates.png/18px-Socrates.png" decoding="async" width="18" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Socrates.png/27px-Socrates.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Socrates.png/36px-Socrates.png 2x" data-file-width="326" data-file-height="500" /></span></span> </span><a href="/wiki/Portal:Philosophy" title="Portal:Philosophy">Philosophy portal</a></li></ul> </div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Neuroanatomy_of_memory&oldid=1190113327">https://en.wikipedia.org/w/index.php?title=Neuroanatomy_of_memory&oldid=1190113327</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Category</a>: <ul><li><a href="/wiki/Category:Neuroscience_of_memory" title="Category:Neuroscience of memory">Neuroscience of memory</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks mw-hidden-cats-hidden">Hidden categories: <ul><li><a href="/wiki/Category:Articles_with_short_description" title="Category:Articles with short description">Articles with short description</a></li><li><a href="/wiki/Category:Short_description_is_different_from_Wikidata" title="Category:Short description is different from Wikidata">Short description is different from Wikidata</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> This page was last edited on 16 December 2023, at 01:01<span class="anonymous-show"> (UTC)</span>.</li> <li id="footer-info-copyright">Text is available under the <a href="/wiki/Wikipedia:Text_of_the_Creative_Commons_Attribution-ShareAlike_4.0_International_License" title="Wikipedia:Text of the Creative Commons Attribution-ShareAlike 4.0 International License">Creative Commons Attribution-ShareAlike 4.0 License</a>; additional terms may apply. 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