CINXE.COM

<!DOCTYPE html> <html dir='ltr' lang='en'> <head> <meta name="csrf-param" content="authenticity_token" /> <meta name="csrf-token" content="AhdlXSypQQvojhagBynQDey75JbSYNlYqYYlCqAFVAjYcz6tBQ1naQlVRJDFJ20eQr1vxV21Wznr2OMaC25wAw==" /> <meta content='text/html; charset=UTF-8' http-equiv='Content-Type'> <meta content='width=device-width, initial-scale=1' name='viewport'> <title> Encyclopedia of Life </title> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/button.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/dimmer.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/header.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/icon.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/image.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/input.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/label.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/loader.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/list.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/menu.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/dropdown.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/search.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/segment.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/statistic.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/sticky.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.3/components/transition.min.css" /> <link rel="stylesheet" media="all" href="https://cdnjs.cloudflare.com/ajax/libs/font-awesome/4.7.0/css/font-awesome.min.css" /> <link rel="stylesheet" media="all" href="/packs/css/stylesheets/application-1f29efcb.css" /> <link rel="stylesheet" media="all" href="/assets/jqcloud-f0639a0b65fbc9997c3a768b99fe8ee5611ffee32d1cf23083b63ee861bcc602.css" /> <script src="https://code.jquery.com/jquery-1.12.4.min.js" crossorigin="anonymous" integrity="sha256-ZosEbRLbNQzLpnKIkEdrPv7lOy9C27hHQ+Xp8a4MxAQ=" defer="defer"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/corejs-typeahead/1.2.0/typeahead.bundle.min.js" defer="defer"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/uikit/3.1.6/js/uikit.min.js" defer="defer"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/uikit/3.1.6/js/uikit-icons.min.js" defer="defer"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.1/components/api.min.js" defer="defer"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.1/components/search.min.js" defer="defer"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.1/components/sticky.min.js" defer="defer"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.1/components/dimmer.min.js" defer="defer"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.1/components/dropdown.min.js" defer="defer"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/semantic-ui/2.3.1/components/transition.min.js" defer="defer"></script> <script> I18n = {}; I18n.defaultLocale = "en" I18n.locale = "en" </script> <script src="/packs/js/runtime-138760ce02a10efbb84f.js" defer="defer"></script> <script src="/packs/js/8915-b1697b29d0a6b6d6bf9f.js" defer="defer"></script> <script src="/packs/js/application-431428328720eaf3316c.js" defer="defer"></script> <script async src='https://www.googletagmanager.com/gtag/js?id=UA-3298646-10'></script> <script> window.dataLayer = window.dataLayer || []; function gtag(){dataLayer.push(arguments);} gtag('js', new Date()); gtag('config', 'UA-3298646-10'); </script>  <script> window.webtrendsAsyncInit=function(){ var dcs=new Webtrends.dcs().init({ dcsid:"dcsg0chobadzpxfga2extd7pb_7c2s", domain:"logs1.smithsonian.museum", timezone:-5, i18n:true, fpcdom:".eol.org", plugins:{ } }).track(); }; (function(){ var s=document.createElement("script"); s.async=true; s.src="https://www.si.edu/sites/all/modules/si_webtrends/js/webtrends.min.js"; var s2=document.getElementsByTagName("script")[0]; s2.parentNode.insertBefore(s,s2); }()); </script> <noscript> <img alt='dcsimg' height='1' id='dcsimg' src='//logs1.smithsonian.museum/dcsg0chobadzpxfga2extd7pb_7c2s/njs.gif?dcsuri=/nojavascript&amp;WT.js=No&amp;WT.tv=10.4.23&amp;dcssip=www.eol.org' width='1'> </noscript>  <meta name="csrf-param" content="authenticity_token" /> <meta name="csrf-token" content="GyG2/oR5BCFwV9u7MZDcWJ3Bfb6cgbKLhup+WgqWPfvBRe0Ord0iQ5GMiYvznmFLM8f27RNUMOrEtLhKof0Z8A==" /> </head> <body> <div id='mobile-menu' uk-offcanvas='overlay: true; flip: true'> <div class='uk-offcanvas-bar'> <ul class='uk-nav'> <li><a href="/docs/what-is-eol">About</a></li> <li><a href="http://education.eol.org">Education</a></li> <li><a href="http://discuss.eol.org">Discuss</a></li> <li><a href="/traitbank">TraitBank</a></li> <li><a href="/users/sign_in">Sign In</a></li> <li><a href="/users/sign_up">Sign Up</a></li> <li class='uk-nav-header'>Language</li> <li><a href="/set_locale?set_locale=de">Deutsch</a></li> <li><a href="/set_locale?set_locale=en">English</a></li> <li><a href="/set_locale?set_locale=es">Español</a></li> <li><a href="/set_locale?set_locale=fr">français</a></li> <li><a href="/set_locale?set_locale=it">italiano</a></li> <li><a href="/set_locale?set_locale=nl">Nederlands</a></li> <li><a href="/set_locale?set_locale=pms">Piemontèis</a></li> <li><a href="/set_locale?set_locale=pt-BR">Português do Brasil</a></li> <li><a href="/set_locale?set_locale=fi">suomi</a></li> <li><a href="/set_locale?set_locale=tr">Türkçe</a></li> <li><a href="/set_locale?set_locale=cs">čeština</a></li> <li><a href="/set_locale?set_locale=el">Ελληνικά</a></li> <li><a href="/set_locale?set_locale=mk">македонски</a></li> <li><a href="/set_locale?set_locale=uk">Українська</a></li> <li><a href="/set_locale?set_locale=ar">العربية</a></li> <li><a href="/set_locale?set_locale=zh-CN">简体中文</a></li> <li><a href="/set_locale?set_locale=zh-TW">繁體中文</a></li> <li class='uk-nav-header'>names in breadcrumbs</li> <li> <form class="js-bread-type-toggle" action="/breadcrumb_type" accept-charset="UTF-8" method="post"><input name="utf8" type="hidden" value="✓" autocomplete="off" /><input type="hidden" name="authenticity_token" value="eZbybBAu7DaO4SfdilQKGZmFKR17wNtvFQvFwWwJ8XzZpNCcuSGrrG3Z7S2EpIWQ8gSkryNqC/kfoFUmPLgkrQ==" autocomplete="off" /> <select name="type" id="type"><option selected="selected" value="0">vernacular</option> <option value="1">scientific</option></select> </form> </li> </ul> <div class='uk-offcanvas-close' uk-close></div> </div> </div> <div class='l-nav js-navbar'> <div class='nav-left'> <div class='nav-item nav-menu-anchor uk-visible@s'> <a class="nav-logo" href="/"><img alt="EOL" src="/assets/blue_logo-5c5dbb19100961ce1612fdb8fc090b944f11a79066a7163558d3ba52ecb36746.png" /></a> <span> </span> <i class='fa fa-chevron-down'></i> </div> <div uk-dropdown='mode: click; pos: bottom-left; offset: 0'> <ul class='uk-nav uk-dropdown-nav uk-nav-primary'> <li><a href="/docs/what-is-eol">About</a></li> <li><a href="http://education.eol.org">Education</a></li> <li><a href="http://discuss.eol.org">Discuss</a></li> <li><a href="/traitbank">TraitBank</a></li> <li><a href="/users/sign_in">Sign In</a></li> <li><a href="/users/sign_up">Sign Up</a></li> </ul> </div> <div class='nav-item nav-menu-anchor uk-visible@s'> <i class='fa fa-globe'></i>   en   <i class='fa fa-chevron-down'></i> </div> <div uk-dropdown='mode: click; pos: bottom-left; offset: 0'> <ul class='uk-nav uk-dropdown-nav'> <li><a href="/set_locale?set_locale=de">Deutsch</a></li> <li><a href="/set_locale?set_locale=en">English</a></li> <li><a href="/set_locale?set_locale=es">Español</a></li> <li><a href="/set_locale?set_locale=fr">français</a></li> <li><a href="/set_locale?set_locale=it">italiano</a></li> <li><a href="/set_locale?set_locale=nl">Nederlands</a></li> <li><a href="/set_locale?set_locale=pms">Piemontèis</a></li> <li><a href="/set_locale?set_locale=pt-BR">Português do Brasil</a></li> <li><a href="/set_locale?set_locale=fi">suomi</a></li> <li><a href="/set_locale?set_locale=tr">Türkçe</a></li> <li><a href="/set_locale?set_locale=cs">čeština</a></li> <li><a href="/set_locale?set_locale=el">Ελληνικά</a></li> <li><a href="/set_locale?set_locale=mk">македонски</a></li> <li><a href="/set_locale?set_locale=uk">Українська</a></li> <li><a href="/set_locale?set_locale=ar">العربية</a></li> <li><a href="/set_locale?set_locale=zh-CN">简体中文</a></li> <li><a href="/set_locale?set_locale=zh-TW">繁體中文</a></li> </ul> </div> <div class='nav-item nav-menu-anchor uk-hidden@s'> <a class="nav-logo" href="/"><img alt="EOL" src="/assets/blue_logo-5c5dbb19100961ce1612fdb8fc090b944f11a79066a7163558d3ba52ecb36746.png" /></a> </div> <div class='nav-item nav-menu-anchor uk-visible@s'> <i class='fa fa-cog'></i> <i class='fa fa-chevron-down'></i> </div> <div uk-dropdown='mode: click; pos: bottom-left; offset: 0'> <div class='uk-nav uk-dropdown-nav uk-nav-primary'> <div class='uk-nav-header'>names in breadcrumbs</div> <form class="js-bread-type-toggle" action="/breadcrumb_type" accept-charset="UTF-8" method="post"><input name="utf8" type="hidden" value="✓" autocomplete="off" /><input type="hidden" name="authenticity_token" value="AX/rqGWMEC3LsAxMqoBdB4G7vzPz/NviHRMEwH3hU4ehTclYzINXtyiIxrykcNKO6joygatWC3QXuJQnLVCGVg==" autocomplete="off" /> <div class='uk-grid-small uk-child-width-auto uk-grid'> <label> <input type="radio" name="type" id="type_0" value="0" class="uk-radio" checked="checked" /> vernacular </label> <label> <input type="radio" name="type" id="type_1" value="1" class="uk-radio" /> scientific </label> </div> </form> </div> </div> </div> <form class="nav-search-form" action="/search" accept-charset="UTF-8" method="get"><input name="utf8" type="hidden" value="✓" autocomplete="off" /> <input type="text" name="q" id="q" class="nav-search js-nav-search" data-no-results-text="<strong>No confirmed results</strong><br />Hit Enter to see uncurated results" autocomplete="off" /> <i class='fa fa-search fa-lg nav-item nav-search-icon'></i> </form> <div class='nav-right nav-item nav-menu-anchor nav-menu-anchor-bars uk-hidden@s' uk-toggle='#mobile-menu'> <i class='fa fa-bars'></i> </div> </div> <img class="print-logo" src="/assets/print_logo-d9be0e683b5dbbd007c371799af3aeb9d7444008a39bd56c9e383ce79312350c.png" /> <div class='uk-container'> <div id='main'> <div class='uk-section uk-padding-remove-bottom'> <h1>Internal server error (500)</h1> </div> </div> </div> <div class='footer'> <ul class='footer-section footer-section-eol-info'> <li><a href="/docs/what-is-eol">What is EOL?</a></li> <li><a href="/docs/what-is-eol/whats-new">What's New?</a></li> <li><a href="http://discuss.eol.org">The EOL Forum</a></li> <li><a href="https://education.eol.org">Education</a></li> <li><a href="/docs/what-is-eol/citing-eol-and-eol-content">Citing EOL</a></li> <li><a href="/docs/what-is-eol/terms-of-use">Terms of Use</a></li> <li><a href="/docs/what-is-eol/data-services">Data Services</a></li> <li><a href="/docs/discover/food-webs">Food Webs</a></li> <li><a href="/docs/what-is-eol/language-support">Language Support</a></li> <li><a href="/docs/what-is-eol/donate">Donate</a></li> <li><a href="http://discuss.eol.org/t/contact-us-at-eol/181">Contact Us</a></li> </ul> <ul class='footer-section footer-section-discover'> <li><a href="/docs/discover/animals">Animals</a></li> <li><a href="/docs/discover/mammals">Mammals</a></li> <li><a href="/docs/discover/birds">Birds</a></li> <li><a href="/docs/discover/amphibians">Amphibians</a></li> <li><a href="/docs/discover/reptiles">Reptiles</a></li> <li><a href="/docs/discover/fishes">Fishes</a></li> <li><a href="/docs/discover/invertebrates">Invertebrates</a></li> <li><a href="/docs/discover/crustaceans">Crustaceans</a></li> <li><a href="/docs/discover/mollusks">Mollusks</a></li> <li><a href="/docs/discover/insects">Insects</a></li> <li><a href="/docs/discover/spiders">Spiders</a></li> <li><a href="/docs/discover/worms">Worms</a></li> <li><a href="/docs/discover/plants">Plants</a></li> <li><a href="/docs/discover/flowering-plants">Flowering Plants</a></li> <li><a href="/docs/discover/trees">Trees</a></li> <li><a href="/docs/discover/fungi">Fungi</a></li> <li><a href="/docs/discover/mushrooms">Mushrooms</a></li> <li><a href="/docs/discover/molds">Molds</a></li> <li><a href="/docs/discover/bacteria">Bacteria</a></li> <li><a href="/docs/discover/algae">Algae</a></li> <li><a href="/docs/discover/protists-or-protozoa">Protists</a></li> <li><a href="/docs/discover/archaea">Archaea</a></li> <li><a href="/docs/discover/viruses">Viruses</a></li> </ul> <div class='footer-section footer-section-nmnh'> <a href="https://naturalhistory.si.edu/research/eol"><div class='nmnh-header'>EOL is hosted by:</div> <img class="nmnh-logo" src="/assets/nmnh_logo_bw-72cc01336737d61fba9c8c5cda8e2f549bd19ae44f2b0568a236f378761d53b3.png" /> </a></div> </div> </body> </html>