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Portugalosuchus - Wikipedia
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searchaux" style="display:none">Extinct genus of reptiles</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><i>Portugalosuchus</i><br /><div style="font-size: 85%;">Temporal range: <a href="/wiki/Late_Cretaceous" title="Late Cretaceous">Late Cretaceous</a>: <a href="/wiki/Cenomanian" title="Cenomanian">Cenomanian</a>, <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">99.7–94.3 <a href="/wiki/Megaannum" class="mw-redirect" title="Megaannum">Ma</a></span><sup id="cite_ref-Rio2021_1-0" class="reference"><a href="#cite_note-Rio2021-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:186.25538461538px; width:1.8276923076923px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:186.25538461538px; width:1.8276923076923px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:187.25538461538px; width:-0.17230769230769px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Portugalosuchus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Reptile" title="Reptile">Reptilia</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Archosaur" title="Archosaur">Archosauria</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Pseudosuchia" title="Pseudosuchia">Pseudosuchia</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Crocodylomorpha" title="Crocodylomorpha">Crocodylomorpha</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Crocodyliformes" title="Crocodyliformes">Crocodyliformes</a> </td></tr> <tr> <td>Genus: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a class="mw-selflink selflink"><i>Portugalosuchus</i></a><br /><small>Mateus <i>et al.</i>, 2018</small> </td></tr> <tr> <td>Species: </td> <td><div style="display:inline" class="species">†<i><b>P. azenhae</b></i></div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Binomial_nomenclature" title="Binomial nomenclature">Binomial name</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><b><span class="binomial"><span style="font-weight:normal;">†</span><i>Portugalosuchus azenhae</i></span></b><br /><div style="font-size: 85%;"><a href="/wiki/Oct%C3%A1vio_Mateus" title="Octávio Mateus">Mateus</a> <i>et al.</i>, <a href="/wiki/2018_in_archosaur_paleontology" title="2018 in archosaur paleontology">2018</a></div> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Portugalosuchus</b></i> (meaning "crocodile from <a href="/wiki/Portugal" title="Portugal">Portugal</a>") is an <a href="/wiki/Extinct" class="mw-redirect" title="Extinct">extinct</a> <a href="/wiki/Genus" title="Genus">genus</a> of <a href="/wiki/Eusuchia" title="Eusuchia">eusuchian</a> <a href="/wiki/Crocodyliform" class="mw-redirect" title="Crocodyliform">crocodyliform</a> that was possibly a <a href="/wiki/Basal_(phylogeny)" class="mw-redirect" title="Basal (phylogeny)">basal</a> <a href="/wiki/Crocodylia" class="mw-redirect" title="Crocodylia">crocodylian</a> – if so then it would be the oldest known crocodylian to date.<sup id="cite_ref-Mateus2018_2-0" class="reference"><a href="#cite_note-Mateus2018-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The <a href="/wiki/Type_species" title="Type species">type species</a> is <i><b>P. azenhae</b></i>, <a href="/wiki/Scientifically_described" class="mw-redirect" title="Scientifically described">described</a> in 2018,<sup id="cite_ref-Mateus2018_2-1" class="reference"><a href="#cite_note-Mateus2018-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> and it is known from the <a href="/wiki/Late_Cretaceous" title="Late Cretaceous">Late Cretaceous</a> (<a href="/wiki/Cenomanian" title="Cenomanian">Cenomanian</a>)-aged <a href="/w/index.php?title=Tentugal_Formation&action=edit&redlink=1" class="new" title="Tentugal Formation (page does not exist)">Tentugal Formation</a> in <a href="/wiki/Portugal" title="Portugal">Portugal</a>.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> A 2021 <a href="/wiki/Morphology_(biology)" title="Morphology (biology)">morphological</a> study recovered <i>Portugalosuchus</i> within <a href="/wiki/Crocodylia" class="mw-redirect" title="Crocodylia">Crocodylia</a> as a member of <a href="/wiki/Gavialidae" title="Gavialidae">Gavialidae</a> closely related to similar "thoracosaurs" (e.g. <i><a href="/wiki/Thoracosaurus" title="Thoracosaurus">Thoracosaurus</a></i>), while also noting that it might also possibly be outside of Crocodylia completely.<sup id="cite_ref-Rio2021_1-1" class="reference"><a href="#cite_note-Rio2021-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> A 2022 <a href="/wiki/Tip_dating" title="Tip dating">tip dating</a> analysis incorporating both morphological and <a href="/wiki/DNA" title="DNA">DNA</a> data placed <i>Portugalosuchus</i> outside of Crocodylia, as the sister taxon of the family <a href="/wiki/Allodaposuchidae" title="Allodaposuchidae">Allodaposuchidae</a>. A <a href="/wiki/Cladogram" title="Cladogram">cladogram</a> simplified after that analysis is shown below:<sup id="cite_ref-Darlim2022_4-0" class="reference"><a href="#cite_note-Darlim2022-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1258728058">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="font-size:85%;line-height:85%"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Eusuchia" title="Eusuchia">Eusuchia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Hylaeochampsa" title="Hylaeochampsa">Hylaeochampsa</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Acynodon" title="Acynodon">Acynodon</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Iharkutosuchus" title="Iharkutosuchus">Iharkutosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Portugalosuchus</b></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Agaresuchus" title="Agaresuchus">Agaresuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Lohuecosuchus" title="Lohuecosuchus">Lohuecosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Allodaposuchus" title="Allodaposuchus">Allodaposuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Arenysuchus" title="Arenysuchus">Arenysuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p>"Thoracosaurs" </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Borealosuchus" title="Borealosuchus">Borealosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Planocraniidae" title="Planocraniidae">Planocraniidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Crocodylia" class="mw-redirect" title="Crocodylia">Crocodylia</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <p>The morphology-based phylogenetic analyses based on the new neuroanatomical data obtained from its skull using micro-CT scans suggested that this taxon is a crown group crocodilian and a member of the 'thoracosaurs', recovered as a sister taxon of <i><a href="/wiki/Thoracosaurus" title="Thoracosaurus">Thoracosaurus</a></i> within Gavialoidea,<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> though it is uncertain whether 'thoracosaurs' were true gavialoids.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Portugalosuchus&action=edit&section=1" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-Rio2021-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-Rio2021_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Rio2021_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFRioMannion2021" class="citation journal cs1">Rio, Jonathan P.; Mannion, Philip D. (6 September 2021). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8428266">"Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem"</a>. <i><a href="/wiki/PeerJ" title="PeerJ">PeerJ</a></i>. <b>9</b>: e12094. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.7717%2Fpeerj.12094">10.7717/peerj.12094</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8428266">8428266</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/34567843">34567843</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=PeerJ&rft.atitle=Phylogenetic+analysis+of+a+new+morphological+dataset+elucidates+the+evolutionary+history+of+Crocodylia+and+resolves+the+long-standing+gharial+problem&rft.volume=9&rft.pages=e12094&rft.date=2021-09-06&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC8428266%23id-name%3DPMC&rft_id=info%3Apmid%2F34567843&rft_id=info%3Adoi%2F10.7717%2Fpeerj.12094&rft.aulast=Rio&rft.aufirst=Jonathan+P.&rft.au=Mannion%2C+Philip+D.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC8428266&rfr_id=info%3Asid%2Fen.wikipedia.org%3APortugalosuchus" class="Z3988"></span></span> </li> <li id="cite_note-Mateus2018-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-Mateus2018_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Mateus2018_2-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFOctávio_MateusEduardo_Puértolas-PascualPedro_M._Callapez2018" class="citation journal cs1">Octávio Mateus; Eduardo Puértolas-Pascual; Pedro M. Callapez (2018). "A new eusuchian crocodylomorph from the Cenomanian (Late Cretaceous) of Portugal reveals novel implications on the origin of Crocodylia". <i>Zoological Journal of the Linnean Society</i>. <b>186</b> (2): 501–528. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fzoolinnean%2Fzly064">10.1093/zoolinnean/zly064</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Zoological+Journal+of+the+Linnean+Society&rft.atitle=A+new+eusuchian+crocodylomorph+from+the+Cenomanian+%28Late+Cretaceous%29+of+Portugal+reveals+novel+implications+on+the+origin+of+Crocodylia&rft.volume=186&rft.issue=2&rft.pages=501-528&rft.date=2018&rft_id=info%3Adoi%2F10.1093%2Fzoolinnean%2Fzly064&rft.au=Oct%C3%A1vio+Mateus&rft.au=Eduardo+Pu%C3%A9rtolas-Pascual&rft.au=Pedro+M.+Callapez&rfr_id=info%3Asid%2Fen.wikipedia.org%3APortugalosuchus" class="Z3988"></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=379360">"<i>Portugalosuchus</i>"</a>. Fossilworks<span class="reference-accessdate">. Retrieved <span class="nowrap">17 December</span> 2021</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Portugalosuchus&rft.pub=Fossilworks&rft_id=https%3A%2F%2Fpaleobiodb.org%2Fclassic%2FcheckTaxonInfo%3Ftaxon_no%3D379360&rfr_id=info%3Asid%2Fen.wikipedia.org%3APortugalosuchus" class="Z3988"></span></span> </li> <li id="cite_note-Darlim2022-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-Darlim2022_4-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDarlimLeeWalterRabi2022" class="citation journal cs1">Darlim, G.; Lee, M. S. Y.; Walter, J.; Rabi, M. (2022). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8825999">"The impact of molecular data on the phylogenetic position of the putative oldest crown crocodilian and the age of the clade"</a>. <i>Biology Letters</i>. <b>18</b> (2): 20210603. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frsbl.2021.0603">10.1098/rsbl.2021.0603</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8825999">8825999</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/35135314">35135314</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Biology+Letters&rft.atitle=The+impact+of+molecular+data+on+the+phylogenetic+position+of+the+putative+oldest+crown+crocodilian+and+the+age+of+the+clade&rft.volume=18&rft.issue=2&rft.pages=20210603&rft.date=2022&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC8825999%23id-name%3DPMC&rft_id=info%3Apmid%2F35135314&rft_id=info%3Adoi%2F10.1098%2Frsbl.2021.0603&rft.aulast=Darlim&rft.aufirst=G.&rft.au=Lee%2C+M.+S.+Y.&rft.au=Walter%2C+J.&rft.au=Rabi%2C+M.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC8825999&rfr_id=info%3Asid%2Fen.wikipedia.org%3APortugalosuchus" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPuértolas‐PascualKuzminSerrano‐MartínezMateus2023" class="citation journal cs1">Puértolas‐Pascual, Eduardo; Kuzmin, Ivan T.; Serrano‐Martínez, Alejandro; Mateus, Octávio (2023-02-02). <a rel="nofollow" class="external text" href="https://onlinelibrary.wiley.com/doi/10.1111/joa.13836">"Neuroanatomy of the crocodylomorph <i>Portugalosuchus azenhae</i> from the late cretaceous of Portugal"</a>. <i>Journal of Anatomy</i>: joa.13836. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fjoa.13836">10.1111/joa.13836</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0021-8782">0021-8782</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Anatomy&rft.atitle=Neuroanatomy+of+the+crocodylomorph+Portugalosuchus+azenhae+from+the+late+cretaceous+of+Portugal&rft.pages=joa.13836&rft.date=2023-02-02&rft_id=info%3Adoi%2F10.1111%2Fjoa.13836&rft.issn=0021-8782&rft.aulast=Pu%C3%A9rtolas%E2%80%90Pascual&rft.aufirst=Eduardo&rft.au=Kuzmin%2C+Ivan+T.&rft.au=Serrano%E2%80%90Mart%C3%ADnez%2C+Alejandro&rft.au=Mateus%2C+Oct%C3%A1vio&rft_id=https%3A%2F%2Fonlinelibrary.wiley.com%2Fdoi%2F10.1111%2Fjoa.13836&rfr_id=info%3Asid%2Fen.wikipedia.org%3APortugalosuchus" class="Z3988"></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHekkalaGatesyNarechaniaMeredith2021" class="citation journal cs1">Hekkala, E.; Gatesy, J.; Narechania, A.; Meredith, R.; Russello, M.; Aardema, M. L.; Jensen, E.; Montanari, S.; Brochu, C.; Norell, M.; Amato, G. (2021-04-27). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8079395">"Paleogenomics illuminates the evolutionary history of the extinct Holocene "horned" crocodile of Madagascar, Voay robustus"</a>. <i>Communications Biology</i>. <b>4</b> (1): 505. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fs42003-021-02017-0">10.1038/s42003-021-02017-0</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/2399-3642">2399-3642</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8079395">8079395</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/33907305">33907305</a>.</cite><span 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0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="*_Kingdom:_Animalia_*_Phylum:_Chordata_*_Class:_Sauropsida_*_Clade:_Pseudosuchia_*_Clade:_Metasuchia" style="font-size:114%;margin:0 4em"><span style="font-weight:normal; font-style:normal"><span style="font-size:85%;"> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Class: <a href="/wiki/Sauropsida" title="Sauropsida">Sauropsida</a></li> <li><i>Clade</i>: <a href="/wiki/Pseudosuchia" title="Pseudosuchia">Pseudosuchia</a></li> <li><i>Clade</i>: <a href="/wiki/Metasuchia" title="Metasuchia">Metasuchia</a></li></ul> </span></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pseudosuchia" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pseudosuchia" title="Pseudosuchia">Pseudosuchia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Pseudosuchia" title="Template:Pseudosuchia">Pseudosuchia</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Neosuchia" title="Neosuchia">Neosuchia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Neosuchia" style="font-size:114%;margin:0 4em"><a href="/wiki/Neosuchia" title="Neosuchia">Neosuchia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Neosuchia" scope="row" class="navbox-group" style="width:1%"><div style="position: absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Neosuchia" title="Neosuchia">Neosuchia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Burkesuchus" title="Burkesuchus">Burkesuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Isisfordia" title="Isisfordia">Isisfordia</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Karatausuchus" title="Karatausuchus">Karatausuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Khoratosuchus" title="Khoratosuchus">Khoratosuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Montsecosuchus" title="Montsecosuchus">Montsecosuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Stolokrosuchus" title="Stolokrosuchus">Stolokrosuchus</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Susisuchus" title="Susisuchus">Susisuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Tadzhikosuchus" title="Tadzhikosuchus">Tadzhikosuchus</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Wahasuchus" title="Wahasuchus">Wahasuchus</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Zhyrasuchus" title="Zhyrasuchus">Zhyrasuchus</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><b><a href="/wiki/Thalattosuchia" title="Thalattosuchia">Thalattosuchia</a></b>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Tethysuchia" title="Tethysuchia">Tethysuchia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pholidosauridae" title="Pholidosauridae">Pholidosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Anglosuchus" title="Anglosuchus">Anglosuchus</a></i></li> <li><i><a href="/wiki/Chalawan_(reptile)" title="Chalawan (reptile)">Chalawan</a></i></li> <li><i><a href="/wiki/Crocodilaemus" title="Crocodilaemus">Crocodilaemus</a></i></li> <li><i><a href="/wiki/Dakotasuchus" title="Dakotasuchus">Dakotasuchus</a></i></li> <li><i><a href="/wiki/Elosuchus" title="Elosuchus">Elosuchus</a></i></li> <li><i><a href="/wiki/Meridiosaurus" title="Meridiosaurus">Meridiosaurus</a></i></li> <li><i><a href="/wiki/Oceanosuchus" title="Oceanosuchus">Oceanosuchus</a></i></li> <li><i><a href="/wiki/Pholidosaurus" title="Pholidosaurus">Pholidosaurus</a></i></li> <li><i><a href="/wiki/Sarcosuchus" title="Sarcosuchus">Sarcosuchus</a></i></li> <li><i><a href="/wiki/Terminonaris" title="Terminonaris">Terminonaris</a></i></li> <li><i><a href="/wiki/Vectisuchus" title="Vectisuchus">Vectisuchus</a></i></li> <li><i><a href="/wiki/Woodbinesuchus" title="Woodbinesuchus">Woodbinesuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Dyrosauridae" title="Dyrosauridae">Dyrosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Acherontisuchus" title="Acherontisuchus">Acherontisuchus</a></i></li> <li><i><a href="/wiki/Anthracosuchus" title="Anthracosuchus">Anthracosuchus</a></i></li> <li><i><a href="/wiki/Arambourgisuchus" title="Arambourgisuchus">Arambourgisuchus</a></i></li> <li><i><a href="/wiki/Atlantosuchus" title="Atlantosuchus">Atlantosuchus</a></i></li> <li><i><a href="/wiki/Brachiosuchus" title="Brachiosuchus">Brachiosuchus</a></i></li> <li><i><a href="/wiki/Cerrejonisuchus" title="Cerrejonisuchus">Cerrejonisuchus</a></i></li> <li><i><a href="/wiki/Chenanisuchus" title="Chenanisuchus">Chenanisuchus</a></i></li> <li><i><a href="/wiki/Congosaurus" title="Congosaurus">Congosaurus</a></i></li> <li><i><a href="/wiki/Dorbignysuchus" title="Dorbignysuchus">Dorbignysuchus</a></i></li> <li><i><a href="/wiki/Dyrosaurus" title="Dyrosaurus">Dyrosaurus</a></i></li> <li><i><a href="/wiki/Fortignathus" title="Fortignathus">Fortignathus</a></i></li> <li><i><a href="/wiki/Guarinisuchus" title="Guarinisuchus">Guarinisuchus</a></i></li> <li><i><a href="/wiki/Hyposaurus" title="Hyposaurus">Hyposaurus</a></i></li> <li><i><a href="/wiki/Luciasuchus" title="Luciasuchus">Luciasuchus</a></i></li> <li><i><a href="/wiki/Phosphatosaurus" title="Phosphatosaurus">Phosphatosaurus</a></i></li> <li><i><a href="/wiki/Rhabdognathus" title="Rhabdognathus">Rhabdognathus</a></i></li> <li><i><a href="/wiki/Rodeosuchus" title="Rodeosuchus">Rodeosuchus</a></i></li> <li><i><a href="/wiki/Sabinosuchus" title="Sabinosuchus">Sabinosuchus</a></i>?</li> <li><i><a href="/wiki/Sokotosuchus" title="Sokotosuchus">Sokotosuchus</a></i></li> <li><i><a href="/wiki/Tilemsisuchus" title="Tilemsisuchus">Tilemsisuchus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Atoposauridae" title="Atoposauridae">Atoposauridae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alligatorellus" title="Alligatorellus">Alligatorellus</a></i></li> <li><i><a href="/wiki/Alligatorium" title="Alligatorium">Alligatorium</a></i></li> <li><i><a href="/wiki/Aprosuchus" title="Aprosuchus">Aprosuchus</a></i></li> <li><i><a href="/wiki/Atoposaurus" title="Atoposaurus">Atoposaurus</a></i></li> <li><i><a href="/wiki/Knoetschkesuchus" title="Knoetschkesuchus">Knoetschkesuchus</a></i></li> <li><i><a href="/wiki/Sabresuchus" title="Sabresuchus">Sabresuchus</a></i></li> <li><i><a href="/wiki/Theriosuchus" title="Theriosuchus">Theriosuchus</a></i></li> <li><i><a href="/wiki/Varanosuchus" title="Varanosuchus">Varanosuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Stomatosuchidae" title="Stomatosuchidae">Stomatosuchidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Laganosuchus" title="Laganosuchus">Laganosuchus</a></i></li> <li><i><a href="/wiki/Stomatosuchus" title="Stomatosuchus">Stomatosuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Paluxysuchidae" class="mw-redirect" title="Paluxysuchidae">Paluxysuchidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Deltasuchus" title="Deltasuchus">Deltasuchus</a></i></li> <li><i><a href="/wiki/Paluxysuchus" title="Paluxysuchus">Paluxysuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Goniopholididae" title="Goniopholididae">Goniopholididae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amphicotylus" title="Amphicotylus">Amphicotylus</a></i></li> <li><i><a href="/wiki/Anteophthalmosuchus" title="Anteophthalmosuchus">Anteophthalmosuchus</a></i></li> <li><i><a href="/wiki/Calsoyasuchus" title="Calsoyasuchus">Calsoyasuchus</a></i></li> <li><i><a href="/wiki/Coelosuchus" title="Coelosuchus">Coelosuchus</a></i></li> <li><i><a href="/wiki/Denazinosuchus" title="Denazinosuchus">Denazinosuchus</a></i></li> <li><i><a href="/wiki/Diplosaurus" title="Diplosaurus">Diplosaurus</a></i></li> <li><i><a href="/wiki/Eutretauranosuchus" title="Eutretauranosuchus">Eutretauranosuchus</a></i></li> <li><i><a href="/wiki/Goniopholis" title="Goniopholis">Goniopholis</a></i></li> <li><i><a href="/wiki/Hulkepholis" title="Hulkepholis">Hulkepholis</a></i></li> <li><i><a href="/wiki/Nannosuchus" title="Nannosuchus">Nannosuchus</a></i></li> <li><i><a href="/wiki/Ophiussasuchus" title="Ophiussasuchus">Ophiussasuchus</a></i></li> <li><i><a href="/wiki/Oweniasuchus" title="Oweniasuchus">Oweniasuchus</a></i></li> <li><i><a href="/wiki/Pinacosuchus" title="Pinacosuchus">Pinacosuchus</a></i>?</li> <li><i><a href="/wiki/Siamosuchus" title="Siamosuchus">Siamosuchus</a></i></li> <li><i><a href="/wiki/Sunosuchus" title="Sunosuchus">Sunosuchus</a></i></li> <li><i><a href="/wiki/Symptosuchus" title="Symptosuchus">Symptosuchus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Bernissartiidae" title="Bernissartiidae">Bernissartiidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bernissartia" title="Bernissartia">Bernissartia</a></i></li> <li><i><a href="/wiki/Koumpiodontosuchus" title="Koumpiodontosuchus">Koumpiodontosuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Paralligatoridae" title="Paralligatoridae">Paralligatoridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Batrachomimus" title="Batrachomimus">Batrachomimus</a></i></li> <li><i><a href="/wiki/Brillanceausuchus" title="Brillanceausuchus">Brillanceausuchus</a></i></li> <li><i><a href="/wiki/Kansajsuchus" title="Kansajsuchus">Kansajsuchus</a></i></li> <li><i><a href="/wiki/Paralligator" title="Paralligator">Paralligator</a></i></li> <li><i><a href="/wiki/Rugosuchus" title="Rugosuchus">Rugosuchus</a></i></li> <li><i><a href="/wiki/Scolomastax" title="Scolomastax">Scolomastax</a></i></li> <li><i><a href="/wiki/Shamosuchus" title="Shamosuchus">Shamosuchus</a></i></li> <li><i><a href="/wiki/Tarsomordeo" title="Tarsomordeo">Tarsomordeo</a></i></li> <li><i><a href="/wiki/Turanosuchus" title="Turanosuchus">Turanosuchus</a></i></li> <li><i><a href="/wiki/Wannchampsus" title="Wannchampsus">Wannchampsus</a></i></li> <li><i><a href="/wiki/Yanjisuchus" title="Yanjisuchus">Yanjisuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Eusuchia" title="Eusuchia">Eusuchia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Oceanosuchus.jpg" class="mw-file-description" title="Oceanosuchus boecensis"><img alt="Oceanosuchus boecensis" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/08/Oceanosuchus.jpg/140px-Oceanosuchus.jpg" decoding="async" width="140" height="52" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/08/Oceanosuchus.jpg/210px-Oceanosuchus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/08/Oceanosuchus.jpg/280px-Oceanosuchus.jpg 2x" data-file-width="1000" data-file-height="368" /></a></span></span> <p><span typeof="mw:File"><a href="/wiki/File:Dyrosaurus_BW.jpg" class="mw-file-description" title="Dyrosaurus phosphaticus"><img alt="Dyrosaurus phosphaticus" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/94/Dyrosaurus_BW.jpg/140px-Dyrosaurus_BW.jpg" decoding="async" width="140" height="43" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/94/Dyrosaurus_BW.jpg/210px-Dyrosaurus_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/94/Dyrosaurus_BW.jpg/280px-Dyrosaurus_BW.jpg 2x" data-file-width="800" data-file-height="247" /></a></span> <span style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Isisfordia.jpg" class="mw-file-description" title="Isisfordia duncani"><img alt="Isisfordia duncani" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3a/Isisfordia.jpg/140px-Isisfordia.jpg" decoding="async" width="140" height="69" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3a/Isisfordia.jpg/210px-Isisfordia.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3a/Isisfordia.jpg/280px-Isisfordia.jpg 2x" data-file-width="1452" data-file-height="716" /></a></span></span> <span style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Goniopholis_BW.jpg" class="mw-file-description" title="Goniopholis simus"><img alt="Goniopholis simus" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Goniopholis_BW.jpg/140px-Goniopholis_BW.jpg" decoding="async" width="140" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Goniopholis_BW.jpg/210px-Goniopholis_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Goniopholis_BW.jpg/280px-Goniopholis_BW.jpg 2x" data-file-width="800" data-file-height="226" /></a></span></span> </p> <span style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Bernissartia_BW.jpg" class="mw-file-description" title="Bernissartia fagesii"><img alt="Bernissartia fagesii" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d0/Bernissartia_BW.jpg/140px-Bernissartia_BW.jpg" decoding="async" width="140" height="45" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d0/Bernissartia_BW.jpg/210px-Bernissartia_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d0/Bernissartia_BW.jpg/280px-Bernissartia_BW.jpg 2x" data-file-width="800" data-file-height="258" /></a></span></span></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Eusuchia" style="font-size:114%;margin:0 4em"><a href="/wiki/Eusuchia" title="Eusuchia">Eusuchia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Eusuchia" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg);;"><a href="/wiki/Eusuchia" title="Eusuchia">Eusuchia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Borealosuchus" title="Borealosuchus">Borealosuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Confractosuchus" title="Confractosuchus">Confractosuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Dolichochampsa" title="Dolichochampsa">Dolichochampsa</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Gilchristosuchus" title="Gilchristosuchus">Gilchristosuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Heterosuchus" title="Heterosuchus">Heterosuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Isisfordia" title="Isisfordia">Isisfordia</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a class="mw-selflink selflink">Portugalosuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Pristichampsus" title="Pristichampsus">Pristichampsus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Susisuchus" title="Susisuchus">Susisuchus</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Titanochampsa" title="Titanochampsa">Titanochampsa</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><b><a href="/wiki/Atoposauridae" title="Atoposauridae">Atoposauridae</a></b>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><b><a href="/wiki/Bernissartiidae" title="Bernissartiidae">Bernissartiidae</a></b>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><b><a href="/wiki/Paralligatoridae" title="Paralligatoridae">Paralligatoridae</a></b>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Hylaeochampsidae" title="Hylaeochampsidae">Hylaeochampsidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Acynodon" title="Acynodon">Acynodon</a></i>?</li> <li><i><a href="/wiki/Hylaeochampsa" title="Hylaeochampsa">Hylaeochampsa</a></i></li> <li><i><a href="/wiki/Iharkutosuchus" title="Iharkutosuchus">Iharkutosuchus</a></i></li> <li><i><a href="/wiki/Pachycheilosuchus" title="Pachycheilosuchus">Pachycheilosuchus</a></i>?</li> <li><i><a href="/wiki/Pietraroiasuchus" title="Pietraroiasuchus">Pietraroiasuchus</a></i></li> <li><i><a href="/wiki/Turcosuchus" title="Turcosuchus">Turcosuchus</a></i></li> <li><i><a href="/wiki/Unasuchus" title="Unasuchus">Unasuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Allodaposuchidae" title="Allodaposuchidae">Allodaposuchidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Agaresuchus" title="Agaresuchus">Agaresuchus</a></i></li> <li><i><a href="/wiki/Allodaposuchus" title="Allodaposuchus">Allodaposuchus</a></i></li> <li><i><a href="/wiki/Arenysuchus" title="Arenysuchus">Arenysuchus</a></i>?</li> <li><i><a href="/wiki/Ischyrochampsa" title="Ischyrochampsa">Ischyrochampsa</a></i></li> <li><i><a href="/wiki/Lohuecosuchus" title="Lohuecosuchus">Lohuecosuchus</a></i></li> <li><i><a href="/wiki/Massaliasuchus" title="Massaliasuchus">Massaliasuchus</a></i></li> <li><i><a href="/wiki/Musturzabalsuchus" title="Musturzabalsuchus">Musturzabalsuchus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Aegyptosuchidae" title="Aegyptosuchidae">Aegyptosuchidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aegyptosuchus" title="Aegyptosuchus">Aegyptosuchus</a></i></li> <li><i><a href="/wiki/Aegisuchus" title="Aegisuchus">Aegisuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr>"Thoracosaurs"</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Argochampsa" title="Argochampsa">Argochampsa</a></i></li> <li><i><a href="/wiki/Eogavialis" title="Eogavialis">Eogavialis</a></i></li> <li><i><a href="/wiki/Eosuchus" title="Eosuchus">Eosuchus</a></i></li> <li><i><a href="/wiki/Eothoracosaurus" title="Eothoracosaurus">Eothoracosaurus</a></i></li> <li><i><a href="/wiki/Thoracosaurus" title="Thoracosaurus">Thoracosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Planocraniidae" title="Planocraniidae">Planocraniidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Boverisuchus" title="Boverisuchus">Boverisuchus</a></i></li> <li><i><a href="/wiki/Duerosuchus" title="Duerosuchus">Duerosuchus</a></i></li> <li><i><a href="/wiki/Planocrania" title="Planocrania">Planocrania</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Crocodilia" title="Crocodilia">Crocodilia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Crocodilia" title="Template:Crocodilia">Crocodilia</a> for extant taxa, and <a href="/wiki/Template:Extinct_Crocodilia" title="Template:Extinct Crocodilia">Extinct Crocodilia</a> for extinct ones</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Allodaposuchus_BW.jpg" class="mw-file-description" title="Allodaposuchus precedens"><img alt="Allodaposuchus precedens" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9c/Allodaposuchus_BW.jpg/140px-Allodaposuchus_BW.jpg" decoding="async" width="140" height="59" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9c/Allodaposuchus_BW.jpg/210px-Allodaposuchus_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9c/Allodaposuchus_BW.jpg/280px-Allodaposuchus_BW.jpg 2x" data-file-width="800" data-file-height="338" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>See also: <a href="/wiki/Coelognathosuchia" title="Coelognathosuchia">Coelognathosuchia</a></li> <li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Neosuchians" title="Category:Neosuchians">Category</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Portugalosuchus</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q105833531" class="extiw" title="wikidata:Q105833531">Q105833531</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/61142149">61142149</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/11149308">11149308</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=379360">379360</a></span></span></li></ul> </div></td></tr></tbody></table></div> <p><br /> </p> <style data-mw-deduplicate="TemplateStyles:r1012311289">.mw-parser-output .asbox{position:relative;overflow:hidden}.mw-parser-output .asbox table{background:transparent}.mw-parser-output .asbox p{margin:0}.mw-parser-output .asbox p+p{margin-top:0.25em}.mw-parser-output .asbox-body{font-style:italic}.mw-parser-output .asbox-note{font-size:smaller}.mw-parser-output .asbox .navbar{position:absolute;top:-0.75em;right:1em;display:none}</style><div role="note" class="metadata plainlinks asbox stub"><table role="presentation"><tbody><tr class="noresize"><td><span typeof="mw:File"><a href="/wiki/File:Quetzalcoatlus07.jpg" class="mw-file-description"><img alt="Stub icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/42/Quetzalcoatlus07.jpg/60px-Quetzalcoatlus07.jpg" decoding="async" width="60" height="37" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/42/Quetzalcoatlus07.jpg/90px-Quetzalcoatlus07.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/42/Quetzalcoatlus07.jpg/120px-Quetzalcoatlus07.jpg 2x" data-file-width="947" data-file-height="579" /></a></span></td><td><p class="asbox-body">This article about a <a href="/wiki/Cretaceous" title="Cretaceous">Cretaceous</a> <a href="/wiki/Reptile" title="Reptile">reptile</a> is a <a href="/wiki/Wikipedia:Stub" title="Wikipedia:Stub">stub</a>. You can help Wikipedia by <a class="external text" href="https://en.wikipedia.org/w/index.php?title=Portugalosuchus&action=edit">expanding it</a>.</p></td></tr></tbody></table><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Cretaceous-reptile-stub" title="Template:Cretaceous-reptile-stub"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Cretaceous-reptile-stub" title="Template talk:Cretaceous-reptile-stub"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Cretaceous-reptile-stub" title="Special:EditPage/Template:Cretaceous-reptile-stub"><abbr title="Edit this template">e</abbr></a></li></ul></div></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1012311289"><div role="note" class="metadata plainlinks asbox stub"><table role="presentation"><tbody><tr class="noresize"><td><span typeof="mw:File"><a href="/wiki/File:Postosuchus_kirkpatricki_flipped.jpg" class="mw-file-description"><img alt="Stub icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Postosuchus_kirkpatricki_flipped.jpg/90px-Postosuchus_kirkpatricki_flipped.jpg" decoding="async" width="90" height="23" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Postosuchus_kirkpatricki_flipped.jpg/135px-Postosuchus_kirkpatricki_flipped.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Postosuchus_kirkpatricki_flipped.jpg/180px-Postosuchus_kirkpatricki_flipped.jpg 2x" data-file-width="2948" data-file-height="744" /></a></span></td><td><p class="asbox-body">This article about a <a href="/wiki/Prehistoric" class="mw-redirect" title="Prehistoric">prehistoric</a> <a href="/wiki/Archosaur" title="Archosaur">archosaur</a> is a <a href="/wiki/Wikipedia:Stub" title="Wikipedia:Stub">stub</a>. You can help Wikipedia by <a class="external text" href="https://en.wikipedia.org/w/index.php?title=Portugalosuchus&action=edit">expanding it</a>.</p></td></tr></tbody></table><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Paleo-archosaur-stub" title="Template:Paleo-archosaur-stub"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Paleo-archosaur-stub" title="Template talk:Paleo-archosaur-stub"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Paleo-archosaur-stub" title="Special:EditPage/Template:Paleo-archosaur-stub"><abbr title="Edit this template">e</abbr></a></li></ul></div></div> <!-- NewPP limit report Parsed by mw‐web.codfw.canary‐84779d6bf6‐jf9pb Cached time: 20241125052500 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1] CPU time usage: 1.117 seconds Real time usage: 1.369 seconds Preprocessor visited node count: 28158/1000000 Post‐expand include size: 321116/2097152 bytes Template argument size: 121031/2097152 bytes Highest expansion depth: 28/100 Expensive parser function count: 4/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 73213/5000000 bytes Lua time usage: 0.701/10.000 seconds Lua memory usage: 5700386/52428800 bytes Number of Wikibase entities loaded: 4/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 1264.016 1 -total 56.54% 714.626 1 Template:Speciesbox 55.95% 707.249 1 Template:Taxobox/core 21.13% 267.041 72 Template:Period_start 20.45% 258.443 12 Template:Clade 15.93% 201.334 1 Template:Geological_range 15.72% 198.682 2 Template:Phanerozoic_220px 14.74% 186.329 72 Template:Period_id 13.88% 175.485 22 Template:Fossil_range/bar 11.82% 149.458 1 Template:Reflist --> <!-- Saved in parser cache with key enwiki:pcache:idhash:66859656-0!canonical and timestamp 20241125052500 and revision id 1258135935. 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