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data-event-name="pinnable-header.vector-toc.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">hide</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">(Top)</div> </a> </li> <li id="toc-Carlson's_Trophic_State_Index" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Carlson's_Trophic_State_Index"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Carlson's Trophic State Index</span> </div> </a> <button aria-controls="toc-Carlson's_Trophic_State_Index-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Carlson's Trophic State Index subsection</span> </button> <ul id="toc-Carlson's_Trophic_State_Index-sublist" class="vector-toc-list"> <li id="toc-Index_variable" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Index_variable"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.1</span> <span>Index variable</span> </div> </a> <ul id="toc-Index_variable-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Trophic_classifications" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Trophic_classifications"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Trophic classifications</span> </div> </a> <button aria-controls="toc-Trophic_classifications-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Trophic classifications subsection</span> </button> <ul id="toc-Trophic_classifications-sublist" class="vector-toc-list"> <li id="toc-Oligotrophic" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Oligotrophic"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1</span> <span>Oligotrophic</span> </div> </a> <ul id="toc-Oligotrophic-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Mesotrophic" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Mesotrophic"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.2</span> <span>Mesotrophic</span> </div> </a> <ul id="toc-Mesotrophic-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Eutrophic_and_hypertrophic" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Eutrophic_and_hypertrophic"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.3</span> <span>Eutrophic and hypertrophic</span> </div> </a> <ul id="toc-Eutrophic_and_hypertrophic-sublist" class="vector-toc-list"> <li id="toc-Eutrophic" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Eutrophic"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.3.1</span> <span>Eutrophic</span> </div> </a> <ul id="toc-Eutrophic-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Hypertrophic" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Hypertrophic"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.3.2</span> <span>Hypertrophic</span> </div> </a> <ul id="toc-Hypertrophic-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> </ul> </li> <li id="toc-Trophic_index_drivers" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Trophic_index_drivers"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Trophic index drivers</span> </div> </a> <button 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id="toc-Management_targets" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Management_targets"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Management targets</span> </div> </a> <ul id="toc-Management_targets-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Notes" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Notes"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>Notes</span> </div> </a> <ul id="toc-Notes-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" 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searchaux" style="display:none">Measure of the ability of water to sustain biological productivity</div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Lake_George_from_village_beach.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e1/Lake_George_from_village_beach.jpg/240px-Lake_George_from_village_beach.jpg" decoding="async" width="240" height="157" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e1/Lake_George_from_village_beach.jpg/360px-Lake_George_from_village_beach.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e1/Lake_George_from_village_beach.jpg/480px-Lake_George_from_village_beach.jpg 2x" data-file-width="2292" data-file-height="1497" /></a><figcaption><a href="/wiki/Lake_George_(lake),_New_York" title="Lake George (lake), New York">Lake George, New York</a>, an oligotrophic lake</figcaption></figure> <p>The <b>Trophic State Index</b> (<b>TSI</b>) is a classification system designed to rate water bodies based on the amount of <a href="/wiki/Biological_productivity" class="mw-redirect" title="Biological productivity">biological productivity</a> they sustain.<sup id="cite_ref-USFWaterAtlas_1-0" class="reference"><a href="#cite_note-USFWaterAtlas-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Although the term "trophic index" is commonly applied to lakes, any surface water body may be indexed. </p><p>The TSI of a water body is rated on a scale from zero to one hundred.<sup id="cite_ref-USFWaterAtlas_1-1" class="reference"><a href="#cite_note-USFWaterAtlas-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Under the TSI scale, water bodies may be defined as:<sup id="cite_ref-USFWaterAtlas_1-2" class="reference"><a href="#cite_note-USFWaterAtlas-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <ul><li><b>oligotrophic</b> (TSI 0–40, having the least amount of biological productivity, "good" water quality);</li> <li><b>mesotrophic</b> (TSI 40–60, having a moderate level of biological productivity, "fair" water quality); or</li> <li><b>eutrophic</b> to <b>hypereutrophic</b> (TSI 60–100, having the highest amount of biological productivity, "poor" water quality).</li></ul> <p>The quantities of <a href="/wiki/Nitrogen" title="Nitrogen">nitrogen</a>, <a href="/wiki/Phosphorus" title="Phosphorus">phosphorus</a>, and other biologically useful nutrients are the primary determinants of a water body's TSI. Nutrients such as nitrogen and phosphorus tend to be <a href="/wiki/Limiting_resource" class="mw-redirect" title="Limiting resource">limiting resources</a> in standing water bodies, so increased concentrations tend to result in increased plant growth, followed by corollary increases in subsequent <a href="/wiki/Trophic_level" title="Trophic level">trophic levels</a>.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>a<span class="cite-bracket">]</span></a></sup> Consequently, trophic index may sometimes be used to make a rough estimate of biological condition of water bodies.<sup id="cite_ref-USEPA_3-0" class="reference"><a href="#cite_note-USEPA-3"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Carlson's_Trophic_State_Index"><span id="Carlson.27s_Trophic_State_Index"></span>Carlson's Trophic State Index</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=1" title="Edit section: Carlson's Trophic State Index"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Carlson's index was proposed by Robert Carlson in his 1977 seminal paper, "A trophic state index for lakes".<sup id="cite_ref-Carlson1977_4-0" class="reference"><a href="#cite_note-Carlson1977-4"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> It is one of the more commonly used trophic indices and is the trophic index used by the <a href="/wiki/United_States_Environmental_Protection_Agency" title="United States Environmental Protection Agency">United States Environmental Protection Agency</a>.<sup id="cite_ref-USEPA_3-1" class="reference"><a href="#cite_note-USEPA-3"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The <i>trophic state</i> is defined as the total weight of <a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">biomass</a> in a given water body at the time of measurement. Because they are of public concern, the Carlson index uses the algal biomass as an objective classifier of a lake or other water body's trophic status.<sup id="cite_ref-Carlson1977_4-1" class="reference"><a href="#cite_note-Carlson1977-4"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> According to the US EPA, the Carlson Index should only be used with lakes that have relatively few rooted plants and non-algal turbidity sources.<sup id="cite_ref-USEPA_3-2" class="reference"><a href="#cite_note-USEPA-3"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Index_variable">Index variable</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=2" title="Edit section: Index variable"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Because they tend to correlate, three independent variables can be used to calculate the Carlson Index: <i><a href="/wiki/Chlorophyll" title="Chlorophyll">chlorophyll pigments</a></i>, <i>total phosphorus</i> and <i><a href="/wiki/Secchi_depth" class="mw-redirect" title="Secchi depth">Secchi depth</a></i>. Of these three, chlorophyll will probably yield the most accurate measures, as it is the most accurate predictor of biomass. Phosphorus may be a more accurate estimation of a water body's summer trophic status than chlorophyll if the measurements are made during the winter. Finally, the Secchi depth is probably the least accurate measure, but also the most affordable and expedient one. Consequently, citizen monitoring programs and other volunteer or large-scale surveys will often use the Secchi depth. By translating the Secchi transparency values to a log base 2 scale, each successive doubling of biomass is represented as a whole integer index number.<sup id="cite_ref-Carlson1996_5-0" class="reference"><a href="#cite_note-Carlson1996-5"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> The Secchi depth, which measures water transparency, indicates the concentration of dissolved and particulate material in the water, which in turn can be used to derive the biomass. This relationship is expressed in the following equation: </p> <dl><dd><span class="mwe-math-element"><span class="mwe-math-mathml-inline mwe-math-mathml-a11y" style="display: none;"><math xmlns="http://www.w3.org/1998/Math/MathML" alttext="{\displaystyle \left({\frac {1}{z}}\right)\left(\ln {\frac {I_{0}}{I_{z}}}\right)=k_{w}+\alpha C}"> <semantics> <mrow class="MJX-TeXAtom-ORD"> <mstyle displaystyle="true" scriptlevel="0"> <mrow> <mo>(</mo> <mrow class="MJX-TeXAtom-ORD"> <mfrac> <mn>1</mn> <mi>z</mi> </mfrac> </mrow> <mo>)</mo> </mrow> <mrow> <mo>(</mo> <mrow> <mi>ln</mi> <mo>⁡</mo> <mrow class="MJX-TeXAtom-ORD"> <mfrac> <msub> <mi>I</mi> <mrow class="MJX-TeXAtom-ORD"> <mn>0</mn> </mrow> </msub> <msub> <mi>I</mi> <mrow class="MJX-TeXAtom-ORD"> <mi>z</mi> </mrow> </msub> </mfrac> </mrow> </mrow> <mo>)</mo> </mrow> <mo>=</mo> <msub> <mi>k</mi> <mrow class="MJX-TeXAtom-ORD"> <mi>w</mi> </mrow> </msub> <mo>+</mo> <mi>α</mi> <mi>C</mi> </mstyle> </mrow> <annotation encoding="application/x-tex">{\displaystyle \left({\frac {1}{z}}\right)\left(\ln {\frac {I_{0}}{I_{z}}}\right)=k_{w}+\alpha C}</annotation> </semantics> </math></span><img src="https://wikimedia.org/api/rest_v1/media/math/render/svg/1c75ee66b92b12104f906ba8d62cf88f105cd11c" class="mwe-math-fallback-image-inline mw-invert skin-invert" aria-hidden="true" style="vertical-align: -2.505ex; width:26.281ex; height:6.176ex;" alt="{\displaystyle \left({\frac {1}{z}}\right)\left(\ln {\frac {I_{0}}{I_{z}}}\right)=k_{w}+\alpha C}"></span></dd> <dd>where <i>z</i> = the depth at which the disk disappears,</dd> <dd><i>I</i><sub>0</sub> is the intensity of light striking the water's surface,</dd> <dd><i>I</i><sub><i>z</i></sub> is about 10% of <i>I</i><sub>0</sub> and is considered a constant,</dd> <dd><i>k</i><sub><i>w</i></sub> is a coefficient for the attenuation of light by water and dissolved substances,</dd> <dd><i>α</i> is treated as a constant with the units of square meters per milligram and</dd> <dd><i>C</i> is the concentration of particulate matter in units for milligrams per cubic meter.<sup id="cite_ref-Carlson1977_4-2" class="reference"><a href="#cite_note-Carlson1977-4"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></dd></dl> <div class="mw-heading mw-heading2"><h2 id="Trophic_classifications">Trophic classifications</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=3" title="Edit section: Trophic classifications"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>A lake is usually classified as being in one of three possible classes: <i>oligotrophic</i>, <i>mesotrophic</i> or <i>eutrophic</i>. Lakes with extreme trophic indices may also be considered <i>hyperoligotrophic</i> or <i>hypereutrophic</i> (also "hypertrophic"). The table below demonstrates how the index values translate into trophic classes. </p> <table class="wikitable" style="text-align:center;"> <caption><i>Relationships between Trophic State Index, chlorophyll, phosphorus, Secchi depth, and trophic class (after Carlson 1996)</i><sup id="cite_ref-Carlson1996_5-1" class="reference"><a href="#cite_note-Carlson1996-5"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </caption> <tbody><tr> <th>Trophic State Index </th> <th>Chlorophyll (μg/L) </th> <th>Phosphorus (μg/L) </th> <th>Secchi depth (m) </th> <th>Trophic Class </th></tr> <tr> <td>< 30—40 </td> <td>0—2.6 </td> <td>0—12 </td> <td>> 8—4 </td> <td align="left">Oligotrophic or hipotrophic </td></tr> <tr> <td>40—50 </td> <td>2.6—7.3 </td> <td>12—24 </td> <td>4—2 </td> <td align="left">Mesotrophic </td></tr> <tr> <td>50—70 </td> <td>7.3—56 </td> <td>24—96 </td> <td>2—0.5 </td> <td align="left">Eutrophic </td></tr> <tr> <td>70—100+ </td> <td>56—155+ </td> <td>96—384+ </td> <td>0.5— < 0.25 </td> <td align="left">Hypertrophic </td></tr></tbody></table> <p><span class="anchor" id="Oligotrophic"></span> Oligotrophic lakes generally host very little or no <a href="/wiki/Aquatic_plant" title="Aquatic plant">aquatic vegetation</a> and are relatively clear, while eutrophic lakes tend to host large quantities of organisms, including algal blooms. Each trophic class supports different types of fish and other organisms, as well. If the algal biomass in a lake or other water body reaches too high a concentration (say >80 TSI), massive <a href="/wiki/Fish_die-off" class="mw-redirect" title="Fish die-off">fish die-offs</a> may occur as decomposing biomass deoxygenates the water. </p> <div class="mw-heading mw-heading3"><h3 id="Oligotrophic">Oligotrophic</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=4" title="Edit section: Oligotrophic"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:20161001_Kurtkowiec_i_Czerwone_Stawy_G%C4%85sienicowe_1730.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d9/20161001_Kurtkowiec_i_Czerwone_Stawy_G%C4%85sienicowe_1730.jpg/220px-20161001_Kurtkowiec_i_Czerwone_Stawy_G%C4%85sienicowe_1730.jpg" decoding="async" width="220" height="177" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d9/20161001_Kurtkowiec_i_Czerwone_Stawy_G%C4%85sienicowe_1730.jpg/330px-20161001_Kurtkowiec_i_Czerwone_Stawy_G%C4%85sienicowe_1730.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d9/20161001_Kurtkowiec_i_Czerwone_Stawy_G%C4%85sienicowe_1730.jpg/440px-20161001_Kurtkowiec_i_Czerwone_Stawy_G%C4%85sienicowe_1730.jpg 2x" data-file-width="3924" data-file-height="3160" /></a><figcaption><a href="/wiki/Kurtkowiec_Lake" title="Kurtkowiec Lake">Kurtkowiec Lake</a>, an oligotrophic lake in the <a href="/wiki/Tatra_Mountains" title="Tatra Mountains">Tatra Mountains</a> of southern <a href="/wiki/Poland" title="Poland">Poland</a></figcaption></figure> <p><a href="/wiki/Limnology" title="Limnology">Limnologists</a> use the term "<a href="/wiki/Oligotroph" title="Oligotroph">oligotrophic</a>" or "hipotrophic" to describe lakes that have low <a href="/wiki/Primary_productivity" class="mw-redirect" title="Primary productivity">primary productivity</a> due to <a href="/wiki/Nutrient" title="Nutrient">nutrient</a> deficiency. (This contrasts against eutrophic lakes, which are highly productive due to an ample supply of nutrients, as can arise from human activities such as agriculture in the watershed.) </p><p>Oligotrophic lakes are most common in cold, sparsely developed regions that are underlain by crystalline <a href="/wiki/Igneous" class="mw-redirect" title="Igneous">igneous</a>, <a href="/wiki/Granitic" class="mw-redirect" title="Granitic">granitic</a> bedrock. Due to their low <a href="/wiki/Algal" class="mw-redirect" title="Algal">algal</a> production, these lakes consequently have very clear waters, with high <a href="/wiki/Drinking_water" title="Drinking water">drinking-water</a> quality. </p><p>Lakes that have intermixing of their layers are classified into the category of <a href="/wiki/Holomictic_lake" title="Holomictic lake">holomictic</a>, whereas lakes that do not have interlayer mixing are permanently stratified and thus are termed <a href="/wiki/Meromictic_lake" title="Meromictic lake">meromictic</a>. </p><p>Generally, in a holomictic lake, during the fall, the cooling of the epilimnion reduces lake stratification, thereby allowing for mixing to occur. Winds aid in this process.<sup id="cite_ref-Dodds,_Walter_K._Walter_Kennedy,_1958–2010_6-0" class="reference"><a href="#cite_note-Dodds,_Walter_K._Walter_Kennedy,_1958–2010-6"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> Thus it is the deep mixing of lakes (which occurs most often during the fall and early winter, in holomictic lakes of the monomictic subtype) that allows oxygen to be transported from the <a href="/wiki/Epilimnion" title="Epilimnion">epilimnion</a> to the hypolimnion.<sup id="cite_ref-sciencedirect.com_7-0" class="reference"><a href="#cite_note-sciencedirect.com-7"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p>In this way, oligotrophic lakes can have significant <a href="/wiki/Oxygen" title="Oxygen">oxygen</a> down to the depth to which the aforementioned seasonal mixing occurs, but they will be oxygen deficient below this depth. Therefore, oligotrophic lakes often support <a href="/wiki/Fish" title="Fish">fish</a> species such as <a href="/wiki/Lake_trout" title="Lake trout">lake trout</a>, which require cold, well-<a href="/wiki/Oxygenation_(environmental)" class="mw-redirect" title="Oxygenation (environmental)">oxygenated</a> waters. The oxygen content of these lakes is a function of their seasonally mixed <a href="/wiki/Hypolimnion" title="Hypolimnion">hypolimnetic</a> volume. Hypolimnetic volumes that are anoxic will result in fish congregating in areas where oxygen is sufficient for their needs.<sup id="cite_ref-sciencedirect.com_7-1" class="reference"><a href="#cite_note-sciencedirect.com-7"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>Anoxia is more common in the hypolimnion during the summer when mixing does not occur.<sup id="cite_ref-Dodds,_Walter_K._Walter_Kennedy,_1958–2010_6-1" class="reference"><a href="#cite_note-Dodds,_Walter_K._Walter_Kennedy,_1958–2010-6"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> In the absence of oxygen from the epilimnion, <a href="/wiki/Decomposition" title="Decomposition">decomposition</a> can cause hypoxia in the hypolimnion.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Mesotrophic">Mesotrophic</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=5" title="Edit section: Mesotrophic"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Mesotrophic lakes are lakes with an intermediate level of productivity. These lakes are commonly clear water lakes and ponds with beds of submerged aquatic plants and medium levels of nutrients. </p><p>The term mesotrophic is also applied to terrestrial habitats. Mesotrophic soils have moderate nutrient levels. </p> <div class="mw-heading mw-heading3"><h3 id="Eutrophic_and_hypertrophic">Eutrophic and hypertrophic</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=6" title="Edit section: Eutrophic and hypertrophic"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading4"><h4 id="Eutrophic">Eutrophic</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=7" title="Edit section: Eutrophic"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Eutrophication" title="Eutrophication">Eutrophication</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:River_algae_Sichuan.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/74/River_algae_Sichuan.jpg/170px-River_algae_Sichuan.jpg" decoding="async" width="170" height="227" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/74/River_algae_Sichuan.jpg/255px-River_algae_Sichuan.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/74/River_algae_Sichuan.jpg/340px-River_algae_Sichuan.jpg 2x" data-file-width="1536" data-file-height="2048" /></a><figcaption>Algal bloom in a village river in the mountains near <a href="/wiki/Chengdu" title="Chengdu">Chengdu</a>, <a href="/wiki/Sichuan" title="Sichuan">Sichuan</a>, China</figcaption></figure> <p>A eutrophic water body, commonly a lake or pond, has high biological productivity. Due to excessive nutrients, especially nitrogen and phosphorus, these water bodies are able to support an abundance of aquatic plants. Usually, the water body will be dominated either by aquatic plants or algae. When aquatic plants dominate, the water tends to be clear. When algae dominate, the water tends to be darker. The algae engage in photosynthesis which supplies oxygen to the fish and biota which inhabit these waters. Occasionally, an excessive algal bloom will occur and can ultimately result in fish death, due to respiration by algae and bottom-living bacteria. The process of <a href="/wiki/Eutrophication" title="Eutrophication">eutrophication</a> can occur naturally and by <a href="/wiki/Human_impact_on_the_environment" title="Human impact on the environment">human impact on the environment</a>. </p><p>Eutrophic comes from the <a href="/wiki/Greek_language" title="Greek language">Greek</a> <i>eutrophos</i> meaning "well-nourished", from <i>eu</i> meaning good and <i>trephein</i> meaning "to nourish".<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Hypertrophic">Hypertrophic</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=8" title="Edit section: Hypertrophic"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hypertrophic or hypereutrophic lakes are very nutrient-rich lakes characterized by frequent and severe nuisance <a href="/wiki/Algal_blooms" class="mw-redirect" title="Algal blooms">algal blooms</a> and low transparency. Hypereutrophic lakes have a visibility depth of less than 3 feet (90 cm), they have greater than 40 micrograms/litre total <a href="/wiki/Chlorophyll" title="Chlorophyll">chlorophyll</a> and greater than 100 micrograms/litre <a href="/wiki/Phosphorus" title="Phosphorus">phosphorus</a>. </p><p>The excessive algal blooms can also significantly reduce oxygen levels and prevent life from functioning at lower depths creating dead zones beneath the surface. </p><p>Likewise, large algal blooms can cause <a href="/wiki/Biodilution" title="Biodilution">biodilution</a> to occur, which is a decrease in the concentration of a pollutant with an increase in <a href="/wiki/Trophic_level" title="Trophic level">trophic level</a>. This is opposed to <a href="/wiki/Biomagnification" title="Biomagnification">biomagnification</a> and is due to a decreased concentration from increased algal uptake. </p> <div class="mw-heading mw-heading2"><h2 id="Trophic_index_drivers">Trophic index drivers</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=9" title="Edit section: Trophic index drivers"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Both natural and <a href="/wiki/Human_impact_on_the_environment" title="Human impact on the environment">anthropogenic</a> factors can influence a lake or other water body's trophic index. A water body situated in a nutrient-rich region with high <a href="/wiki/Net_primary_productivity" class="mw-redirect" title="Net primary productivity">net primary productivity</a> may be naturally eutrophic. Nutrients carried into water bodies from <a href="/wiki/Non-point_sources" class="mw-redirect" title="Non-point sources">non-point sources</a> such as agricultural runoff, residential fertilisers, and sewage will all increase the algal <a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">biomass</a>, and can easily cause an oligotrophic lake to become hypereutrophic.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Freshwater_lakes">Freshwater lakes</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=10" title="Edit section: Freshwater lakes"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Although there is no absolute consensus as to which nutrients contribute the most to increasing primary productivity, phosphorus concentration is thought to be the main limiting factor in freshwater lakes.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:0_17-0" class="reference"><a href="#cite_note-:0-17"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> This is likely due to the prevalence of nitrogen-fixing microorganisms in these systems, which can compensate for a lack of readily available fixed nitrogen.<sup id="cite_ref-:0_17-1" class="reference"><a href="#cite_note-:0-17"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Marine_ecosystems">Marine ecosystems</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=11" title="Edit section: Marine ecosystems"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In some coastal marine ecosystems, research has found nitrogen to be the key limiting nutrient, driving primary production independently of phosphorus.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> Nitrogen fixation cannot adequately supply these marine ecosystems, because the nitrogen fixing microbes are themselves limited by the availability of various <a href="/wiki/Abiotic_component" title="Abiotic component">abiotic factors</a> like sunlight and dissolved oxygen.<sup id="cite_ref-:1_20-0" class="reference"><a href="#cite_note-:1-20"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> However, marine ecosystems are too broad a range of environments for one nutrient to limit all marine primary productivity. The limiting nutrient may vary in different marine environments according to a variety of factors like depth, distance from shore, or availability of organic matter.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:1_20-1" class="reference"><a href="#cite_note-:1-20"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Management_targets">Management targets</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=12" title="Edit section: Management targets"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Often, the desired trophic index differs between stakeholders. Water-fowl enthusiasts (e.g. duck hunters) may want a lake to be eutrophic so that it will support a large population of waterfowl. Residents, though, may want the same lake to be oligotrophic, as this is more pleasant for swimming and boating. Natural resource agencies are generally responsible for reconciling these conflicting uses and determining what a water body's trophic index should be. </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=13" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/40px-Wiktionary-logo-en-v2.svg.png" decoding="async" width="40" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/60px-Wiktionary-logo-en-v2.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/80px-Wiktionary-logo-en-v2.svg.png 2x" data-file-width="512" data-file-height="512" /></span></span></div> <div class="side-box-text plainlist">Look up <i><b><a href="https://en.wiktionary.org/wiki/oligotrophic" class="extiw" title="wiktionary:oligotrophic">oligotrophic</a></b></i>, <i><b><a href="https://en.wiktionary.org/wiki/mesotrophic" class="extiw" title="wiktionary:mesotrophic">mesotrophic</a></b></i>, or <i><b><a href="https://en.wiktionary.org/wiki/hypertrophic" class="extiw" title="wiktionary:hypertrophic">hypertrophic</a></b></i> in Wiktionary, the free dictionary.</div></div> </div> <ul><li><a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">Biomass (ecology)</a></li> <li><a href="/wiki/Eutrophication" title="Eutrophication">Eutrophication</a></li> <li><a href="/wiki/Nonpoint_source_pollution" title="Nonpoint source pollution">Nonpoint source pollution</a></li> <li><a href="/wiki/Secchi_disk" title="Secchi disk">Secchi disk</a></li> <li><a href="/wiki/Surface_runoff" title="Surface runoff">Surface runoff</a></li> <li><a href="/wiki/Trophic_level" title="Trophic level">Trophic level</a></li> <li><a href="/wiki/Trophic_level_index" title="Trophic level index">Trophic level index</a>, a similar measure used in New Zealand</li> <li><a href="/wiki/Water_quality" title="Water quality">Water quality</a></li> <li><a href="/wiki/List_of_biological_development_disorders" title="List of biological development disorders">List of biological development disorders</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="Notes">Notes</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=14" title="Edit section: Notes"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-lower-alpha"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text">Note that this use of trophic levels refers to feeding dynamics, and has a much different meaning than the <i>trophic index</i> of water bodies.</span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_state_index&action=edit&section=15" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239543626"><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-USFWaterAtlas-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-USFWaterAtlas_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-USFWaterAtlas_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-USFWaterAtlas_1-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free 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.navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Aquatic_ecosystems" title="Template:Aquatic ecosystems"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Aquatic_ecosystems" title="Template talk:Aquatic ecosystems"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Aquatic_ecosystems" title="Special:EditPage/Template:Aquatic ecosystems"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Aquatic_ecosystems" style="font-size:114%;margin:0 4em"><a href="/wiki/Aquatic_ecosystem" title="Aquatic ecosystem">Aquatic ecosystems</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="General_components_and_freshwater_ecosystems" style="font-size:114%;margin:0 4em">General components and <a href="/wiki/Freshwater_ecosystem" title="Freshwater ecosystem">freshwater ecosystems</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acoustic_ecology" title="Acoustic ecology">Acoustic ecology</a></li> <li><a href="/wiki/Algal_bloom" title="Algal bloom">Algal bloom</a></li> <li><a href="/wiki/Anoxic_waters" title="Anoxic waters">Anoxic waters</a></li> <li><a href="/wiki/Aquatic_adaptation" class="mw-redirect" title="Aquatic adaptation">Aquatic adaptation</a></li> <li><a href="/wiki/Aquatic_animal" title="Aquatic animal">Aquatic animal</a> <ul><li><a href="/wiki/Aquatic_insect" title="Aquatic insect">Insect</a></li> <li><a href="/wiki/Aquatic_mammal" title="Aquatic mammal">Mammal</a></li> <li><a href="/wiki/Water_bird" title="Water bird">Water bird</a></li></ul></li> <li><a href="/wiki/Aquatic_biomonitoring" title="Aquatic biomonitoring">Aquatic biomonitoring</a></li> <li><a href="/wiki/Aquatic_plant" title="Aquatic plant">Aquatic plant</a></li> <li><a href="/wiki/Agent-based_model_in_biology#Aquatic_population_dynamics" title="Agent-based model in biology">Aquatic population dynamics</a></li> <li><a href="/wiki/Aquatic_predation" class="mw-redirect" title="Aquatic predation">Aquatic predation</a></li> <li><a href="/wiki/Aquatic_respiration" title="Aquatic respiration">Aquatic respiration</a></li> <li><a href="/wiki/Aquatic_science" title="Aquatic science">Aquatic science</a></li> <li><a href="/wiki/Aquatic_toxicology" title="Aquatic toxicology">Aquatic toxicology</a></li> <li><a href="/wiki/Benthos" title="Benthos">Benthos</a></li> <li><a href="/wiki/Bioluminescence" title="Bioluminescence">Bioluminescence</a></li> <li><a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">Biomass</a></li> <li><a href="/wiki/Cascade_effect_(ecology)" title="Cascade effect (ecology)">Cascade effect</a></li> <li><a href="/wiki/Colored_dissolved_organic_matter" title="Colored dissolved organic matter">Colored dissolved organic matter</a></li> <li><a href="/wiki/Dead_zone_(ecology)" title="Dead zone (ecology)">Dead zone</a></li> <li><a href="/wiki/Ecohydrology" title="Ecohydrology">Ecohydrology</a></li> <li><a href="/wiki/Eutrophication" title="Eutrophication">Eutrophication</a></li> <li><a href="/wiki/Fisheries_science" title="Fisheries science">Fisheries science</a></li> <li><a href="/wiki/Food_chain" title="Food chain">Food chain</a></li> <li><a href="/wiki/Food_web" title="Food web">Food web</a></li> <li><a href="/wiki/GIS_and_aquatic_science" title="GIS and aquatic science">GIS and aquatic science</a></li> <li><a href="/wiki/Hydrobiology" title="Hydrobiology">Hydrobiology</a></li> <li><a href="/wiki/Hypoxia_(environmental)" title="Hypoxia (environmental)">Hypoxia</a></li> <li><a href="/wiki/Macrobenthos" title="Macrobenthos">Macrobenthos</a></li> <li><a href="/wiki/Meiobenthos" title="Meiobenthos">Meiobenthos</a></li> <li><a href="/wiki/Microbial_ecology" title="Microbial ecology">Microbial ecology</a></li> <li><a href="/wiki/Microbial_food_web" title="Microbial food web">Microbial food web</a></li> <li><a href="/wiki/Microbial_loop" title="Microbial loop">Microbial loop</a></li> <li><a href="/wiki/Nekton" title="Nekton">Nekton</a></li> <li><a href="/wiki/Neuston" title="Neuston">Neuston</a></li> <li><a href="/wiki/Particle_(ecology)" title="Particle (ecology)">Particle</a></li> <li><a href="/wiki/Pelagic_zone" title="Pelagic zone">Pelagic zone</a></li> <li><a href="/wiki/Photic_zone" title="Photic zone">Photic zone</a></li> <li><a href="/wiki/Phytoplankton" title="Phytoplankton">Phytoplankton</a></li> <li><a href="/wiki/Plankton" title="Plankton">Plankton</a></li> <li><a href="/wiki/Pleuston" class="mw-redirect" title="Pleuston">Pleuston</a></li> <li><a href="/wiki/Productivity_(ecology)" title="Productivity (ecology)">Productivity</a></li> <li><a href="/wiki/Ramsar_Convention" title="Ramsar Convention">Ramsar Convention</a></li> <li><a href="/wiki/Sediment_trap" title="Sediment trap">Sediment trap</a></li> <li><a href="/wiki/Shoaling_and_schooling" title="Shoaling and schooling">Shoaling and schooling</a></li> <li><a href="/wiki/Siltation" title="Siltation">Siltation</a></li> <li><a href="/wiki/Spawn_(biology)" class="mw-redirect" title="Spawn (biology)">Spawn</a></li> <li><a href="/wiki/Isotope_analysis#Stable_isotope_analysis_in_aquatic_ecosystems" title="Isotope analysis">Stable isotope analysis in aquatic ecosystems</a></li> <li><a href="/wiki/Substrate_(marine_biology)" class="mw-redirect" title="Substrate (marine biology)">Substrate</a></li> <li><a href="/wiki/Thermal_pollution" title="Thermal pollution">Thermal pollution</a></li> <li><a href="/wiki/Trophic_level" title="Trophic level">Trophic level</a></li> <li><a href="/wiki/Underwater_camouflage" title="Underwater camouflage">Underwater camouflage</a></li> <li><a href="/wiki/Water_column" title="Water column">Water column</a></li> <li><a href="/wiki/Zooplankton" title="Zooplankton">Zooplankton</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="3" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg/140px-Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg" decoding="async" width="140" height="202" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg/210px-Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg/280px-Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg 2x" data-file-width="382" data-file-height="551" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Freshwater</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Freshwater_biology" title="Freshwater biology">Freshwater biology</a></li> <li><a href="/wiki/Freshwater_biome" class="mw-redirect" title="Freshwater biome">Freshwater biome</a></li> <li><a href="/wiki/Freshwater_environmental_quality_parameters" title="Freshwater environmental quality parameters">Freshwater environmental quality parameters</a></li> <li><a href="/wiki/Freshwater_fish" title="Freshwater fish">Freshwater fish</a></li> <li><a href="/wiki/Hyporheic_zone" title="Hyporheic zone">Hyporheic zone</a></li> <li><a href="/wiki/Limnology" title="Limnology">Limnology</a></li> <li><a href="/wiki/Lake_ecosystem" title="Lake ecosystem">Lake ecosystem</a></li> <li><a href="/wiki/Lake_stratification" title="Lake stratification">Lake stratification</a></li> <li><a href="/wiki/Macrophyte" class="mw-redirect" title="Macrophyte">Macrophyte</a></li> <li><a href="/wiki/Pond" title="Pond">Pond</a> <ul><li><a href="/wiki/Fish_pond" title="Fish pond">Fish pond</a></li></ul></li> <li><a href="/wiki/Rheotaxis" title="Rheotaxis">Rheotaxis</a></li> <li><a href="/wiki/River_ecosystem" title="River ecosystem">River ecosystem</a></li> <li><a href="/wiki/Stream_bed" title="Stream bed">Stream bed</a></li> <li><a href="/wiki/Stream_pool" title="Stream pool">Stream pool</a></li> <li><a class="mw-selflink selflink">Trophic state index</a></li> <li><a href="/wiki/Upland_and_lowland" title="Upland and lowland">Upland and lowland</a></li> <li><a href="/wiki/Water_garden" title="Water garden">Water garden</a></li> <li><a href="/wiki/Wetland" title="Wetland">Wetland</a> <ul><li><a href="/wiki/Bog" title="Bog">Bog</a></li> <li><a href="/wiki/Brackish_marsh" title="Brackish marsh">Brackish marsh</a></li> <li><a href="/wiki/Fen" title="Fen">Fen</a></li> <li><a href="/wiki/Freshwater_marsh" title="Freshwater marsh">Freshwater marsh</a></li> <li><a href="/wiki/Freshwater_swamp_forest" title="Freshwater swamp forest">Freshwater swamp forest</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Ecoregions</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/List_of_freshwater_ecoregions_(WWF)" title="List of freshwater ecoregions (WWF)">List of freshwater ecoregions (WWF)</a> <ul><li><a href="/wiki/List_of_freshwater_ecoregions_in_Africa_and_Madagascar" title="List of freshwater ecoregions in Africa and Madagascar">Africa and Madagascar</a></li> <li><a href="/wiki/List_of_freshwater_ecoregions_of_Latin_America_and_the_Caribbean" title="List of freshwater ecoregions of Latin America and the Caribbean">Latin America and the Caribbean</a></li></ul></li> <li><a href="/wiki/List_of_marine_ecoregions" title="List of marine ecoregions">List of marine ecoregions</a></li> <li>Specific examples <ul><li><a href="/wiki/Geography_and_ecology_of_the_Everglades" title="Geography and ecology of the Everglades">Everglades</a></li> <li><a href="/wiki/Freshwater_ecology_of_Maharashtra" title="Freshwater ecology of Maharashtra">Maharashtra</a></li> <li><a href="/wiki/Ecosystem_of_the_North_Pacific_Subtropical_Gyre" title="Ecosystem of the North Pacific Subtropical Gyre">North Pacific Subtropical Gyre</a></li> <li><a href="/wiki/Ecology_of_the_San_Francisco_Estuary" title="Ecology of the San Francisco Estuary">San Francisco Estuary</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Marine_ecosystems" style="font-size:114%;margin:0 4em"><a href="/wiki/Marine_ecosystem" title="Marine ecosystem">Marine ecosystems</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Deep_scattering_layer" title="Deep scattering layer">Deep scattering layer</a></li> <li><a href="/wiki/Diel_vertical_migration" title="Diel vertical migration">Diel vertical migration</a></li> <li><a href="/wiki/F-ratio_(oceanography)" title="F-ratio (oceanography)">f-ratio</a></li> <li><a href="/wiki/Iron_fertilization" title="Iron fertilization">Iron fertilization</a></li> <li><a href="/wiki/Large_marine_ecosystem" class="mw-redirect" title="Large marine ecosystem">Large marine ecosystem</a></li> <li><a href="/wiki/Marine_biology" title="Marine biology">Marine biology</a></li> <li><a href="/wiki/Marine_chemistry" title="Marine chemistry">Marine chemistry</a></li> <li><a href="/wiki/Marine_food_web" title="Marine food web">Marine food web</a></li> <li><a href="/wiki/Marine_primary_production" title="Marine primary production">Marine primary production</a></li> <li><a href="/wiki/Marine_snow" title="Marine snow">Marine snow</a></li> <li><a href="/wiki/Ocean_fertilization" title="Ocean fertilization">Ocean fertilization</a></li> <li><a href="/wiki/Oceanic_physical-biological_process" title="Oceanic physical-biological process">Oceanic physical-biological process</a></li> <li><a href="/wiki/Ocean_turbidity" title="Ocean turbidity">Ocean turbidity</a></li> <li><a href="/wiki/Photophore" title="Photophore">Photophore</a></li> <li><a href="/wiki/Thorson%27s_rule" title="Thorson's rule">Thorson's rule</a></li> <li><a href="/wiki/Upwelling" title="Upwelling">Upwelling</a></li> <li><a href="/wiki/Viral_shunt" title="Viral shunt">Viral shunt</a></li> <li><a href="/wiki/Whale_fall" title="Whale fall">Whale fall</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="4" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Maldivesfish2.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/35/Maldivesfish2.jpg/140px-Maldivesfish2.jpg" decoding="async" width="140" height="95" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/35/Maldivesfish2.jpg/210px-Maldivesfish2.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/35/Maldivesfish2.jpg/280px-Maldivesfish2.jpg 2x" data-file-width="1818" data-file-height="1228" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_life" title="Marine life">Marine life</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Census_of_Marine_Life" title="Census of Marine Life">Census of Marine Life</a></li> <li><a href="/wiki/Deep-sea_community" title="Deep-sea community">Deep-sea community</a></li> <li><a href="/wiki/Deep-water_coral" title="Deep-water coral">Deep-water coral</a></li> <li><a href="/wiki/Marine_fungi" title="Marine fungi">Marine fungi</a></li> <li><a href="/wiki/Marine_invertebrates" title="Marine invertebrates">Marine invertebrates</a></li> <li><a href="/wiki/Marine_larval_ecology" title="Marine larval ecology">Marine larval ecology</a></li> <li><a href="/wiki/Seagrass" title="Seagrass">Seagrass</a></li> <li><a href="/wiki/Seashore_wildlife" title="Seashore wildlife">Seashore wildlife</a></li> <li><a href="/wiki/Wild_fisheries" title="Wild fisheries">Wild fisheries</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_microorganisms" title="Marine microorganisms">Microorganisms</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Marine_bacteriophage" class="mw-redirect" title="Marine bacteriophage">Marine bacteriophage</a></li> <li><a href="/wiki/Marine_prokaryotes" title="Marine prokaryotes">Marine prokaryotes</a></li> <li><a href="/wiki/Marine_protists" title="Marine protists">Marine protists</a></li> <li><a href="/wiki/Marine_viruses" title="Marine viruses">Marine viruses</a></li> <li><a href="/wiki/Paradox_of_the_plankton" title="Paradox of the plankton">Paradox of the plankton</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_vertebrate" title="Marine vertebrate">Vertebrates</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Marine_mammal" title="Marine mammal">Marine mammal</a></li> <li><a href="/wiki/Marine_reptile" title="Marine reptile">Marine reptile</a></li> <li><a href="/wiki/Saltwater_fish" title="Saltwater fish">Saltwater fish</a> <ul><li><a href="/wiki/Coastal_fish" title="Coastal fish">Coastal fish</a></li> <li><a href="/wiki/Coral_reef_fish" title="Coral reef fish">Coral reef fish</a></li> <li><a href="/wiki/Deep-sea_fish" title="Deep-sea fish">Deep-sea fish</a></li> <li><a href="/wiki/Demersal_fish" title="Demersal fish">Demersal fish</a></li> <li><a href="/wiki/Pelagic_fish" title="Pelagic fish">Pelagic fish</a></li></ul></li> <li><a href="/wiki/Seabird" title="Seabird">Seabird</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_habitat" title="Marine habitat">Marine habitats</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bay_mud" title="Bay mud">Bay mud</a></li> <li><a href="/wiki/Marine_coastal_ecosystem" title="Marine coastal ecosystem">Marine coastal ecosystem</a></li> <li><a href="/wiki/Coastal_biogeomorphology" title="Coastal biogeomorphology">Coastal biogeomorphology</a></li> <li><a href="/wiki/Cold_seep" title="Cold seep">Cold seep</a></li> <li><a href="/wiki/Coral_reef" title="Coral reef">Coral reef</a></li> <li><a href="/wiki/Davidson_Seamount#Ecology" title="Davidson Seamount">Davidson Seamount § Ecology</a></li> <li><a href="/wiki/Estuary" title="Estuary">Estuary</a></li> <li><a href="/wiki/Intertidal_ecology" title="Intertidal ecology">Intertidal ecology</a></li> <li><a href="/wiki/Intertidal_wetland" title="Intertidal wetland">Intertidal wetland</a></li> <li><a href="/wiki/Kelp_forest" title="Kelp forest">Kelp forest</a></li> <li><a href="/wiki/Hydrothermal_vent" title="Hydrothermal vent">Hydrothermal vent</a></li> <li><a href="/wiki/Lagoon" title="Lagoon">Lagoon</a></li> <li><a href="/wiki/Mangrove" title="Mangrove">Mangrove</a></li> <li><a href="/wiki/Marine_biomes" class="mw-redirect" title="Marine biomes">Marine biomes</a></li> <li><a href="/wiki/Mudflat" title="Mudflat">Mudflat</a></li> <li><a href="/wiki/Oyster_reef" title="Oyster reef">Oyster reef</a></li> <li><a href="/wiki/Rocky_shore" title="Rocky shore">Rocky shore</a></li> <li><a href="/wiki/Salt_marsh" title="Salt marsh">Salt marsh</a></li> <li><a href="/wiki/Salt_pannes_and_pools" title="Salt pannes and pools">Salt pannes and pools</a></li> <li><a href="/wiki/Seagrass_meadow" title="Seagrass meadow">Seagrass meadow</a></li> <li><a href="/wiki/Sponge_ground" title="Sponge ground">Sponge ground</a></li> <li><a href="/wiki/Sponge_reef" title="Sponge reef">Sponge reef</a></li> <li><a href="/wiki/Tide_pool" title="Tide pool">Tide pool</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_conservation" title="Marine conservation">Conservation</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Coral_bleaching" title="Coral bleaching">Coral bleaching</a></li> <li><a href="/wiki/Ecological_values_of_mangroves" title="Ecological values of mangroves">Ecological values of mangroves</a></li> <li><a href="/wiki/Fisheries_and_climate_change" class="mw-redirect" title="Fisheries and climate change">Fisheries and climate change</a></li> <li><a href="/wiki/Hotspot_Ecosystem_Research_and_Man%27s_Impact_On_European_Seas" title="Hotspot Ecosystem Research and Man's Impact On European Seas">HERMIONE</a></li> <li><a href="/wiki/Human_impact_on_marine_life" title="Human impact on marine life">Human impact on marine life</a></li> <li><a href="/wiki/Marine_conservation_activism" title="Marine conservation activism">Marine conservation activism</a></li> <li><a href="/wiki/Marine_pollution" title="Marine pollution">Marine pollution</a></li> <li><a href="/wiki/Marine_protected_area" title="Marine protected area">Marine protected area</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><span class="nowrap"><span class="nowrap"><span class="noviewer" typeof="mw:File"><span><img alt="image" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png/16px-Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png/24px-Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png/32px-Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png 2x" data-file-width="1500" data-file-height="1500" /></span></span> </span><a href="/wiki/Portal:Lakes" title="Portal:Lakes">Lakes portal</a></span></li> <li><span class="nowrap"><span class="nowrap"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Waves_in_pacifica_1.jpg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/Waves_in_pacifica_1.jpg/16px-Waves_in_pacifica_1.jpg" decoding="async" width="16" height="11" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/Waves_in_pacifica_1.jpg/24px-Waves_in_pacifica_1.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/45/Waves_in_pacifica_1.jpg/32px-Waves_in_pacifica_1.jpg 2x" data-file-width="2000" data-file-height="1358" /></a></span> </span><a href="/wiki/Portal:Oceans" title="Portal:Oceans">Oceans portal</a></span></li> <li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Aquatic_ecology" title="Category:Aquatic ecology">Category</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Ecology:_Modelling_ecosystems:_Trophic_components" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Modelling_ecosystems" title="Template:Modelling ecosystems"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Modelling_ecosystems" title="Template talk:Modelling ecosystems"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Modelling_ecosystems" title="Special:EditPage/Template:Modelling ecosystems"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Ecology:_Modelling_ecosystems:_Trophic_components" style="font-size:114%;margin:0 4em"><a href="/wiki/Ecology" title="Ecology">Ecology</a>: <a href="/wiki/Ecosystem_model" title="Ecosystem model">Modelling ecosystems</a>: <a href="/wiki/Trophic_level" title="Trophic level">Trophic</a> components</div></th></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abiotic_component" title="Abiotic component">Abiotic component</a></li> <li><a href="/wiki/Abiotic_stress" title="Abiotic stress">Abiotic stress</a></li> <li><a href="/wiki/Behavioral_ecology" title="Behavioral ecology">Behaviour</a></li> <li><a href="/wiki/Biogeochemical_cycle" title="Biogeochemical cycle">Biogeochemical cycle</a></li> <li><a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">Biomass</a></li> <li><a href="/wiki/Biotic_component" class="mw-redirect" title="Biotic component">Biotic component</a></li> <li><a href="/wiki/Biotic_stress" title="Biotic stress">Biotic stress</a></li> <li><a href="/wiki/Carrying_capacity" title="Carrying capacity">Carrying capacity</a></li> <li><a href="/wiki/Competition_(biology)" title="Competition (biology)">Competition</a></li> <li><a href="/wiki/Ecosystem" title="Ecosystem">Ecosystem</a></li> <li><a href="/wiki/Ecosystem_ecology" title="Ecosystem ecology">Ecosystem ecology</a></li> <li><a href="/wiki/Ecosystem_model" title="Ecosystem model">Ecosystem model</a></li> <li><a href="/wiki/Green_world_hypothesis" title="Green world hypothesis">Green world hypothesis</a></li> <li><a href="/wiki/Keystone_species" title="Keystone species">Keystone species</a></li> <li><a href="/wiki/List_of_feeding_behaviours" title="List of feeding behaviours">List of feeding behaviours</a></li> <li><a href="/wiki/Metabolic_theory_of_ecology" title="Metabolic theory of ecology">Metabolic theory of ecology</a></li> <li><a href="/wiki/Productivity_(ecology)" title="Productivity (ecology)">Productivity</a></li> <li><a href="/wiki/Resource_(biology)" title="Resource (biology)">Resource</a></li> <li><a href="/wiki/Restoration_ecology" class="mw-redirect" title="Restoration ecology">Restoration</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Autotroph" title="Autotroph">Producers</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Autotroph" title="Autotroph">Autotrophs</a></li> <li><a href="/wiki/Chemosynthesis" title="Chemosynthesis">Chemosynthesis</a></li> <li><a href="/wiki/Chemotroph" title="Chemotroph">Chemotrophs</a></li> <li><a href="/wiki/Foundation_species" title="Foundation species">Foundation species</a></li> <li><a href="/wiki/Kinetotroph" title="Kinetotroph">Kinetotrophs</a></li> <li><a href="/wiki/Mixotroph" title="Mixotroph">Mixotrophs</a></li> <li><a href="/wiki/Myco-heterotrophy" title="Myco-heterotrophy">Myco-heterotrophy</a></li> <li><a href="/wiki/Mycotroph" title="Mycotroph">Mycotroph</a></li> <li><a href="/wiki/Organotroph" title="Organotroph">Organotrophs</a></li> <li><a href="/wiki/Photoheterotroph" title="Photoheterotroph">Photoheterotrophs</a></li> <li><a href="/wiki/Photosynthesis" title="Photosynthesis">Photosynthesis</a></li> <li><a href="/wiki/Photosynthetic_efficiency" title="Photosynthetic efficiency">Photosynthetic efficiency</a></li> <li><a href="/wiki/Phototroph" title="Phototroph">Phototrophs</a></li> <li><a href="/wiki/Primary_nutritional_groups" title="Primary nutritional groups">Primary nutritional groups</a></li> <li><a href="/wiki/Primary_production" title="Primary production">Primary production</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Consumer_(food_chain)" title="Consumer (food chain)">Consumers</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Apex_predator" title="Apex predator">Apex predator</a></li> <li><a href="/wiki/Bacterivore" title="Bacterivore">Bacterivore</a></li> <li><a href="/wiki/Carnivore" title="Carnivore">Carnivores</a></li> <li><a href="/wiki/Chemoorganotroph" class="mw-redirect" title="Chemoorganotroph">Chemoorganotroph</a></li> <li><a href="/wiki/Foraging" title="Foraging">Foraging</a></li> <li><a href="/wiki/Generalist_and_specialist_species" title="Generalist and specialist species">Generalist and specialist species</a></li> <li><a href="/wiki/Intraguild_predation" title="Intraguild predation">Intraguild predation</a></li> <li><a href="/wiki/Herbivore" title="Herbivore">Herbivores</a></li> <li><a href="/wiki/Heterotroph" title="Heterotroph">Heterotroph</a></li> <li><a href="/wiki/Heterotrophic_nutrition" title="Heterotrophic nutrition">Heterotrophic nutrition</a></li> <li><a href="/wiki/Insectivore" title="Insectivore">Insectivore</a></li> <li><a href="/wiki/Mesopredator" title="Mesopredator">Mesopredators</a></li> <li><a href="/wiki/Mesopredator_release_hypothesis" title="Mesopredator release hypothesis">Mesopredator release hypothesis</a></li> <li><a href="/wiki/Omnivore" title="Omnivore">Omnivores</a></li> <li><a href="/wiki/Optimal_foraging_theory" title="Optimal foraging theory">Optimal foraging theory</a></li> <li><a href="/wiki/Planktivore" title="Planktivore">Planktivore</a></li> <li><a href="/wiki/Predation" title="Predation">Predation</a></li> <li><a href="/wiki/Prey_switching" title="Prey switching">Prey switching</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Decomposer" title="Decomposer">Decomposers</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Chemoorganoheterotrophy" class="mw-redirect" title="Chemoorganoheterotrophy">Chemoorganoheterotrophy</a></li> <li><a href="/wiki/Decomposition" title="Decomposition">Decomposition</a></li> <li><a href="/wiki/Detritivore" title="Detritivore">Detritivores</a></li> <li><a href="/wiki/Detritus" title="Detritus">Detritus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Microorganism#Habitats_and_ecology" title="Microorganism">Microorganisms</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archaea" title="Archaea">Archaea</a></li> <li><a href="/wiki/Bacteriophage" title="Bacteriophage">Bacteriophage</a></li> <li><a href="/wiki/Lithoautotroph" title="Lithoautotroph">Lithoautotroph</a></li> <li><a href="/wiki/Lithotroph" title="Lithotroph">Lithotrophy</a></li> <li><a href="/wiki/Marine_microorganisms" title="Marine microorganisms">Marine</a></li> <li><a href="/wiki/Microbial_cooperation" title="Microbial cooperation">Microbial cooperation</a></li> <li><a href="/wiki/Microbial_ecology" title="Microbial ecology">Microbial ecology</a></li> <li><a href="/wiki/Microbial_food_web" title="Microbial food web">Microbial food web</a></li> <li><a href="/wiki/Microbial_intelligence" title="Microbial intelligence">Microbial intelligence</a></li> <li><a href="/wiki/Microbial_loop" title="Microbial loop">Microbial loop</a></li> <li><a href="/wiki/Microbial_mat" title="Microbial mat">Microbial mat</a></li> <li><a href="/wiki/Microbial_metabolism" title="Microbial metabolism">Microbial metabolism</a></li> <li><a href="/wiki/Phage_ecology" title="Phage ecology">Phage ecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Food_web" title="Food web">Food webs</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biomagnification" title="Biomagnification">Biomagnification</a></li> <li><a href="/wiki/Ecological_efficiency" title="Ecological efficiency">Ecological efficiency</a></li> <li><a href="/wiki/Ecological_pyramid" title="Ecological pyramid">Ecological pyramid</a></li> <li><a href="/wiki/Energy_flow_(ecology)" title="Energy flow (ecology)">Energy flow</a></li> <li><a href="/wiki/Food_chain" title="Food chain">Food chain</a></li> <li><a href="/wiki/Trophic_level" title="Trophic level">Trophic level</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Example webs</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lake_ecosystem#Trophic_relationships" title="Lake ecosystem">Lakes</a></li> <li><a href="/wiki/River_ecosystem#Trophic_relationships" title="River ecosystem">Rivers</a></li> <li><a href="/wiki/Soil_food_web" title="Soil food web">Soil</a></li> <li><a href="/wiki/Tritrophic_interactions_in_plant_defense" title="Tritrophic interactions in plant defense">Tritrophic interactions in plant defense</a></li> <li><a href="/wiki/Marine_food_web" title="Marine food web">Marine food webs</a> <ul><li><a href="/wiki/Cold_seep" title="Cold seep">cold seeps</a></li> <li><a href="/wiki/Hydrothermal_vent#Biological_communities" title="Hydrothermal vent">hydrothermal vents</a></li> <li><a href="/wiki/Intertidal_ecology" title="Intertidal ecology">intertidal</a></li> <li><a href="/wiki/Kelp_forest#Trophic_ecology" title="Kelp forest">kelp forests</a></li> <li><a href="/wiki/North_Pacific_Gyre" title="North Pacific Gyre">North Pacific Gyre</a></li> <li><a href="/wiki/Ecology_of_the_San_Francisco_Estuary#Food_web" title="Ecology of the San Francisco Estuary">San Francisco Estuary</a></li> <li><a href="/wiki/Tide_pool" title="Tide pool">tide pool</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Processes</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ascendency" title="Ascendency">Ascendency</a></li> <li><a href="/wiki/Bioaccumulation" title="Bioaccumulation">Bioaccumulation</a></li> <li><a href="/wiki/Cascade_effect_(ecology)" title="Cascade effect (ecology)">Cascade effect</a></li> <li><a href="/wiki/Climax_community" title="Climax community">Climax community</a></li> <li><a href="/wiki/Competitive_exclusion_principle" title="Competitive exclusion principle">Competitive exclusion principle</a></li> <li><a href="/wiki/Consumer%E2%80%93resource_interactions" title="Consumer–resource interactions">Consumer–resource interactions</a></li> <li><a href="/wiki/Copiotroph" title="Copiotroph">Copiotrophs</a></li> <li><a href="/wiki/Dominance_(ecology)" title="Dominance (ecology)">Dominance</a></li> <li><a href="/wiki/Ecological_network" title="Ecological network">Ecological network</a></li> <li><a href="/wiki/Ecological_succession" title="Ecological succession">Ecological succession</a></li> <li><a href="/wiki/Energy_quality" title="Energy quality">Energy quality</a></li> <li><a href="/wiki/Energy_systems_language" title="Energy systems language">Energy systems language</a></li> <li><a href="/wiki/F-ratio_(oceanography)" title="F-ratio (oceanography)">f-ratio</a></li> <li><a href="/wiki/Feed_conversion_ratio" title="Feed conversion ratio">Feed conversion ratio</a></li> <li><a href="/wiki/Feeding_frenzy" title="Feeding frenzy">Feeding frenzy</a></li> <li><a href="/wiki/Mesotrophic_soil" title="Mesotrophic soil">Mesotrophic soil</a></li> <li><a href="/wiki/Nutrient_cycle" title="Nutrient cycle">Nutrient cycle</a></li> <li><a href="/wiki/Oligotroph" title="Oligotroph">Oligotroph</a></li> <li><a href="/wiki/Paradox_of_the_plankton" title="Paradox of the plankton">Paradox of the plankton</a></li> <li><a href="/wiki/Trophic_cascade" title="Trophic cascade">Trophic cascade</a></li> <li><a href="/wiki/Trophic_mutualism" title="Trophic mutualism">Trophic mutualism</a></li> <li><a class="mw-selflink selflink">Trophic state index</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Defense,<br />counter</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Animal_coloration" title="Animal coloration">Animal coloration</a></li> <li><a href="/wiki/Anti-predator_adaptation" title="Anti-predator adaptation">Anti-predator adaptations</a></li> <li><a href="/wiki/Camouflage" title="Camouflage">Camouflage</a></li> <li><a href="/wiki/Deimatic_behaviour" title="Deimatic behaviour">Deimatic behaviour</a></li> <li><a href="/wiki/Herbivore_adaptations_to_plant_defense" title="Herbivore adaptations to plant defense">Herbivore adaptations to plant defense</a></li> <li><a href="/wiki/Mimicry" title="Mimicry">Mimicry</a></li> <li><a href="/wiki/Plant_defense_against_herbivory" title="Plant defense against herbivory">Plant defense against herbivory</a></li> <li><a href="/wiki/Shoaling_and_schooling#Predator_avoidance" title="Shoaling and schooling">Predator avoidance in schooling fish</a></li></ul> </div></td></tr></tbody></table></div><div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Ecology:_Modelling_ecosystems:_Other_components" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" 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ecology">Population<br />ecology</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abundance_(ecology)" title="Abundance (ecology)">Abundance</a></li> <li><a href="/wiki/Allee_effect" title="Allee effect">Allee effect</a></li> <li><a href="/wiki/Consumer-resource_model" title="Consumer-resource model">Consumer-resource model</a></li> <li><a href="/wiki/Depensation" title="Depensation">Depensation</a></li> <li><a href="/wiki/Ecological_yield" title="Ecological yield">Ecological yield</a></li> <li><a href="/wiki/Effective_population_size" title="Effective population size">Effective population size</a></li> <li><a href="/wiki/Intraspecific_competition" title="Intraspecific competition">Intraspecific competition</a></li> <li><a href="/wiki/Logistic_function" title="Logistic function">Logistic function</a></li> <li><a href="/wiki/Malthusian_growth_model" title="Malthusian growth model">Malthusian growth model</a></li> <li><a href="/wiki/Maximum_sustainable_yield" title="Maximum sustainable yield">Maximum sustainable yield</a></li> <li><a href="/wiki/Overpopulation" title="Overpopulation">Overpopulation</a></li> <li><a href="/wiki/Overexploitation" title="Overexploitation">Overexploitation</a></li> <li><a href="/wiki/Population_cycle" title="Population cycle">Population cycle</a></li> <li><a href="/wiki/Population_dynamics" title="Population dynamics">Population dynamics</a></li> <li><a href="/wiki/Population_model" title="Population model">Population modeling</a></li> <li><a href="/wiki/Population_size" title="Population size">Population size</a></li> <li><a href="/wiki/Lotka%E2%80%93Volterra_equations" title="Lotka–Volterra equations">Predator–prey (Lotka–Volterra) equations</a></li> <li><a href="/wiki/Recruitment_(biology)" title="Recruitment (biology)">Recruitment</a></li> <li><a href="/wiki/Small_population_size" title="Small population size">Small population size</a></li> <li><a href="/wiki/Ecological_stability" title="Ecological stability">Stability</a> <ul><li><a href="/wiki/Ecological_resilience" title="Ecological resilience">Resilience</a></li> <li><a href="/wiki/Resistance_(ecology)" title="Resistance (ecology)">Resistance</a></li></ul></li> <li><a href="/wiki/Random_generalized_Lotka%E2%80%93Volterra_model" title="Random generalized Lotka–Volterra model">Random generalized Lotka–Volterra model</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Species</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biodiversity" title="Biodiversity">Biodiversity</a></li> <li><a href="/wiki/Density_dependence" title="Density dependence">Density-dependent inhibition</a></li> <li><a href="/wiki/Ecological_effects_of_biodiversity" title="Ecological effects of biodiversity">Ecological effects of biodiversity</a></li> <li><a href="/wiki/Ecological_extinction" title="Ecological extinction">Ecological extinction</a></li> <li><a href="/wiki/Endemism" title="Endemism">Endemic species</a></li> <li><a href="/wiki/Flagship_species" title="Flagship species">Flagship species</a></li> <li><a href="/wiki/Ordination_(statistics)" title="Ordination (statistics)">Gradient analysis</a></li> <li><a href="/wiki/Bioindicator" title="Bioindicator">Indicator species</a></li> <li><a href="/wiki/Introduced_species" title="Introduced species">Introduced species</a></li> <li><a href="/wiki/Invasive_species" title="Invasive species">Invasive species</a> / <a href="/wiki/Native_species" title="Native species">Native species</a></li> <li><a href="/wiki/Latitudinal_gradients_in_species_diversity" title="Latitudinal gradients in species diversity">Latitudinal gradients in species diversity</a></li> <li><a href="/wiki/Minimum_viable_population" title="Minimum viable population">Minimum viable population</a></li> <li><a href="/wiki/Unified_neutral_theory_of_biodiversity" title="Unified neutral theory of biodiversity">Neutral theory</a></li> <li><a href="/wiki/Occupancy%E2%80%93abundance_relationship" title="Occupancy–abundance relationship">Occupancy–abundance relationship</a></li> <li><a href="/wiki/Population_viability_analysis" title="Population viability analysis">Population viability analysis</a></li> <li><a href="/wiki/Priority_effect" title="Priority effect">Priority effect</a></li> <li><a href="/wiki/Rapoport%27s_rule" title="Rapoport's rule">Rapoport's rule</a></li> <li><a href="/wiki/Relative_abundance_distribution" title="Relative abundance distribution">Relative abundance distribution</a></li> <li><a href="/wiki/Relative_species_abundance" title="Relative species abundance">Relative species abundance</a></li> <li><a href="/wiki/Species_diversity" title="Species diversity">Species diversity</a></li> <li><a href="/wiki/Species_homogeneity" title="Species homogeneity">Species homogeneity</a></li> <li><a href="/wiki/Species_richness" title="Species richness">Species richness</a></li> <li><a href="/wiki/Species_distribution" title="Species distribution">Species distribution</a></li> <li><a href="/wiki/Species%E2%80%93area_relationship" title="Species–area relationship">Species–area curve</a></li> <li><a href="/wiki/Umbrella_species" title="Umbrella species">Umbrella species</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Species<br />interaction</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Antibiosis" title="Antibiosis">Antibiosis</a></li> <li><a href="/wiki/Biological_interaction" title="Biological interaction">Biological interaction</a></li> <li><a href="/wiki/Commensalism" title="Commensalism">Commensalism</a></li> <li><a href="/wiki/Community_(ecology)" title="Community (ecology)">Community ecology</a></li> <li><a href="/wiki/Ecological_facilitation" title="Ecological facilitation">Ecological facilitation</a></li> <li><a href="/wiki/Interspecific_competition" title="Interspecific competition">Interspecific competition</a></li> <li><a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">Mutualism</a></li> <li><a href="/wiki/Parasitism" title="Parasitism">Parasitism</a></li> <li><a href="/wiki/Storage_effect" title="Storage effect">Storage effect</a></li> <li><a href="/wiki/Symbiosis" title="Symbiosis">Symbiosis</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Spatial_ecology" title="Spatial ecology">Spatial<br />ecology</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a href="/wiki/Cross-boundary_subsidy" title="Cross-boundary subsidy">Cross-boundary subsidy</a></li> <li><a href="/wiki/Cline_(biology)" title="Cline (biology)">Ecocline</a></li> <li><a href="/wiki/Ecotone" title="Ecotone">Ecotone</a></li> <li><a href="/wiki/Ecotype" title="Ecotype">Ecotype</a></li> <li><a href="/wiki/Disturbance_(ecology)" title="Disturbance (ecology)">Disturbance</a></li> <li><a href="/wiki/Edge_effects" title="Edge effects">Edge effects</a></li> <li><a href="/wiki/Foster%27s_rule" title="Foster's rule">Foster's rule</a></li> <li><a href="/wiki/Habitat_fragmentation" title="Habitat fragmentation">Habitat fragmentation</a></li> <li><a href="/wiki/Ideal_free_distribution" title="Ideal free distribution">Ideal free distribution</a></li> <li><a href="/wiki/Intermediate_disturbance_hypothesis" title="Intermediate disturbance hypothesis">Intermediate disturbance hypothesis</a></li> <li><a href="/wiki/Insular_biogeography" title="Insular biogeography">Insular biogeography</a></li> <li><a href="/wiki/Land_change_modeling" title="Land change modeling">Land change modeling</a></li> <li><a href="/wiki/Landscape_ecology" title="Landscape ecology">Landscape ecology</a></li> <li><a href="/wiki/Landscape_epidemiology" title="Landscape epidemiology">Landscape epidemiology</a></li> <li><a href="/wiki/Landscape_limnology" title="Landscape limnology">Landscape limnology</a></li> <li><a href="/wiki/Metapopulation" title="Metapopulation">Metapopulation</a></li> <li><a href="/wiki/Patch_dynamics" title="Patch dynamics">Patch dynamics</a></li> <li><a href="/wiki/R/K_selection_theory" title="R/K selection theory"><i>r</i>/<i>K</i> selection theory</a></li> <li><a href="/wiki/Resource_selection_function" title="Resource selection function">Resource selection function</a></li> <li><a href="/wiki/Source%E2%80%93sink_dynamics" title="Source–sink dynamics">Source–sink dynamics</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Ecological_niche" title="Ecological niche">Niche</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ecological_niche" title="Ecological niche">Ecological niche</a></li> <li><a href="/wiki/Ecological_trap" title="Ecological trap">Ecological trap</a></li> <li><a href="/wiki/Ecosystem_engineer" title="Ecosystem engineer">Ecosystem engineer</a></li> <li><a href="/wiki/Species_distribution_modelling" title="Species distribution modelling">Environmental niche modelling</a></li> <li><a href="/wiki/Guild_(ecology)" title="Guild (ecology)">Guild</a></li> <li><a href="/wiki/Habitat" title="Habitat">Habitat</a> <ul><li><a href="/wiki/Marine_habitat" title="Marine habitat">marine habitats</a></li></ul></li> <li><a href="/wiki/Limiting_similarity" title="Limiting similarity">Limiting similarity</a></li> <li><a href="/wiki/Niche_apportionment_models" title="Niche apportionment models">Niche apportionment models</a></li> <li><a href="/wiki/Niche_construction" title="Niche construction">Niche construction</a></li> <li><a href="/wiki/Niche_differentiation" class="mw-redirect" title="Niche differentiation">Niche differentiation</a></li> <li><a href="/wiki/Ontogenetic_niche_shift" title="Ontogenetic niche shift">Ontogenetic niche shift</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Non-trophic_networks" title="Non-trophic networks">Other<br />networks</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Assembly_rules" title="Assembly rules">Assembly rules</a></li> <li><a href="/wiki/Bateman%27s_principle" title="Bateman's principle">Bateman's principle</a></li> <li><a href="/wiki/Bioluminescence" title="Bioluminescence">Bioluminescence</a></li> <li><a href="/wiki/Ecological_collapse" class="mw-redirect" title="Ecological collapse">Ecological collapse</a></li> <li><a href="/wiki/Ecological_debt" title="Ecological debt">Ecological debt</a></li> <li><a href="/wiki/Ecological_debt" title="Ecological debt">Ecological deficit</a></li> <li><a href="/wiki/Energy_flow_(ecology)" title="Energy flow (ecology)">Ecological energetics</a></li> <li><a href="/wiki/Ecological_indicator" title="Ecological indicator">Ecological indicator</a></li> <li><a href="/wiki/Ecological_threshold" title="Ecological threshold">Ecological threshold</a></li> <li><a href="/wiki/Ecosystem_diversity" title="Ecosystem diversity">Ecosystem diversity</a></li> <li><a href="/wiki/Emergence" title="Emergence">Emergence</a></li> <li><a href="/wiki/Extinction_debt" title="Extinction debt">Extinction debt</a></li> <li><a href="/wiki/Kleiber%27s_law" title="Kleiber's law">Kleiber's law</a></li> <li><a href="/wiki/Liebig%27s_law_of_the_minimum" title="Liebig's law of the minimum">Liebig's law of the minimum</a></li> <li><a href="/wiki/Marginal_value_theorem" title="Marginal value theorem">Marginal value theorem</a></li> <li><a href="/wiki/Thorson%27s_rule" title="Thorson's rule">Thorson's rule</a></li> <li><a href="/wiki/Xerosere" title="Xerosere">Xerosere</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Allometry" title="Allometry">Allometry</a></li> <li><a href="/wiki/Alternative_stable_state" title="Alternative stable state">Alternative stable state</a></li> <li><a href="/wiki/Balance_of_nature" title="Balance of nature">Balance of nature</a></li> <li><a href="/wiki/Biological_data_visualization" title="Biological data visualization">Biological data visualization</a></li> <li><a href="/wiki/Ecological_economics" title="Ecological economics">Ecological economics</a></li> <li><a href="/wiki/Ecological_footprint" title="Ecological footprint">Ecological footprint</a></li> <li><a href="/wiki/Ecological_forecasting" title="Ecological forecasting">Ecological forecasting</a></li> <li><a href="/wiki/Environmental_humanities" title="Environmental humanities">Ecological humanities</a></li> <li><a href="/wiki/Ecological_stoichiometry" title="Ecological stoichiometry">Ecological stoichiometry</a></li> <li><a href="/wiki/Ecopath" title="Ecopath">Ecopath</a></li> <li><a href="/wiki/Ecosystem_based_fisheries" class="mw-redirect" title="Ecosystem based fisheries">Ecosystem based fisheries</a></li> <li><a href="/wiki/Endolith" title="Endolith">Endolith</a></li> <li><a href="/wiki/Evolutionary_ecology" title="Evolutionary ecology">Evolutionary ecology</a></li> <li><a href="/wiki/Functional_ecology" title="Functional ecology">Functional ecology</a></li> <li><a href="/wiki/Industrial_ecology" title="Industrial ecology">Industrial ecology</a></li> <li><a href="/wiki/Macroecology" title="Macroecology">Macroecology</a></li> <li><a href="/wiki/Microecosystem" title="Microecosystem">Microecosystem</a></li> <li><a href="/wiki/Natural_environment" title="Natural environment">Natural environment</a></li> <li><a href="/wiki/Regime_shift" title="Regime shift">Regime shift</a></li> <li><a href="/wiki/Sexecology" title="Sexecology">Sexecology</a></li> <li><a href="/wiki/Systems_ecology" title="Systems ecology">Systems ecology</a></li> <li><a href="/wiki/Urban_ecology" title="Urban ecology">Urban ecology</a></li> <li><a href="/wiki/Theoretical_ecology" title="Theoretical ecology">Theoretical ecology</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><a href="/wiki/Outline_of_ecology" title="Outline of ecology">Outline of ecology</a></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" 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