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Beschikbaar in 57 talen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-57" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">57 talen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AC%D8%A7%D9%83_%D8%B2%D9%88%D8%B3%D8%AA%D8%A7%D9%83" title="جاك زوستاك – Arabisch" lang="ar" hreflang="ar" data-title="جاك زوستاك" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%AC%D8%A7%D9%83_%D8%B2%D9%88%D8%B3%D8%AA%D8%A7%D9%83" title="جاك زوستاك – Egyptisch Arabisch" lang="arz" hreflang="arz" data-title="جاك زوستاك" data-language-autonym="مصرى" data-language-local-name="Egyptisch Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%AC%DA%A9_%DA%98%D9%88%D8%B3%D8%AA%D8%A7%DA%A9" title="جک ژوستاک – South Azerbaijani" lang="azb" hreflang="azb" data-title="جک ژوستاک" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%94%D0%B6%D1%8D%D0%BA_%D0%A8%D0%BE%D1%81%D1%82%D0%B0%D0%BA" title="Джэк Шостак – Belarussisch" lang="be" hreflang="be" data-title="Джэк Шостак" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%9C%E0%A7%8D%E0%A6%AF%E0%A6%BE%E0%A6%95_%E0%A6%B6%E0%A6%B8%E0%A6%9F%E0%A7%8D%E0%A6%AF%E0%A6%BE%E0%A6%95" title="জ্যাক শসট্যাক – Bengaals" lang="bn" hreflang="bn" data-title="জ্যাক শসট্যাক" data-language-autonym="বাংলা" data-language-local-name="Bengaals" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Catalaans" lang="ca" hreflang="ca" data-title="Jack W. Szostak" data-language-autonym="Català" data-language-local-name="Catalaans" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D8%AC%D8%A7%DA%A9_%D8%B4%DB%86%D8%B3%D8%AA%D8%A7%DA%A9" title="جاک شۆستاک – Soranî" lang="ckb" hreflang="ckb" data-title="جاک شۆستاک" data-language-autonym="کوردی" data-language-local-name="Soranî" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Tsjechisch" lang="cs" hreflang="cs" data-title="Jack W. Szostak" data-language-autonym="Čeština" data-language-local-name="Tsjechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Deens" lang="da" hreflang="da" data-title="Jack W. Szostak" data-language-autonym="Dansk" data-language-local-name="Deens" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Duits" lang="de" hreflang="de" data-title="Jack Szostak" data-language-autonym="Deutsch" data-language-local-name="Duits" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Engels" lang="en" hreflang="en" data-title="Jack W. Szostak" data-language-autonym="English" data-language-local-name="Engels" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Esperanto" lang="eo" hreflang="eo" data-title="Jack Szostak" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Spaans" lang="es" hreflang="es" data-title="Jack Szostak" data-language-autonym="Español" data-language-local-name="Spaans" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Estisch" lang="et" hreflang="et" data-title="Jack Szostak" data-language-autonym="Eesti" data-language-local-name="Estisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Baskisch" lang="eu" hreflang="eu" data-title="Jack W. Szostak" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%AC%DA%A9_%DA%98%D9%88%D8%B3%D8%AA%D8%A7%DA%A9" title="جک ژوستاک – Perzisch" lang="fa" hreflang="fa" data-title="جک ژوستاک" data-language-autonym="فارسی" data-language-local-name="Perzisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Fins" lang="fi" hreflang="fi" data-title="Jack W. Szostak" data-language-autonym="Suomi" data-language-local-name="Fins" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Frans" lang="fr" hreflang="fr" data-title="Jack Szostak" data-language-autonym="Français" data-language-local-name="Frans" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Galicisch" lang="gl" hreflang="gl" data-title="Jack Szostak" data-language-autonym="Galego" data-language-local-name="Galicisch" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%92%27%D7%A7_%D7%A9%D7%95%D7%A1%D7%98%D7%A7" title="ג'ק שוסטק – Hebreeuws" lang="he" hreflang="he" data-title="ג'ק שוסטק" data-language-autonym="עברית" data-language-local-name="Hebreeuws" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%9C%E0%A5%88%E0%A4%95_%E0%A4%B6%E0%A5%89%E0%A4%B8%E0%A5%8D%E0%A4%A4%E0%A4%95" title="जैक शॉस्तक – Hindi" lang="hi" hreflang="hi" data-title="जैक शॉस्तक" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Kroatisch" lang="hr" hreflang="hr" data-title="Jack W. Szostak" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Hongaars" lang="hu" hreflang="hu" data-title="Jack W. Szostak" data-language-autonym="Magyar" data-language-local-name="Hongaars" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%8B%D5%A5%D5%AF_%D5%87%D5%B8%D5%BD%D5%BF%D5%A1%D5%AF" title="Ջեկ Շոստակ – Armeens" lang="hy" hreflang="hy" data-title="Ջեկ Շոստակ" data-language-autonym="Հայերեն" data-language-local-name="Armeens" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Indonesisch" lang="id" hreflang="id" data-title="Jack W. Szostak" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Jack_William_Szostak" title="Jack William Szostak – Ido" lang="io" hreflang="io" data-title="Jack William Szostak" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – IJslands" lang="is" hreflang="is" data-title="Jack W. Szostak" data-language-autonym="Íslenska" data-language-local-name="IJslands" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Italiaans" lang="it" hreflang="it" data-title="Jack W. Szostak" data-language-autonym="Italiano" data-language-local-name="Italiaans" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B8%E3%83%A3%E3%83%83%E3%82%AF%E3%83%BBW%E3%83%BB%E3%82%B7%E3%83%A7%E3%82%B9%E3%82%BF%E3%82%AF" title="ジャック・W・ショスタク – Japans" lang="ja" hreflang="ja" data-title="ジャック・W・ショスタク" data-language-autonym="日本語" data-language-local-name="Japans" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%94%D0%B6%D0%B5%D0%BA_%D0%A8%D0%BE%D1%81%D1%82%D0%B0%D0%BA" title="Джек Шостак – Kazachs" lang="kk" hreflang="kk" data-title="Джек Шостак" data-language-autonym="Қазақша" data-language-local-name="Kazachs" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%9E%AD_%EC%87%BC%EC%8A%A4%ED%83%9D" title="잭 쇼스택 – Koreaans" lang="ko" hreflang="ko" data-title="잭 쇼스택" data-language-autonym="한국어" data-language-local-name="Koreaans" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Latijn" lang="la" hreflang="la" data-title="Jack Szostak" data-language-autonym="Latina" data-language-local-name="Latijn" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Jack_William_Szostak" title="Jack William Szostak – Litouws" lang="lt" hreflang="lt" data-title="Jack William Szostak" data-language-autonym="Lietuvių" data-language-local-name="Litouws" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%9C%E0%B4%BE%E0%B4%95%E0%B5%8D%E0%B4%95%E0%B5%8D._%E0%B4%A1%E0%B4%AC%E0%B5%8D%E0%B4%B2%E0%B5%8D%E0%B4%AF%E0%B5%82._%E0%B4%B7%E0%B5%8B%E0%B4%B8%E0%B5%8D%E0%B4%B1%E0%B5%8D%E0%B4%B1%E0%B4%BE%E0%B4%95%E0%B5%8D%E0%B4%95%E0%B5%8D" title="ജാക്ക്. ഡബ്ല്യൂ. ഷോസ്റ്റാക്ക് – Malayalam" lang="ml" hreflang="ml" data-title="ജാക്ക്. ഡബ്ല്യൂ. ഷോസ്റ്റാക്ക്" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-mr mw-list-item"><a href="https://mr.wikipedia.org/wiki/%E0%A4%9C%E0%A5%85%E0%A4%95_%E0%A4%A1%E0%A4%AC%E0%A4%B2%E0%A5%8D%E0%A4%AF%E0%A5%82._%E0%A4%9D%E0%A5%8B%E0%A4%B8%E0%A5%8D%E0%A4%9F%E0%A4%BE%E0%A4%95" title="जॅक डबल्यू. झोस्टाक – Marathi" lang="mr" hreflang="mr" data-title="जॅक डबल्यू. झोस्टाक" data-language-autonym="मराठी" data-language-local-name="Marathi" class="interlanguage-link-target"><span>मराठी</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Maleis" lang="ms" hreflang="ms" data-title="Jack W. Szostak" data-language-autonym="Bahasa Melayu" data-language-local-name="Maleis" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%AC%DA%A9_%DA%98%D9%88%D8%B3%D8%AA%D8%A7%DA%A9" title="جک ژوستاک – Mazanderani" lang="mzn" hreflang="mzn" data-title="جک ژوستاک" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Nedersaksisch" lang="nds" hreflang="nds" data-title="Jack Szostak" data-language-autonym="Plattdüütsch" data-language-local-name="Nedersaksisch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Noors - Bokmål" lang="nb" hreflang="nb" data-title="Jack Szostak" data-language-autonym="Norsk bokmål" data-language-local-name="Noors - Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Occitaans" lang="oc" hreflang="oc" data-title="Jack W. Szostak" data-language-autonym="Occitan" data-language-local-name="Occitaans" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Pools" lang="pl" hreflang="pl" data-title="Jack Szostak" data-language-autonym="Polski" data-language-local-name="Pools" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%AC%DB%8C%DA%A9_%D8%B2%D9%88%D8%B3%D8%AA%D8%A7%DA%A9" title="جیک زوستاک – Western Punjabi" lang="pnb" hreflang="pnb" data-title="جیک زوستاک" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Portugees" lang="pt" hreflang="pt" data-title="Jack Szostak" data-language-autonym="Português" data-language-local-name="Portugees" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Roemeens" lang="ro" hreflang="ro" data-title="Jack Szostak" data-language-autonym="Română" data-language-local-name="Roemeens" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A8%D0%BE%D1%81%D1%82%D0%B0%D0%BA,_%D0%94%D0%B6%D0%B5%D0%BA" title="Шостак, Джек – Russisch" lang="ru" hreflang="ru" data-title="Шостак, Джек" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-scn mw-list-item"><a href="https://scn.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Siciliaans" lang="scn" hreflang="scn" data-title="Jack W. Szostak" data-language-autonym="Sicilianu" data-language-local-name="Siciliaans" class="interlanguage-link-target"><span>Sicilianu</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Simple English" lang="en-simple" hreflang="en-simple" data-title="Jack Szostak" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Sloveens" lang="sl" hreflang="sl" data-title="Jack W. Szostak" data-language-autonym="Slovenščina" data-language-local-name="Sloveens" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Zweeds" lang="sv" hreflang="sv" data-title="Jack Szostak" data-language-autonym="Svenska" data-language-local-name="Zweeds" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Swahili" lang="sw" hreflang="sw" data-title="Jack Szostak" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%9C%E0%AE%BE%E0%AE%95%E0%AF%8D_%E0%AE%9A%E0%AF%8B%E0%AE%B8%E0%AF%8D%E0%AE%9F%E0%AE%BE%E0%AE%95%E0%AF%8D" title="ஜாக் சோஸ்டாக் – Tamil" lang="ta" hreflang="ta" data-title="ஜாக் சோஸ்டாக்" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Jack_Szostak" title="Jack Szostak – Turks" lang="tr" hreflang="tr" data-title="Jack Szostak" data-language-autonym="Türkçe" data-language-local-name="Turks" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%94%D0%B6%D0%B5%D0%BA_%D0%A8%D0%BE%D1%81%D1%82%D0%B0%D0%BA" title="Джек Шостак – Oekraïens" lang="uk" hreflang="uk" data-title="Джек Шостак" data-language-autonym="Українська" data-language-local-name="Oekraïens" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%AC%DB%8C%DA%A9_%DA%88%D8%A8%D9%84%DB%8C%D9%88._%D8%B3%D8%B2%D9%88%D8%B3%D9%B9%DA%A9" title="جیک ڈبلیو. سزوسٹک – Urdu" lang="ur" hreflang="ur" data-title="جیک ڈبلیو. سزوسٹک" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Vietnamees" lang="vi" hreflang="vi" data-title="Jack W. Szostak" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamees" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Jack_W._Szostak" title="Jack W. Szostak – Yoruba" lang="yo" hreflang="yo" data-title="Jack W. 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src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/25px-Nobel_prize_medal.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/38px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/50px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span>  Jack W. Szostak </th></tr> <tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center;" colspan="3"><a href="/wiki/9_november" title="9 november">9 november</a> <a href="/wiki/1952" title="1952">1952</a> </th></tr> <tr> <td style="text-align:center; padding:0;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:JSzostak.jpg" class="mw-file-description"><img alt="Jack William Szostak" src="//upload.wikimedia.org/wikipedia/commons/8/88/JSzostak.jpg" decoding="async" width="266" height="381" class="mw-file-element" data-file-width="143" data-file-height="205" /></a></span> </td></tr> <tr> <td style="text-align:center;" colspan="3"><div style="font-size:90%">Jack William Szostak</div> </td></tr> <tr> <th>Geboorteland </th> <td colspan="2"><a href="/wiki/Verenigd_Koninkrijk" title="Verenigd Koninkrijk">Verenigd Koninkrijk</a> </td></tr> <tr> <th>Geboorteplaats </th> <td colspan="2"><a href="/wiki/Londen" title="Londen">Londen</a> </td></tr> <tr> <th>Nationaliteit </th> <td colspan="2"><a href="/wiki/Verenigde_Staten" title="Verenigde Staten">Amerikaans</a> </td></tr> <tr> <th>Nobelprijs </th> <td colspan="2"><a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Fysiologie of Geneeskunde</a> </td></tr> <tr> <th>Jaar </th> <td colspan="2"><a href="/wiki/2009" title="2009">2009</a> </td></tr> <tr> <th>Reden </th> <td colspan="2">Voor hun ontdekking hoe <a href="/wiki/Chromosoom" title="Chromosoom">chromosomen</a> worden beschermd door <a href="/wiki/Telomeer" title="Telomeer">telomeren</a> en het <a href="/wiki/Enzym" title="Enzym">enzym</a> <a href="/wiki/Telomerase" title="Telomerase">telomerase</a> </td></tr> <tr> <th>Samen met </th> <td colspan="2"><a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a><br /><a href="/wiki/Carol_Greider" title="Carol Greider">Carol Greider</a> </td></tr> <tr> <th><span class="nowrap">Voorganger(s)</span> </th> <td colspan="2"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a><br /><a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a><br /><a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> </td></tr> <tr> <th>Opvolger(s) </th> <td colspan="2"><a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Robert Edwards</a> </td></tr> <tr> <td class="infobox-kop notheme center" style="background-color:lightsteelblue; color:inherit;" colspan="3"> <table cellspacing="0" cellpadding="0"> <tbody><tr> <td style="vertical-align:middle;"><b>Portaal</b>  <span class="noviewer" typeof="mw:File"><a href="/wiki/Bestand:Portal.svg" class="mw-file-description" title="Portaalicoon"><img alt="Portaalicoon" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/22px-Portal.svg.png" decoding="async" width="22" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/33px-Portal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/44px-Portal.svg.png 2x" data-file-width="36" data-file-height="32" /></a></span>   </td> <td class="ta-left"><b><a href="/wiki/Portaal:Geneeskunde" title="Portaal:Geneeskunde">Geneeskunde</a> </b> </td></tr></tbody></table> </td></tr> </tbody></table> <p><b>Jack William Szostak</b> (<a href="/wiki/Londen" title="Londen">Londen</a>, <a href="/wiki/9_november" title="9 november">9 november</a> <a href="/wiki/1952" title="1952">1952</a>) is een <a href="/wiki/Verenigde_Staten" title="Verenigde Staten">Amerikaans</a> bioloog van <a href="/wiki/Engeland" title="Engeland">Engelse</a> komaf. Szostak is sinds 2022 als universiteitsprofessor verbonden aan de <a href="/wiki/University_of_Chicago" class="mw-redirect" title="University of Chicago">University of Chicago</a>, daarvoor was hij werkzaam als professor <a href="/wiki/Genetica" title="Genetica">Genetica</a> aan de <a href="/wiki/Harvard_Medical_School" title="Harvard Medical School">Harvard Medical School</a>. In 2009 ontving hij, samen met <a href="/wiki/Carol_Greider" title="Carol Greider">Carol Greider</a> en <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a> "<i>voor hun ontdekking hoe <a href="/wiki/Chromosoom" title="Chromosoom">chromosomen</a> worden beschermd door <a href="/wiki/Telomeer" title="Telomeer">telomeren</a> en het <a href="/wiki/Enzym" title="Enzym">enzym</a> <a href="/wiki/Telomerase" title="Telomerase">telomerase</a></i>". </p> <div class="mw-heading mw-heading2"><h2 id="Biografie">Biografie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Jack_Szostak&veaction=edit&section=1" title="Bewerk dit kopje: Biografie" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Jack_Szostak&action=edit&section=1" title="De broncode bewerken van de sectie: Biografie"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hoewel hij in Londen geboren werd, groeide Szostak op in Canada. Hij behaalde op zijn vijftiende zijn middelbareschooldiploma in Québec en op zijn negentiende zijn bachelordiploma in de celbiologie aan de <a href="/wiki/McGill-universiteit" title="McGill-universiteit">McGill-universiteit</a>. Szostak promoveerde aan de <a href="/wiki/Cornell-universiteit" title="Cornell-universiteit">Cornell-universiteit</a>, waar hij tot 1979 aanbleef als onderzoeksassistent. </p><p>Vervolgens ging hij naar de Harvard Medical School, waar hij in 1988 werd benoemd tot hoogleraar aan de faculteit Genetica en een eigen laboratorium stichtte: het "Sidney Farber Cancer Institute". Daarnaast is hij als moleculair bioloog verbonden aan het Department of Moleculair Biology van het Massachusetts General Hospital en sinds 1998 is hij "Investigator" bij het Howard Hughes Medical Institute (HHMI). </p><p>In september 2022 trad Szostak toe tot de faculteit van de Universiteit van Chicago als universiteitsprofessor, waar hij leiding geeft aan een nieuw interdisciplinair programma genaamd het <i>Origins of Life Initiative</i>. </p> <div class="mw-heading mw-heading2"><h2 id="Werk">Werk</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Jack_Szostak&veaction=edit&section=2" title="Bewerk dit kopje: Werk" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Jack_Szostak&action=edit&section=2" title="De broncode bewerken van de sectie: Werk"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Samen met Elizabeth Blackburn verrichtte Szostak pionierswerk op het gebied van de DNA-uiteinden van chromosomen, telomeren, die een beschermde werking hebben tijdens celdelingen. Samen met Blackburn en Greider won Szostak in 2009 hiervoor de Nobelprijs. In 2006 wonnen ze samen ook al de <a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award for Basic Medical Research</a>.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> In 2008 ontving hij de <a href="/wiki/Dr._A.H._Heinekenprijs" title="Dr. A.H. Heinekenprijs">Dr. A.H. Heinekenprijs</a> voor biochemie en biofysica, hij ontving deze prijs "<i>Voor zijn zeer originele inzichten in fundamentele processen van het leven</i>".<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="toccolours appendix" role="presentation" style="font-size:90%; margin:1em 0 -0.5em; clear:both;"> <div><span style="font-weight:bold">Bronnen, noten en/of referenties</span></div> <ul><li><style data-mw-deduplicate="TemplateStyles:r67679320">.mw-parser-output .taalaanduiding{font-family:sans-serif;font-size:85%;cursor:help;color:var(--color-subtle,#555)}.mw-parser-output .taalaanduiding span{border-bottom:1px dotted var(--color-subtle,#555)}</style><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/2009/summary/">The Nobel Prize in Physiology or Medicine 2009</a>. NobelPrize.org. Nobel Media AB 2018</li> <li><span class="citation" id="CITEREFJ.W._Szostak_&_E.H._Blackburn1982"><span style="font-variant:small-caps;">J.W. Szostak & E.H. Blackburn</span> (1982). <cite>Cloning yeast telomeres on linear plasmid vectors</cite>. <i>Cell</i> <b>29</b> (1): 254-255. <a href="/wiki/PubMed" title="PubMed">PMID</a> <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/6286143">6286143</a>. <a href="/wiki/Digital_object_identifier" title="Digital object identifier">DOI</a>: <a rel="nofollow" class="external text" href="https://doi.org/10.1016/0092-8674(82)90109-X">10.1016/0092-8674(82)90109-X</a>. </span></li></ul> <div class="reflist" style="list-style-type: decimal;"><div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20080918121732/http://www.laskerfoundation.org/awards/2006basic.htm">Laskerfoundation.org</a></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.knaw.nl/nl/prijzen/laureaten/dr-h-p-heinekenprijs-voor-biochemie-en-biofysica/jack-szostak-1952-verenigde-staten">Winnaar Dr. A.H. 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srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/24px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/32px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span></div></div> <div id="Winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" class="navigatie-titel">Winnaars van de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a></div> <div class="navigatie-inhoud mw-collapsible-content"> <p>1901: <a href="/wiki/Emil_Adolf_von_Behring" title="Emil Adolf von Behring">Behring</a> · 1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ross</a> · 1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Finsen</a> · 1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Pavlov</a> · 1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Koch</a> · 1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Golgi</a>, <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Ramón y Cajal</a> · 1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Laveran</a> · 1908: <a href="/wiki/Ilja_Iljitsj_Metsjnikov" title="Ilja Iljitsj Metsjnikov">Mechnikov</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> · 1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> · 1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> · 1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> · 1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> · 1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> · 1914: <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> · 1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> · 1920: <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> · 1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a>, <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> · 1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_Macleod_(medicus)" title="John Macleod (medicus)">Macleod</a> · 1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> · 1926: <a href="/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger">Fibiger</a> · 1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> · 1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> · 1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> · 1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> · 1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> · 1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Adrian</a> · 1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> · 1934: <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a>, <a href="/wiki/George_Minot" title="George Minot">Minot</a>, <a href="/wiki/William_Murphy" title="William Murphy">Murphy</a> · 1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> · 1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> · 1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> · 1938: <a href="/wiki/Corneel_Heymans" title="Corneel Heymans">Heymans</a> · 1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> · 1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> · 1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> · 1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> · 1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> · 1947: <span class="nowrap"><a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a></span>, <span class="nowrap"><a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a></span>, <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> · 1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> · 1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> · 1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> · 1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> · 1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> · 1953: <a href="/wiki/Hans_Krebs_(arts)" title="Hans Krebs (arts)">Krebs</a>,<a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> · 1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> · 1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> · 1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> · 1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> · 1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> · 1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> · 1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> · 1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> · 1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> · 1963: <a href="/wiki/John_Eccles" title="John Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> · 1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a>, <a href="/wiki/Feodor_Felix_Konrad_Lynen" title="Feodor Felix Konrad Lynen">Lynen</a> · 1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Michael_Lwoff" title="André Michael Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> · 1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> · 1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> · 1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> · 1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a>, <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a>, <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> · 1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_Svante_von_Euler" title="Ulf Svante von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> · 1971: <a href="/wiki/Earl_Wilbur_Sutherland_jr." title="Earl Wilbur Sutherland jr.">Sutherland</a> · 1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a>, <a href="/wiki/Rodney_Porter" title="Rodney Porter">Porter</a> · 1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> · 1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">De Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> · 1975: <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Temin</a> · 1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> · 1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> · 1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a href="/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith">Smith</a> · 1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> · 1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> · 1981: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> · 1982: <a href="/wiki/Sune_Karl_Bergstr%C3%B6m" title="Sune Karl Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> · 1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> · 1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a href="/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> · 1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Goldstein</a> · 1986: <a href="/wiki/Stanley_Cohen_(biochemicus)" title="Stanley Cohen (biochemicus)">Cohen</a>, <span class="nowrap"><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a></span> · 1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> · 1988: <a href="/wiki/James_Whyte_Black" title="James Whyte Black">Black</a>, <a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Elion</a>, <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> · 1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> · 1990: <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Murray</a>, <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">Thomas</a> · 1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> · 1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a>, <a href="/wiki/Edwin_Gerhard_Krebs" title="Edwin Gerhard Krebs">Krebs</a> · 1993: <a href="/wiki/Richard_John_Roberts" title="Richard John Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> · 1994: <a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> · 1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <span class="nowrap"><a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a></span>, <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> · 1996: <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a>, <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> · 1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> · 1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a>, <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> · 1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> · 2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> · 2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> · 2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a href="/wiki/John_Edward_Sulston" title="John Edward Sulston">Sulston</a> · 2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> · 2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> · 2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> · 2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> · 2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> · 2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Zur Hausen</a>, <span class="nowrap"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré–Sinoussi</a>,</span> <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> · 2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a>, <a class="mw-selflink selflink">Szostak</a> · 2010: <a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Edwards</a> · 2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> · 2012: <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> · 2013: <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> · 2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">O'Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">Moser</a> · 2015: <a href="/wiki/William_C._Campbell_(wetenschapper)" title="William C. Campbell (wetenschapper)">Campbell</a>, <a href="/wiki/Satoshi_Omura" title="Satoshi Omura">Omura</a>, <a href="/wiki/Youyou_Tu" title="Youyou Tu">Tu</a> · 2016: <a href="/wiki/Yoshinori_Osumi" title="Yoshinori Osumi">Osumi</a> · 2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> · 2018: <a href="/wiki/James_Allison" title="James Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> · 2019: <a href="/wiki/William_Kaelin_jr." title="William Kaelin jr.">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> · 2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> · 2021: <a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> · 2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> · 2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> · 2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div></div> <div class="interProject commons mw-list-item" style="display:none;"><a href="https://commons.wikimedia.org/wiki/Category:Jack_W._Szostak#mw-subcategories" class="extiw" title="commons:Category:Jack W. 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