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class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox"><tbody><tr><th colspan="2" class="infobox-above" style="background-color:lemonchiffon">Dorsal cochlear nucleus</th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Gray760.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Gray760.png/250px-Gray760.png" decoding="async" width="250" height="162" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Gray760.png/375px-Gray760.png 1.5x, //upload.wikimedia.org/wikipedia/commons/0/0b/Gray760.png 2x" data-file-width="500" data-file-height="323" /></a></span><div class="infobox-caption">Dorsal cochlear nucleus is #4, at upper left</div></td></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Human_caudal_brainstem_posterior_view_description.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f6/Human_caudal_brainstem_posterior_view_description.JPG/250px-Human_caudal_brainstem_posterior_view_description.JPG" decoding="async" width="250" height="345" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/f/f6/Human_caudal_brainstem_posterior_view_description.JPG 1.5x" data-file-width="344" data-file-height="475" /></a></span><div class="infobox-caption">Human caudal brainstem posterior view (Dorsal cochlear nucleus is #5)</div></td></tr><tr><th colspan="2" class="infobox-header" style="background-color: #efefef">Details</th></tr><tr><th colspan="2" class="infobox-header" style="background-color: #efefef">Identifiers</th></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Latin" title="Latin">Latin</a></th><td class="infobox-data"><i>nucleus cochlearis posterior</i></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/NeuroNames" title="NeuroNames">NeuroNames</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="http://braininfo.rprc.washington.edu/centraldirectory.aspx?ID=721">721</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/NeuroLex" title="NeuroLex">NeuroLex</a> ID</th><td class="infobox-data"><a rel="nofollow" class="external text" href="http://uri.neuinfo.org/nif/nifstd/birnlex_2569">birnlex_2569</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Terminologia_Anatomica" title="Terminologia Anatomica">TA98</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://ifaa.unifr.ch/Public/EntryPage/TA98%20Tree/Entity%20TA98%20EN/14.1.04.248%20Entity%20TA98%20EN.htm">A14.1.04.248</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Terminologia_Anatomica" title="Terminologia Anatomica">TA2</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://ta2viewer.openanatomy.org/?id=6007">6007</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Foundational_Model_of_Anatomy" title="Foundational Model of Anatomy">FMA</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://bioportal.bioontology.org/ontologies/FMA/?p=classes&conceptid=http%3A%2F%2Fpurl.org%2Fsig%2Font%2Ffma%2Ffma54624">54624</a></td></tr><tr><td colspan="2" class="infobox-below"><a href="/wiki/Anatomical_terms_of_neuroanatomy" title="Anatomical terms of neuroanatomy">Anatomical terms of neuroanatomy</a><div style="text-align: right;"><small class="noprint">[<a href="https://www.wikidata.org/wiki/Q5298771" class="extiw" title="d:Q5298771">edit on Wikidata</a>]</small></div></td></tr></tbody></table> <p>The <b>dorsal cochlear nucleus</b> (<b>DCN</b>, also known as the "<b>tuberculum acusticum</b>") is a cortex-like structure on the dorso-lateral surface of the brainstem. Along with the <a href="/wiki/Ventral_cochlear_nucleus" title="Ventral cochlear nucleus">ventral cochlear nucleus</a> (VCN), it forms the <a href="/wiki/Cochlear_nucleus" title="Cochlear nucleus">cochlear nucleus</a> (CN), where all auditory nerve fibers from the cochlea form their first synapses. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Anatomy">Anatomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dorsal_cochlear_nucleus&action=edit&section=1" title="Edit section: Anatomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The DCN differs from the ventral portion of the CN as it not only projects to the central nucleus (a subdivision) of the <a href="/wiki/Inferior_colliculus" title="Inferior colliculus">inferior colliculus</a> (CIC), but also receives efferent innervation from the <a href="/wiki/Auditory_cortex" title="Auditory cortex">auditory cortex</a>, <a href="/wiki/Superior_olivary_complex" title="Superior olivary complex">superior olivary complex</a> and the inferior colliculus. The <a href="/wiki/Cytoarchitecture" title="Cytoarchitecture">cytoarchitecture</a> and <a href="/wiki/Neurochemistry" title="Neurochemistry">neurochemistry</a> of the DCN is similar to that of the <a href="/wiki/Cerebellum" title="Cerebellum">cerebellum</a>, an important concept in theories of DCN function.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Thus, the DCN is thought to be involved with more complex auditory processing, rather than merely transferring information. </p><p>The <a href="/wiki/Pyramidal_cells" class="mw-redirect" title="Pyramidal cells">pyramidal cells</a> or giant cells are a major cell grouping of the DCN. These cells are the target of two different input systems. The first system arises from the auditory nerve, and carries acoustic information. The second set of inputs is relayed through a set of small granule cells in the cochlear nucleus. There are also a great number of neighbouring <a href="/wiki/Cartwheel_cell" title="Cartwheel cell">cartwheel cells</a>.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The granule cells in turn are the target of a number of different inputs, including both those involved in auditory processing and, at least in lower mammals, somatosensory inputs associated with the head, the ear, and the jaw. </p><p>Projections from DCN principal cells form the dorsal acoustic stria, which ultimately terminate in the CIC. This projection overlaps with that of the <a href="/wiki/Lateral_superior_olive" class="mw-redirect" title="Lateral superior olive">lateral superior olive</a> (LSO) in a well-defined manner,<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> where they form the primary excitatory input for ICC type O units.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Physiology">Physiology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dorsal_cochlear_nucleus&action=edit&section=2" title="Edit section: Physiology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Principal cells in the DCN have very complex frequency intensity tuning curves. Classified as <a href="/wiki/Cochlear_nucleus" title="Cochlear nucleus">cochlear nucleus</a> type IV cells,<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> the <a href="/wiki/Action_potential#Biophysical_basis" title="Action potential">firing rate</a> may be very rapid in response to a low intensity sound at one frequency and then fall below the spontaneous rate with only a small increment in stimulus frequency or intensity. The firing rate may then increase with another increment in intensity or frequency. Type IV cells are excited by wide band noise, and particularly excited by a noise-notch stimulus directly below the cell's best frequency (BF). </p><p>While the VCN bushy cells aid in the location of a sound stimulus on the horizontal axis via their inputs to the <a href="/wiki/Superior_olivary_complex" title="Superior olivary complex">superior olivary complex</a>, type IV cells may participate in localization of the sound stimulus on the vertical axis. The <a href="/wiki/Pinna_(anatomy)" class="mw-redirect" title="Pinna (anatomy)">pinna</a> selectively amplifies frequencies, resulting in reduced sound energy at specific frequencies in certain regions of space. The complicated firing patterns of type IV cells makes them especially suited to detecting these notches, and with the combined power of these two localization systems, an ordinary person can locate where a firework explodes without the use of their eyes. </p><p>Somatosensory inputs inhibit type IV cell activity, possibly silencing their activity during head and pinna movements.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> While this has not been studied extensively, it may play an important role in sound source localization in elevation. A similar effect is seen in the visual system in an effect known as <a href="/wiki/Change_blindness" title="Change blindness">change blindness</a>. </p><p>Current auditory models of the DCN employ a two-inhibitor model. Type IV cells receive excitation directly from the auditory nerve, and are inhibited by type II (vertical) cells and a wide band inhibitor (onset-c cells). </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dorsal_cochlear_nucleus&action=edit&section=3" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text">Bell CC, Han V et al. Cerebellum-like structures and their implications for cerebellar function. Annu.Rev.Neurosci. 2008.Vol31.<style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18275284">18275284</a></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text">Wouterlood FG and Mugnaini E. Cartwheel neurons of the dorsal cochlear nucleus: a Golgi-electron microscopic study in rat. J.Comp Neurol 1984 Jul20;227(1) <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/6088594">6088594</a></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFOliverBeckiusBishopKuwada1997" class="citation journal cs1">Oliver, Douglas L.; Beckius, Gretchen E.; Bishop, Deborah C.; Kuwada, Shigeyuki (2 June 1997). "Simultaneous anterograde labeling of axonal layers from lateral superior olive and dorsal cochlear nucleus in the inferior colliculus of cat". <i>The Journal of Comparative Neurology</i>. <b>382</b> (2): 215–229. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1002%2F%28SICI%291096-9861%2819970602%29382%3A2%3C215%3A%3AAID-CNE6%3E3.0.CO%3B2-6">10.1002/(SICI)1096-9861(19970602)382:2<215::AID-CNE6>3.0.CO;2-6</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/9183690">9183690</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Journal+of+Comparative+Neurology&rft.atitle=Simultaneous+anterograde+labeling+of+axonal+layers+from+lateral+superior+olive+and+dorsal+cochlear+nucleus+in+the+inferior+colliculus+of+cat&rft.volume=382&rft.issue=2&rft.pages=215-229&rft.date=1997-06-02&rft_id=info%3Adoi%2F10.1002%2F%28SICI%291096-9861%2819970602%29382%3A2%3C215%3A%3AAID-CNE6%3E3.0.CO%3B2-6&rft_id=info%3Apmid%2F9183690&rft.aulast=Oliver&rft.aufirst=Douglas+L.&rft.au=Beckius%2C+Gretchen+E.&rft.au=Bishop%2C+Deborah+C.&rft.au=Kuwada%2C+Shigeyuki&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADorsal+cochlear+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDavis2002" class="citation journal cs1">Davis, Kevin A. (April 2002). "Evidence of a Functionally Segregated Pathway From Dorsal Cochlear Nucleus to Inferior Colliculus". <i>Journal of Neurophysiology</i>. <b>87</b> (4): 1824–1835. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1152%2Fjn.00769.2001">10.1152/jn.00769.2001</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11929904">11929904</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Neurophysiology&rft.atitle=Evidence+of+a+Functionally+Segregated+Pathway+From+Dorsal+Cochlear+Nucleus+to+Inferior+Colliculus&rft.volume=87&rft.issue=4&rft.pages=1824-1835&rft.date=2002-04&rft_id=info%3Adoi%2F10.1152%2Fjn.00769.2001&rft_id=info%3Apmid%2F11929904&rft.aulast=Davis&rft.aufirst=Kevin+A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADorsal+cochlear+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFShofnerYoung1985" class="citation journal cs1">Shofner, WP; Young, ED (October 1985). "Excitatory/inhibitory response types in the cochlear nucleus: relationships to discharge patterns and responses to electrical stimulation of the auditory nerve". <i>Journal of Neurophysiology</i>. <b>54</b> (4): 917–39. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1152%2Fjn.1985.54.4.917">10.1152/jn.1985.54.4.917</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/4067627">4067627</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Neurophysiology&rft.atitle=Excitatory%2Finhibitory+response+types+in+the+cochlear+nucleus%3A+relationships+to+discharge+patterns+and+responses+to+electrical+stimulation+of+the+auditory+nerve.&rft.volume=54&rft.issue=4&rft.pages=917-39&rft.date=1985-10&rft_id=info%3Adoi%2F10.1152%2Fjn.1985.54.4.917&rft_id=info%3Apmid%2F4067627&rft.aulast=Shofner&rft.aufirst=WP&rft.au=Young%2C+ED&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADorsal+cochlear+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFYoungNelkenConley1995" class="citation journal cs1">Young, ED; Nelken, I; Conley, RA (February 1995). "Somatosensory effects on neurons in dorsal cochlear nucleus". <i>Journal of Neurophysiology</i>. <b>73</b> (2): 743–65. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1152%2Fjn.1995.73.2.743">10.1152/jn.1995.73.2.743</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/7760132">7760132</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Neurophysiology&rft.atitle=Somatosensory+effects+on+neurons+in+dorsal+cochlear+nucleus.&rft.volume=73&rft.issue=2&rft.pages=743-65&rft.date=1995-02&rft_id=info%3Adoi%2F10.1152%2Fjn.1995.73.2.743&rft_id=info%3Apmid%2F7760132&rft.aulast=Young&rft.aufirst=ED&rft.au=Nelken%2C+I&rft.au=Conley%2C+RA&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADorsal+cochlear+nucleus" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dorsal_cochlear_nucleus&action=edit&section=4" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" 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body</a></li> <li><a href="/wiki/Trigeminal_lemniscus" title="Trigeminal lemniscus">Trigeminal lemniscus</a> <ul><li><a href="/wiki/Dorsal_trigeminal_tract" title="Dorsal trigeminal tract">Dorsal trigeminal tract</a></li> <li><a href="/wiki/Anterior_trigeminothalamic_tract" class="mw-redirect" title="Anterior trigeminothalamic tract">Ventral trigeminal tract</a></li></ul></li> <li><a href="/wiki/Medial_lemniscus" title="Medial lemniscus">Medial lemniscus</a></li> <li><a href="/wiki/Lateral_lemniscus" title="Lateral lemniscus">Lateral lemniscus</a></li> <li><a href="/wiki/Central_tegmental_tract" title="Central tegmental tract">Central tegmental tract</a></li> <li><a href="/wiki/Medial_longitudinal_fasciculus" title="Medial longitudinal fasciculus">Medial longitudinal fasciculus</a> <ul><li><a href="/wiki/Vestibulo-oculomotor_fibers" title="Vestibulo-oculomotor fibers">Vestibulo-oculomotor fibers</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/White_matter" title="White matter">White</a>: Motor</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Inferior_cerebellar_peduncle" title="Inferior cerebellar peduncle">Inferior cerebellar peduncle</a> <ul><li><a href="/wiki/Vestibulocerebellar_tract" title="Vestibulocerebellar tract">Vestibulocerebellar tract</a></li></ul></li> <li><a href="/wiki/Medial_longitudinal_fasciculus" title="Medial longitudinal fasciculus">Medial longitudinal fasciculus</a> <ul><li><a href="/wiki/Vestibulospinal_tract" title="Vestibulospinal tract">Vestibulospinal tract</a></li> <li><a href="/wiki/Medial_vestibulospinal_tract" title="Medial vestibulospinal tract">Medial vestibulospinal tract</a></li> <li><a href="/wiki/Lateral_vestibulospinal_tract" title="Lateral vestibulospinal tract">Lateral vestibulospinal tract</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grey_matter" title="Grey matter">Grey</a>: <a href="/wiki/Cranial_nerve_nucleus" title="Cranial nerve nucleus">Cranial nuclei</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><span class="nobold"><i>afferent:</i></span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/General_somatic_afferent_fibers" class="mw-redirect" title="General somatic afferent fibers">GSA</a> <ul><li>Trigeminal <ul><li><a href="/wiki/Principal_sensory_nucleus_of_trigeminal_nerve" title="Principal sensory nucleus of trigeminal nerve">Principal</a></li> <li><a href="/wiki/Spinal_trigeminal_nucleus" title="Spinal trigeminal nucleus">Spinal</a></li></ul></li></ul></li> <li><a href="/wiki/Cochlear_nucleus" title="Cochlear nucleus">Cochlear nucleus</a> <ul><li><a class="mw-selflink selflink">Dorsal</a></li> <li><a href="/wiki/Anterior_cochlear_nucleus" class="mw-redirect" title="Anterior cochlear nucleus">Anterior</a></li></ul></li> <li><a href="/wiki/Vestibular_nuclei" title="Vestibular nuclei">Vestibular nuclei</a> <ul><li><a href="/wiki/Superior_vestibular_nucleus" title="Superior vestibular nucleus">Superior</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><span class="nobold"><i>efferent:</i></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Special_visceral_efferent" class="mw-redirect" title="Special visceral efferent">SVE</a>: <a href="/wiki/Trigeminal_motor_nucleus" title="Trigeminal motor nucleus">Trigeminal motor nucleus</a></li> <li><a href="/wiki/Facial_motor_nucleus" title="Facial motor nucleus">Facial motor nucleus</a></li> <li><a href="/wiki/General_somatic_efferent_fibers" class="mw-redirect" title="General somatic efferent fibers">GSE</a>: <a href="/wiki/Abducens_nucleus" title="Abducens nucleus">Abducens nucleus</a></li> <li><a href="/wiki/General_visceral_efferent_fibers" class="mw-redirect" title="General visceral efferent fibers">GVE</a>: <a href="/wiki/Superior_salivary_nucleus" class="mw-redirect" title="Superior salivary nucleus">Superior salivary nucleus</a></li> <li><a href="/wiki/Inferior_salivary_nucleus" class="mw-redirect" title="Inferior salivary nucleus">Inferior salivary nucleus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Grey: Other nuclei</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pedunculopontine_nucleus" title="Pedunculopontine nucleus">Pedunculopontine nucleus</a></li> <li><a href="/wiki/Apneustic_center" class="mw-redirect" title="Apneustic center">Apneustic center</a></li> <li><a href="/wiki/Pneumotaxic_center" class="mw-redirect" title="Pneumotaxic center">Pneumotaxic center</a></li> <li><a href="/wiki/Parabrachial_nuclei" title="Parabrachial nuclei">Parabrachial nuclei</a> <ul><li><a href="/wiki/Subparabrachial_nucleus" class="mw-redirect" title="Subparabrachial nucleus">Subparabrachial nucleus</a></li> <li><a href="/wiki/Medial_parabrachial_nucleus" class="mw-redirect" title="Medial parabrachial nucleus">Medial parabrachial nucleus</a></li> <li><a href="/wiki/Lateral_parabrachial_nucleus" class="mw-redirect" title="Lateral parabrachial nucleus">Lateral parabrachial nucleus</a></li></ul></li> <li><a href="/wiki/Superior_olivary_complex" title="Superior olivary complex">Superior olivary nucleus</a></li> <li><a href="/wiki/Locus_coeruleus" title="Locus coeruleus">Locus coeruleus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Ventral/<br />(<a href="/wiki/Basilar_part_of_pons" title="Basilar part of pons">base</a>)</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grey_matter" title="Grey matter">Grey</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pontine_nuclei" title="Pontine nuclei">Pontine nuclei</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/White_matter" title="White matter">White</a>: Motor/descending</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pyramidal_tracts" title="Pyramidal tracts">Corticospinal tract</a></li> <li><a href="/wiki/Corticobulbar_tract" title="Corticobulbar tract">Corticobulbar tract</a></li> <li><a href="/wiki/Corticopontine_fibers" title="Corticopontine fibers">Corticopontine fibers</a></li> <li><a href="/wiki/Middle_cerebellar_peduncle" title="Middle cerebellar peduncle">MCP</a> <ul><li><a href="/wiki/Pontocerebellar_fibers" title="Pontocerebellar fibers">Pontocerebellar fibers</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Surface</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Basilar_sulcus" title="Basilar sulcus">Basilar sulcus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other grey: Raphe/<br />reticular</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Reticular_formation" title="Reticular formation">Reticular formation</a> <ul><li><a href="/wiki/Caudal_pontine_reticular_nucleus" title="Caudal pontine reticular nucleus">Caudal</a></li> <li><a href="/wiki/Oral_pontine_reticular_nucleus" title="Oral pontine reticular nucleus">Oral</a></li> <li><a href="/wiki/Tegmental_pontine_reticular_nucleus" class="mw-redirect" title="Tegmental pontine reticular nucleus">Tegmental</a></li> <li><a href="/wiki/Paramedian_pontine_reticular_formation" title="Paramedian pontine reticular formation">Paramedian</a></li></ul></li> <li><a href="/wiki/Raphe_nuclei" title="Raphe nuclei">Raphe nuclei</a> <ul><li><a href="/wiki/Median_raphe_nucleus" title="Median raphe nucleus">Median</a></li></ul></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Physiology_of_balance_and_hearing" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Physiology_of_hearing_and_balance" title="Template:Physiology of hearing and balance"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Physiology_of_hearing_and_balance" title="Template talk:Physiology of hearing and balance"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Physiology_of_hearing_and_balance" title="Special:EditPage/Template:Physiology of hearing and balance"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Physiology_of_balance_and_hearing" style="font-size:114%;margin:0 4em">Physiology of <a href="/wiki/Balance_(ability)" title="Balance (ability)">balance</a> and <a href="/wiki/Hearing" title="Hearing">hearing</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Hearing</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Auditory_system" title="Auditory system">Auditory system</a></li> <li><a href="/wiki/Bone_conduction" title="Bone conduction">Bone conduction</a></li> <li><a href="/wiki/Otoacoustic_emission" title="Otoacoustic emission">Otoacoustic emission</a></li> <li><a href="/wiki/Tullio_phenomenon" title="Tullio phenomenon">Tullio phenomenon</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Neural_pathway" title="Neural pathway">Pathway</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Inner_ear" title="Inner ear">inner ear</a>:</i> <a href="/wiki/Hair_cell" title="Hair cell">Hair cells</a> → <a href="/wiki/Spiral_ganglion" title="Spiral ganglion">Spiral ganglion</a> → <a href="/wiki/Cochlear_nerve" title="Cochlear nerve">Cochlear nerve</a> <sup><a href="/wiki/Vestibulocochlear_nerve" title="Vestibulocochlear nerve">VIII</a></sup> →</li></ul> <ul><li><i><a href="/wiki/Pons" title="Pons">pons</a>:</i> <a href="/wiki/Cochlear_nucleus" title="Cochlear nucleus">Cochlear nucleus</a> (<a href="/wiki/Anterior_cochlear_nucleus" class="mw-redirect" title="Anterior cochlear nucleus">Anterior</a>, <a class="mw-selflink selflink">Dorsal</a>) → <a href="/wiki/Trapezoid_body" title="Trapezoid body">Trapezoid body</a> → <a href="/wiki/Superior_olivary_complex" title="Superior olivary complex">Superior olivary nuclei</a> →</li></ul> <ul><li><i><a href="/wiki/Midbrain" title="Midbrain">midbrain</a>:</i> <a href="/wiki/Lateral_lemniscus" title="Lateral lemniscus">Lateral lemniscus</a> → <a href="/wiki/Inferior_colliculus" title="Inferior colliculus">Inferior colliculi</a> →</li></ul> <ul><li><i><a href="/wiki/Thalamus" title="Thalamus">thalamus</a>:</i> <a href="/wiki/Medial_geniculate_nucleus" title="Medial geniculate nucleus">Medial geniculate nuclei</a> →</li></ul> <ul><li><i><a href="/wiki/Cerebrum" title="Cerebrum">cerebrum</a>:</i> <a href="/wiki/Acoustic_radiation" title="Acoustic radiation">Acoustic radiation</a> → <a href="/wiki/Primary_auditory_cortex" class="mw-redirect" title="Primary auditory cortex">Primary auditory cortex</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Balance</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Vestibular_system" title="Vestibular system">Vestibular system</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Pathway</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Inner_ear" title="Inner ear">inner ear</a>:</i> <a href="/wiki/Vestibular_nerve" title="Vestibular nerve">Vestibular nerve</a> <sup><a href="/wiki/Vestibulocochlear_nerve" title="Vestibulocochlear nerve">VIII</a></sup> →</li></ul> <ul><li><i><a href="/wiki/Pons" title="Pons">pons</a>:</i> <a href="/wiki/Vestibular_nuclei" title="Vestibular nuclei">Vestibular nuclei</a> (<a href="/wiki/Medial_vestibular_nucleus" title="Medial vestibular nucleus">Medial vestibular nucleus</a>, <a href="/wiki/Lateral_vestibular_nucleus" title="Lateral vestibular nucleus">Lateral vestibular nucleus</a>)</li></ul> <ul><li><i><a href="/wiki/Cerebellum" title="Cerebellum">cerebellum</a>:</i> <a href="/wiki/Flocculonodular_lobe" title="Flocculonodular lobe">Flocculonodular lobe</a></li></ul> <ul><li><i><a href="/wiki/Spinal_cord" title="Spinal cord">spinal cord</a>:</i> <a href="/wiki/Vestibulospinal_tract" title="Vestibulospinal tract">Vestibulospinal tract</a> (<a href="/wiki/Medial_vestibulospinal_tract" title="Medial vestibulospinal tract">Medial vestibulospinal tract</a>, <a href="/wiki/Lateral_vestibulospinal_tract" title="Lateral vestibulospinal tract">Lateral vestibulospinal tract</a>)</li></ul> <ul><li><i><a href="/wiki/Thalamus" title="Thalamus">thalamus</a>:</i> <a href="/wiki/Ventral_posterolateral_nucleus" title="Ventral posterolateral nucleus">Ventral posterolateral nucleus</a></li></ul> <ul><li><i><a href="/wiki/Cerebrum" title="Cerebrum">cerebrum</a>:</i> <a href="/wiki/Vestibular_cortex" title="Vestibular cortex">Vestibular cortex</a></li></ul> <ul><li><a href="/wiki/Vestibulo-oculomotor_fibers" title="Vestibulo-oculomotor fibers">Vestibulo-oculomotor fibers</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link 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