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ÉCOLOGIE - Encyclopædia Universalis

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L'individu dans son environnement </a> </li> <li class="list-group-item "> <a class="d-block" title="ÉCOLOGIE, Fonctionnement des populations" href="/encyclopedie/ecologie/2-fonctionnement-des-populations/"> 2. Fonctionnement des populations </a> </li> <li class="list-group-item "> <a class="d-block" title="ÉCOLOGIE, Organisation fonctionnelle des biocénoses" href="/encyclopedie/ecologie/3-organisation-fonctionnelle-des-biocenoses/"> 3. Organisation fonctionnelle des biocénoses </a> </li> <li class="list-group-item "> <a class="d-block" title="ÉCOLOGIE, Stabilité, adaptabilité et évolution des biocénoses" href="/encyclopedie/ecologie/4-stabilite-adaptabilite-et-evolution-des-biocenoses/"> 4. Stabilité, adaptabilité et évolution des biocénoses </a> </li> <li class="list-group-item "> <a class="d-block" title="ÉCOLOGIE, Méthodologie écologique" href="/encyclopedie/ecologie/5-methodologie-ecologique/"> 5. Méthodologie écologique </a> </li> <li class="list-group-item "> <a class="d-block" title="ÉCOLOGIE, Applications de l&#039;écologie" href="/encyclopedie/ecologie/6-applications-de-l-ecologie/"> 6. Applications de l'écologie </a> </li> <li class="list-group-item "> <a class="d-block" title="ÉCOLOGIE, Bibliographie" href="/encyclopedie/ecologie/7-bibliographie/"> 7. Bibliographie </a> </li> </ul> <button class="btn btn-sm btn-universalis" data-bs-toggle="collapse" data-bs-target="#free-toc" aria-expanded="false" title="Afficher / masquer le sommaire"> <i aria-hidden="true" class="fa fa-fw fa-eye"></i> Sommaire </button> </div> <h1 class="mb-1"> <strong>ÉCOLOGIE</strong> </h1> <ul class="pagination justify-content-center public-links "><li class="page-item active"><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 1" class="page-link" title="Abonnez-vous" data-loaded="true">1</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 2" class="page-link" title="Abonnez-vous" data-loaded="true">2</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 3" class="page-link" title="Abonnez-vous" data-loaded="true">3</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 4" class="page-link" title="Abonnez-vous" data-loaded="true">4</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 5" class="page-link" title="Abonnez-vous" data-loaded="true">5</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 6" class="page-link" title="Abonnez-vous" data-loaded="true">6</a></li></ul><div class="content paged type-article" ><div class="block_1" data-ucode="yC071134_0"><i class="anchor" id="yC071134_0"></i><div class="clearfix"></div><div class="media-container float-md-start me-2 col-md-3 text-center"><figure class="figure text-center w-100 mb-0 p-1" style="position: relative"><a data-ucode="PH995132" data-showMediaPlayer="1" class="protected media-reference-copy " title="Média : Ernst Haeckel" href="/media/ernst-haeckel-ph995132/"><img id="PH995132" loading="lazy" src="/typo3temp/assets/_processed_/a/f/csm_ph995132_29b8b812f6.webp" class="figure-img rounded mx-auto media_resized" alt="Ernst Haeckel - crédits : Hulton-Deutsch Collection/ Corbis/ Getty Images" data-description="En diffusant et en popularisant la théorie darwinienne et en énonçant la « loi de la récapitulation » (selon laquelle le développement embryonnaire serait une répétition de l'histoire évolutive), le zoologiste Ernst Haeckel (1834-1919) a joué un rôle important dans l'histoire des sciences de&hellip;" data-copyright="Hulton-Deutsch Collection/ Corbis/ Getty Images" width="864" height="1024" /></a><figcaption class="figure-caption"><p class="m-0 mt-1 fw-bold text-gray-700 text-truncate"> Ernst Haeckel </p><p class="m-0 copyright text-sm text-gray-700 text-truncate"> Hulton-Deutsch Collection/ Corbis/ Getty Images </p></figcaption></figure></div><p>Le terme écologie (du grec <em>oikos</em>, demeure, et <em>logos</em>, science) a été proposé par Ernst Haeckel en 1866 pour désigner la science qui étudie les rapports entre les organismes et le milieu où ils vivent. Cette définition reste encore valable, mais elle demande à être approfondie et précisée, car elle est trop générale. Pour la situer par rapport aux autres sciences biologiques, il est commode de considérer les divers niveaux d&#039;organisation de la matière vivante.</p><p>Le système le plus simple ayant toutes les caractéristiques fondamentales des êtres vivants est la cellule. Des cellules, associées en tissus et en organes, sont intégrées en organismes pluricellulaires, animaux ou végétaux.</p><p>À un niveau d&#039;intégration supérieur, les individus de certaines espèces peuvent constituer des <em>colonies</em>, comme chez les cœlentérés, ou des <em>sociétés</em>, comme chez les termites, les fourmis et les abeilles. À un niveau encore plus élevé se place, pour tous les êtres vivants, la <em>population</em>, ensemble des individus d&#039;une même espèce liés entre eux par des liens parentaux, c&#039;est-à-dire génétiques ; selon que la reproduction est sexuée ou non, ces liens sont évidemment différents.</p><p>Les populations des diverses espèces (bactériennes, végétales, animales...) groupées en un lieu déterminé forment un niveau d&#039;organisation plus complexe encore, la communauté biologique ou <em>biocénose</em> ; le milieu physique et chimique où celle-ci est installée est souvent appelé <em>biotope</em> et leur ensemble est un <em>écosystème</em>. Le niveau ultime d&#039;organisation du <a data-bs-toggle="popover" data-bs-content="Le terme de monde est utilisé dans des contextes très divers pour indiquer, d&#039;une manière ou de l&#039;autre, une totalité d&#039;appartenance. Ainsi on parle du « monde grec » ou du « monde de la Renaissance », pour désigner des ensembles culturels spécifiques, situés dans notre passé historique et auxquels nous avons accès à travers un certain nombre d&#039;œuvres qui sont venues jusqu&#039;à ..." title="Article : MONDE" href="/encyclopedie/monde/">monde</a> vivant est constitué par l&#039;ensemble des <a data-bs-toggle="popover" data-bs-content="Le terme « écosystème » désigne un groupe d&#039;êtres vivants – parfois désigné sous le nom de biocénose (animaux, végétaux et microorganismes) – et de composantes physiques et chimiques – parfois regroupées sous le terme de biotope . Il correspond à une vision intégrative de la nature, centrée sur le concept d’interaction : toute composante de l’écosystème, vi..." title="Article : ÉCOSYSTÈMES" href="/encyclopedie/ecosystemes/">écosystèmes</a> de la planète, c&#039;est-à-dire la <em>biosphère</em>.</p><p>La structure et le fonctionnement des cellules constituent l&#039;objet de la cytologie et de la physiologie cellulaire ; la structure et le fonctionnement des êtres pluricellulaires concernent l&#039;histologie, l&#039;anatomie et la physiologie des organismes. L&#039;écologie a donc pour objet essentiel l&#039;étude des niveaux supérieurs d&#039;organisation de la matière vivante, de la population monospécifique à l&#039;écosystème et à la biosphère : comme pour les niveaux inférieurs, mais cette fois à l&#039;échelle de systèmes particulièrement complexes, il s&#039;agit de décrire des structures, de comprendre des fonctionnements et de reconstituer des évolutions. On ne peut toutefois saisir les phénomènes jouant à un certain niveau d&#039;intégration sans connaître ceux qui interviennent aux niveaux inférieurs : de même que l&#039;organisation des écosystèmes ne peut être comprise qu&#039;en tenant compte du fonctionnement des populations, celui-ci ne peut l&#039;être sans référence aux relations que chaque <a data-bs-toggle="popover" data-bs-content="La notion d&#039;individualité ne se laisse définir ni par les procédés étymologiques habituels aux philosophes depuis que Platon en a établi la coutume dans le Cratyle , ni par une axiomatique rationnelle. Lorsqu&#039;on s&#039;y attache, c&#039;est pour trouver des différences, lui opposer d&#039;autres notions ; tout se passe comme si elle relevait de plusieurs champs sémantiques, sans qu&#039;au premier abord i..." title="Article : INDIVIDU" href="/encyclopedie/individu/">individu</a> entretient avec le milieu.</p><p>Ces relations entre les individus d&#039;une même espèce et le milieu où ils vivent constituent le domaine de l&#039;écologie des populations (autrefois appelée autoécologie). Elles concernent à la fois les individus, pour eux-mêmes, et les populations qu&#039;ils forment. Dans cette perspective, le milieu est conçu comme un ensemble de facteurs dont on distingue deux types : les facteurs <em>abiotiques</em>, liés au milieu physique et chimique, et les facteurs <em>biotiques</em>, liés aux êtres vivants présents dans l&#039;écosystème étudié.</p><p>L&#039;écologie des communautés (autrefois appelée synécologie ou <em>biocénotique</em>) envisage essentiellement la structure et le fonctionnement des écosystèmes, mais son champ s&#039;étend également aux complexes d&#039;écosystèmes, associés par exemple dans un même bassin versant, et finalement à la <a data-bs-toggle="popover" data-bs-content="L&#039;idée de biosphère, ensemble des êtres vivants qui peuplent notre planète, remonte à J.-B. Lamarck. C&#039;est ensuite le géologue autrichien Suess qui, en 1875, a mis en parallèle le terme de biosphère avec ceux d&#039;hydrosphère, d&#039;atmosphère et de lithosphère . Il étendait en fait le concept à tout ce qui constituait ou avait constitué le monde vivant, en y incluant donc aussi, par ..." title="Article : BIOSPHÈRE" href="/encyclopedie/biosphere/">biosphère</a>.</p><p>Discipline biologique, l'écologie ne se contente pas de décrire des structures et d'analyser des fonctionnements : elle tente de les interpréter dans une optique évolutionniste et doit considérer les systèmes qu'elle étudie – populations, écosystèmes – à différentes échelles de temps.<a href="javascript:" title="Lire la suite" class="read-more ms-1">[...]</a></p></div><div class="clearfix"></div></div><ul class="pagination justify-content-center public-links "><li class="page-item active"><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 1" class="page-link" title="Abonnez-vous" data-loaded="true">1</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 2" class="page-link" title="Abonnez-vous" data-loaded="true">2</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 3" class="page-link" title="Abonnez-vous" data-loaded="true">3</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 4" class="page-link" title="Abonnez-vous" data-loaded="true">4</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 5" class="page-link" title="Abonnez-vous" data-loaded="true">5</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 6" class="page-link" title="Abonnez-vous" data-loaded="true">6</a></li></ul><div class="folio-block"><p class="folio-block-title text-center text-uc"><strong>La suite de cet article est accessible aux abonnés</strong></p><ul class="list-unstyled"><li><i class="fa fa-fw fa-check-circle"></i> Des contenus variés, complets et fiables</li><li><i class="fa fa-fw fa-check-circle"></i> Accessible sur tous les écrans</li><li><i class="fa fa-fw fa-check-circle"></i> Pas de publicité</li></ul><p class="text-center"><a href="/abonnement/" class="btn btn-cta text-uc" id="folio-button" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="Block bottom">Découvrez nos offres </a></p><p class="text-center text-sm"> Déjà abonné ? <a href="/identification/" id="folio-login">Se connecter</a></p></div><div id="ad_mpuslot" class="pub-container text-center"><div class="pub-placeholder"></div></div><section class="mt-3 mb-5"><i class="anchor" id="C071134_AUTHORS"></i><p class="title-minor">Écrit par</p><ul class="list-group"><li class="list-group-item" data-dc-author="BLANDIN, Patrick"><a title="Auteur : Patrick BLANDIN" rel="author" href="/auteurs/patrick-blandin/"><strong>Patrick BLANDIN</strong></a> : professeur au Muséum national d'histoire naturelle, directeur de la Grande Galerie de l'évolution </li><li class="list-group-item" data-dc-author="COUVET, Denis"><a title="Auteur : Denis COUVET" rel="author" href="/auteurs/denis-couvet/"><strong>Denis COUVET</strong></a> : professeur au Muséum national d'histoire naturelle, Paris </li><li class="list-group-item" data-dc-author="LAMOTTE, Maxime"><a title="Auteur : Maxime LAMOTTE" rel="author" href="/auteurs/maxime-lamotte/"><strong>Maxime LAMOTTE</strong></a> : professeur honoraire à l'université de Paris-VI-Pierre-et-Marie-Curie (faculté des sciences), ancien directeur du laboratoire de zoologie de l'École normale supérieure </li><li class="list-group-item" data-dc-author="SACCHI, Cesare F."><a title="Auteur : Cesare F. SACCHI" rel="author" href="/auteurs/cesare-f.-sacchi/"><strong>Cesare F. SACCHI</strong></a> : professeur d'écologie à l'université de Paris-VI, directeur de l'Institut d'écologie et d'éthologie de Pavie </li></ul></section><section class="mb-5"><div class="text-center mb-3 d-print-none"><a class="d-inline-flex flex-row justify-content-center align-items-center btn btn-light btn-large text-start" title="Carte mentale : ÉCOLOGIE" href="/carte-mentale/ecologie/"><i class="fa fa-sitemap fa-rotate-270 me-1 fs-1 text-universalis"></i><p class="m-0"><strong class="fs-4">Carte mentale</strong><br /> Élargissez votre recherche dans Universalis </p></a></div></section> <section class="mb-5 article-classification" data-print-label="la classification" data-print-target=".article-classification"> <i class="anchor" id="yC071134_ROOT_LINES"></i> <p class="title-minor">Classification</p> <nav aria-label="Classification 1"> <ol class="breadcrumb"> <li class="breadcrumb-item"> <a title="Classification : Sciences de la vie" href="/classification/sciences-de-la-vie/"> Sciences de la vie </a> </li> <li class="breadcrumb-item active" data-dc-subject="Écologie"> <a title="Classification : Écologie" href="/classification/sciences-de-la-vie/ecologie/"> Écologie </a> </li> </ol> </nav> <nav aria-label="Classification 2"> <ol class="breadcrumb"> <li class="breadcrumb-item"> <a title="Classification : Sciences de la vie" href="/classification/sciences-de-la-vie/"> Sciences de la vie </a> </li> <li class="breadcrumb-item"> <a title="Classification : Écologie" href="/classification/sciences-de-la-vie/ecologie/"> Écologie </a> </li> <li class="breadcrumb-item active" data-dc-subject="Écophysiologie"> <a title="Classification : Écophysiologie" href="/classification/sciences-de-la-vie/ecologie/ecophysiologie/"> Écophysiologie </a> </li> </ol> </nav> <nav aria-label="Classification 3"> <ol class="breadcrumb"> <li class="breadcrumb-item"> <a title="Classification : Sciences de la vie" href="/classification/sciences-de-la-vie/"> Sciences de la vie </a> </li> <li class="breadcrumb-item"> <a title="Classification : Écologie" href="/classification/sciences-de-la-vie/ecologie/"> Écologie </a> </li> <li class="breadcrumb-item active" data-dc-subject="Biologie et dynamique des populations"> <a title="Classification : Biologie et dynamique des populations" href="/classification/sciences-de-la-vie/ecologie/biologie-et-dynamique-des-populations/"> Biologie et dynamique des populations </a> </li> </ol> </nav> <nav aria-label="Classification 4"> <ol class="breadcrumb"> <li class="breadcrumb-item"> <a title="Classification : Sciences de la vie" href="/classification/sciences-de-la-vie/"> Sciences de la vie </a> </li> <li class="breadcrumb-item"> <a title="Classification : Écologie" href="/classification/sciences-de-la-vie/ecologie/"> Écologie </a> </li> <li class="breadcrumb-item active" data-dc-subject="Structure et fonctionnement des écosystèmes"> <a title="Classification : Structure et fonctionnement des écosystèmes" href="/classification/sciences-de-la-vie/ecologie/structure-et-fonctionnement-des-ecosystemes/"> Structure et fonctionnement des écosystèmes </a> </li> </ol> </nav> <nav aria-label="Classification 5"> <ol class="breadcrumb"> <li class="breadcrumb-item"> <a title="Classification : Sciences de la vie" href="/classification/sciences-de-la-vie/"> Sciences de la vie </a> </li> <li class="breadcrumb-item active" data-dc-subject="Sciences de la vie: instruments et techniques expérimentales"> <a title="Classification : Sciences de la vie: instruments et techniques expérimentales" href="/classification/sciences-de-la-vie/sciences-de-la-vie-instruments-et-techniques-experimentales/"> Sciences de la vie: instruments et techniques expérimentales </a> </li> </ol> </nav> <nav aria-label="Classification 6"> <ol class="breadcrumb"> <li class="breadcrumb-item"> <a title="Classification : Sciences de la vie" href="/classification/sciences-de-la-vie/"> Sciences de la vie </a> </li> <li class="breadcrumb-item"> <a title="Classification : Écologie" href="/classification/sciences-de-la-vie/ecologie/"> Écologie </a> </li> <li class="breadcrumb-item active" data-dc-subject="Écologie appliquée"> <a title="Classification : Écologie appliquée" href="/classification/sciences-de-la-vie/ecologie/ecologie-appliquee/"> Écologie appliquée </a> </li> </ol> </nav> </section> <section class="mb-5 d-print-none"> <i class="anchor" id="yC071134_MEDIAS"></i> <p class="title-minor">Médias</p> <div class="row"> <div class="col-12 col-md-4 text-center p-1"> <figure class="figure text-center w-100 mb-0 p-1" style="position: relative"> <a data-ucode="PH995132" data-showMediaPlayer="1" class="protected media-reference-copy " title="Média : Ernst Haeckel" href="/media/ernst-haeckel-ph995132/"> <img id="PH995132" loading="lazy" src="/typo3temp/assets/_processed_/a/f/csm_ph995132_29b8b812f6.webp" class="figure-img rounded mx-auto media_resized" alt="Ernst Haeckel - crédits : Hulton-Deutsch Collection/ Corbis/ Getty Images" data-description="En diffusant et en popularisant la théorie darwinienne et en énonçant la « loi de la récapitulation » (selon laquelle le développement embryonnaire serait une répétition de l'histoire évolutive), le zoologiste Ernst Haeckel (1834-1919) a joué un rôle important dans l'histoire des sciences de&hellip;" data-copyright="Hulton-Deutsch Collection/ Corbis/ Getty Images" width="864" height="1024" /> </a> <figcaption class="figure-caption"> <p class="m-0 mt-1 fw-bold text-gray-700 text-truncate"> Ernst Haeckel </p> <p class="m-0 copyright text-sm text-gray-700 text-truncate"> Hulton-Deutsch Collection/ Corbis/ Getty Images </p> </figcaption> </figure> </div> <div class="col-12 col-md-4 text-center p-1"> <figure class="figure text-center w-100 mb-0 p-1" style="position: relative"> <a data-ucode="V070862A" data-showMediaPlayer="1" class="protected media-reference-copy " title="Média : Flux d&#039;énergie chez les animaux et les végétaux" href="/media/flux-denergie-chez-les-animaux-et-les-vegetaux-v070862a/"> <img id="V070862A" loading="lazy" src="/typo3temp/assets/_processed_/f/5/csm_v070862a_28f2508da1.webp" class="figure-img rounded mx-auto media_resized" alt="Flux d&#039;énergie chez les animaux et les végétaux - crédits : Encyclopædia Universalis France" data-description="Répartition des flux d'énergie : a) chez les végétaux et b) chez les animaux." data-copyright="Encyclopædia Universalis France" width="2000" height="1160" /> </a> <figcaption class="figure-caption"> <p class="m-0 mt-1 fw-bold text-gray-700 text-truncate"> Flux d'énergie chez les animaux et les végétaux </p> <p class="m-0 copyright text-sm text-gray-700 text-truncate"> Encyclopædia Universalis France </p> </figcaption> </figure> </div> <div class="col-12 col-md-4 text-center p-1"> <figure class="figure text-center w-100 mb-0 p-1" style="position: relative"> <a data-ucode="V070862B" data-showMediaPlayer="1" class="protected media-reference-copy " title="Média : Métabolisme d&#039;animaux homéothermes" href="/media/metabolisme-danimaux-homeothermes-v070862b/"> <img id="V070862B" loading="lazy" src="/typo3temp/assets/_processed_/f/4/csm_v070862b_8a9227154c.webp" class="figure-img rounded mx-auto media_resized" alt="Métabolisme d&#039;animaux homéothermes - crédits : Encyclopædia Universalis France" data-description="Variation du métabolisme de base A d'animaux homéothermes en fonction de leur poids W." data-copyright="Encyclopædia Universalis France" width="1220" height="2000" /> </a> <figcaption class="figure-caption"> <p class="m-0 mt-1 fw-bold text-gray-700 text-truncate"> Métabolisme d'animaux homéothermes </p> <p class="m-0 copyright text-sm text-gray-700 text-truncate"> Encyclopædia Universalis France </p> </figcaption> </figure> </div> </div> <div class="text-center"> <a class="btn btn-sm btn-primary" rel="related" href="/medias/ecologie/"> Afficher les 15 médias de l'article <span class="rg">&Eacute;COLOGIE</span> <i aria-hidden="true" class="fa fa-fw fa-chevron-right"></i> </a> </div> </section> <section class="mb-5 d-print-none learn-more"> <i class="anchor" id="yC071134_REFERENCES"></i> <p class="title-minor">Autres références</p> <ul class="list-group"> <li class="list-group-item"> <li class="list-group-item"> <h3 class="fs-5 m-0 mt-1"> <a href="/encyclopedie/ecologie-et-societe/"> <span class="rg">&Eacute;COLOGIE ET SOCI&Eacute;T&Eacute;</span> </a> </h3> <ul class="list-inline list-inline-bullet mb-1 text-muted"> <li class="list-inline-item text-sm">Écrit par <span rel="author">Robert BARBAULT</span> </li> <li class="list-inline-item text-sm">7 804 mots</li> <li class="list-inline-item text-sm">5 médias </li> </ul> <div class="d-none d-md-inline-block float-start me-3"> <a class="protected" title="Article : &amp;Eacute;COLOGIE ET SOCI&amp;Eacute;T&amp;Eacute;" href="/encyclopedie/ecologie-et-societe/"> <img class="rounded shadow" loading="lazy" alt="Déforestation à Guiroutou, Côte d&#039;Ivoire - 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Ces conditions idéales sont loin d'être réunies dans les milieux naturels... </div> </li> <li class="list-group-item"> <h3 class="fs-5 m-0 mt-1"> <a href="/encyclopedie/anthropologie/#c7588"> <span class="rg">ANTHROPOLOGIE</span> </a> </h3> <ul class="list-inline list-inline-bullet mb-1 text-muted"> <li class="list-inline-item text-sm">Écrit par <span rel="author">Élisabeth COPET-ROUGIER</span> et <span rel="author">Christian GHASARIAN</span> </li> <li class="list-inline-item text-sm">16 158 mots</li> <li class="list-inline-item text-sm">1 média </li> </ul> <div class="d-none d-md-inline-block float-start me-3"> <a class="protected" title="Article : ANTHROPOLOGIE" href="/encyclopedie/anthropologie/"> <img class="rounded shadow" loading="lazy" alt="Claude Lévi-Strauss - crédits : Marc Gantier/ Gamma-Rapho/ Getty Images" width="125" height="125" src="/typo3temp/assets/_processed_/f/8/csm_pa082775_36e7ed6627.webp"/> </a> </div> <div class="snippet-content mt-3"> ...des bases matérielles des sociétés humaines en relation avec leur adaptation au milieu. 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