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Anthony Romilio | The University of Queensland, Australia - Academia.edu
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The 3D digital documentation of these sites is in collaboration with the CSIRO and Flinders University.\r\n\r\nAnthony co-supervises two postgraduate students. The research by these students is aimed to deepen our understanding of specific dinosaur track-sites of the Broome Sandstone. This includes (but not limited to) interpreting the types of track-makers, inferences of palaeo-behaviour, and understanding this in the context of the palaeoenvironment.\r\n\r\nAdditionally, Anthony has been involved for several years with UQ's second largest undergraduate course, first year Genetics. With the success of a teaching grant acquisition, he is an active contributor within team of UQ academics, in ways to improve this course, with the results to roll out during the second half of 2015. 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if ($a.is_logged_in() && $viewedUser.is_current_user()) { $('body').addClass('profile-viewed-by-owner'); } $socialProfiles = [{"id":209629,"link":"http://www.uq.edu.au/dinosaurs/index.html?page=128690\u0026pid=31446","name":"Homepage","link_domain":"www.uq.edu.au","icon":"//www.google.com/s2/u/0/favicons?domain=www.uq.edu.au"}]</script><div id="js-react-on-rails-context" style="display:none" data-rails-context="{"inMailer":false,"i18nLocale":"en","i18nDefaultLocale":"en","href":"https://uq.academia.edu/AnthonyRomilio","location":"/AnthonyRomilio","scheme":"https","host":"uq.academia.edu","port":null,"pathname":"/AnthonyRomilio","search":null,"httpAcceptLanguage":null,"serverSide":false}"></div> <div class="js-react-on-rails-component" style="display:none" data-component-name="ProfileCheckPaperUpdate" data-props="{}" data-trace="false" data-dom-id="ProfileCheckPaperUpdate-react-component-a4aec138-47a7-45c3-be32-d30ec00363f9"></div> <div id="ProfileCheckPaperUpdate-react-component-a4aec138-47a7-45c3-be32-d30ec00363f9"></div> <div class="DesignSystem"><div class="onsite-ping" id="onsite-ping"></div></div><div class="profile-user-info DesignSystem"><div class="social-profile-container"><div class="left-panel-container"><div class="user-info-component-wrapper"><div class="user-summary-cta-container"><div class="user-summary-container"><div class="social-profile-avatar-container"><img class="profile-avatar u-positionAbsolute" alt="Anthony Romilio" border="0" onerror="if (this.src != '//a.academia-assets.com/images/s200_no_pic.png') this.src = '//a.academia-assets.com/images/s200_no_pic.png';" width="200" height="200" src="https://0.academia-photos.com/379100/117483/4472530/s200_anthony.romilio.jpg" /></div><div class="title-container"><h1 class="ds2-5-heading-sans-serif-sm">Anthony Romilio</h1><div class="affiliations-container fake-truncate js-profile-affiliations"><div><a class="u-tcGrayDarker" href="https://uq.academia.edu/">The University of Queensland, Australia</a>, <a class="u-tcGrayDarker" href="https://uq.academia.edu/Departments/School_of_Biological_Sciences/Documents">School of Biological Sciences</a>, <span class="u-tcGrayDarker">Research Assistant</span></div></div></div></div><div class="sidebar-cta-container"><button class="ds2-5-button hidden profile-cta-button grow js-profile-follow-button" data-broccoli-component="user-info.follow-button" data-click-track="profile-user-info-follow-button" data-follow-user-fname="Anthony" data-follow-user-id="379100" data-follow-user-source="profile_button" data-has-google="false"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">add</span>Follow</button><button class="ds2-5-button hidden profile-cta-button grow js-profile-unfollow-button" data-broccoli-component="user-info.unfollow-button" data-click-track="profile-user-info-unfollow-button" data-unfollow-user-id="379100"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">done</span>Following</button></div></div><div class="user-stats-container"><a><div class="stat-container js-profile-followers"><p class="label">Followers</p><p class="data">93</p></div></a><a><div class="stat-container js-profile-followees" data-broccoli-component="user-info.followees-count" data-click-track="profile-expand-user-info-following"><p class="label">Following</p><p class="data">136</p></div></a><a><div class="stat-container js-profile-coauthors" data-broccoli-component="user-info.coauthors-count" data-click-track="profile-expand-user-info-coauthors"><p class="label">Co-authors</p><p class="data">7</p></div></a><span><div class="stat-container"><p class="label"><span class="js-profile-total-view-text">Public Views</span></p><p class="data"><span class="js-profile-view-count"></span></p></div></span></div><div class="user-bio-container"><div class="profile-bio fake-truncate js-profile-about" style="margin: 0px;">Anthony is a Postdoctoral Research Assistant for the Vertebrate Palaeontology and Biomechanics laboratory at The University of Queensland (UQ). <br /> <br />Part of his work involves the documentation of the extensive coastal outcrops north of Broome, Western Australia that contain numerous dinosaur tracksites. The 3D digital documentation of these sites is in collaboration with the CSIRO and Flinders University. <br /> <br />Anthony co-supervises two postgraduate students. The research by these students is aimed to deepen our understanding of specific dinosaur track-sites of the Broome Sandstone. This includes (but not limited to) interpreting the types of track-makers, inferences of palaeo-behaviour, and understanding this in the context of the palaeoenvironment. <br /> <br />Additionally, Anthony has been involved for several years with UQ's second largest undergraduate course, first year Genetics. With the success of a teaching grant acquisition, he is an active contributor within team of UQ academics, in ways to improve this course, with the results to roll out during the second half of 2015.<br /><span class="u-fw700">Supervisors: </span>Dr Steve Salisbury<br /><div class="js-profile-less-about u-linkUnstyled u-tcGrayDarker u-textDecorationUnderline u-displayNone">less</div></div></div><div class="suggested-academics-container"><div class="suggested-academics--header"><h3 class="ds2-5-heading-sans-serif-xs">Related Authors</h3></div><ul class="suggested-user-card-list" data-nosnippet="true"><div class="suggested-user-card"><div class="suggested-user-card__avatar social-profile-avatar-container"><a data-nosnippet="" href="https://fiu.academia.edu/RobertLickliter"><img class="profile-avatar u-positionAbsolute" alt="Robert Lickliter related author profile picture" border="0" onerror="if (this.src != '//a.academia-assets.com/images/s200_no_pic.png') this.src = 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data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/126000529/Simulated_range_of_motion_and_hindfoot_posture_of_Rhoetosaurus_brownei_Longman_1926_Sauropoda_Gravisauria_"><img alt="Research paper thumbnail of Simulated range of motion and hindfoot posture of Rhoetosaurus brownei Longman, 1926 (Sauropoda, Gravisauria)" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title">Simulated range of motion and hindfoot posture of Rhoetosaurus brownei Longman, 1926 (Sauropoda, Gravisauria)</div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="126000529"><a 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{"id":126000529,"title":"Simulated range of motion and hindfoot posture of Rhoetosaurus brownei Longman, 1926 (Sauropoda, Gravisauria)","translated_title":"","metadata":{"publication_date":{"day":null,"month":null,"year":2016,"errors":{}}},"translated_abstract":null,"internal_url":"https://www.academia.edu/126000529/Simulated_range_of_motion_and_hindfoot_posture_of_Rhoetosaurus_brownei_Longman_1926_Sauropoda_Gravisauria_","translated_internal_url":"","created_at":"2024-12-01T22:07:37.539-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[],"slug":"Simulated_range_of_motion_and_hindfoot_posture_of_Rhoetosaurus_brownei_Longman_1926_Sauropoda_Gravisauria_","translated_slug":"","page_count":null,"language":"en","content_type":"Work","summary":null,"owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[],"research_interests":[{"id":48,"name":"Engineering","url":"https://www.academia.edu/Documents/in/Engineering"},{"id":148292,"name":"Sauropoda","url":"https://www.academia.edu/Documents/in/Sauropoda"}],"urls":[{"id":45878318,"url":"https://espace.library.uq.edu.au/view/UQ:573775"}]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-126000529-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="123167319"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/123167319/TRACK_RECOGNITION_VIA_ARTIFICIAL_COGNITION_TRAC_PRELIMINARY_REPORT_ON_THE_APPLICATION_OF_MACHINE_LEARNING_TO_IDENITFY_DINOSAUR_TRACKS"><img alt="Research paper thumbnail of TRACK RECOGNITION VIA ARTIFICIAL COGNITION (TRAC): PRELIMINARY REPORT ON THE APPLICATION OF MACHINE LEARNING TO IDENITFY DINOSAUR TRACKS" class="work-thumbnail" src="https://attachments.academia-assets.com/117665830/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/123167319/TRACK_RECOGNITION_VIA_ARTIFICIAL_COGNITION_TRAC_PRELIMINARY_REPORT_ON_THE_APPLICATION_OF_MACHINE_LEARNING_TO_IDENITFY_DINOSAUR_TRACKS">TRACK RECOGNITION VIA ARTIFICIAL COGNITION (TRAC): PRELIMINARY REPORT ON THE APPLICATION OF MACHINE LEARNING TO IDENITFY DINOSAUR TRACKS</a></div><div class="wp-workCard_item wp-workCard--coauthors"><span>by </span><span><a class="" data-click-track="profile-work-strip-authors" href="https://dinotrax.academia.edu/AMilner">Andrew Milner</a>, <a class="" data-click-track="profile-work-strip-authors" href="https://uq.academia.edu/AnthonyRomilio">Anthony Romilio</a>, and <a class="" data-click-track="profile-work-strip-authors" href="https://independent.academia.edu/HouckKaren">Karen Houck</a></span></div><div class="wp-workCard_item"><span>New Mexico Museum of Natural History and Science Bulletin</span><span>, 2024</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Despite the utilization of new technologies to objectively record and document dinosaur tracks in...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Despite the utilization of new technologies to objectively record and document dinosaur tracks in the field, the categorization and identification of dinosaur tracks remains a largely subjective process. Track Recognition via Artificial Cognition (TRAC) is a project that seeks to provide an objective method for categorizing and identifying dinosaur tracks using track surface photographs. TRAC will employ a combination of different iterations of supervised and unsupervised deep machine learning (ML) approaches to categorize track images drawing from a data set of over 1,000 digital photographs of dinosaur footprint images from around the world. Iterations and adjustments to the ML algorithms will identify natural clusters in the image data set, and our ultimate goal is to build a model capable of assigning an ichnotaxonomic identification to new images that TRAC analyzes. If successful, TRAC can be deployed as an application that can identify dinosaur footprints to ichnotaxon for use by professional paleontologists, citizen scientists, and land managers, with the potential of expanding the scope of the project to include more types of fossils. The TRAC Project is divided into two phases. This paper presents a preliminary report on Phase I, the preparation of the image data set.</span></div><div class="wp-workCard_item"><div class="carousel-container carousel-container--sm" id="profile-work-123167319-figures"><div class="prev-slide-container js-prev-button-container"><button aria-label="Previous" class="carousel-navigation-button js-profile-work-123167319-figures-prev"><span class="material-symbols-outlined" style="font-size: 24px" translate="no">arrow_back_ios</span></button></div><div class="slides-container js-slides-container"><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/6289364/figure-1-examples-of-dinosaur-footprint-images-in-the-trac"><img alt="FIGURE 1. A-M, Examples of dinosaur footprint images in the TRAC image dataset. Locality and ichnotaxonomic data are provided in Table 1. Specimens F and M are replicas of the originals. " class="figure-slide-image" src="https://figures.academia-assets.com/117665830/figure_001.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/6289374/figure-2-track-recognition-via-artificial-cognition-trac"><img alt="" class="figure-slide-image" src="https://figures.academia-assets.com/117665830/figure_002.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/6289382/figure-3-during-the-preliminary-supervised-learning-phase-he"><img alt="FIGURE 3. During the preliminary supervised learning phase, he location of the track on the image should be tagged using an application such as CVAT.ai. Additional image features, such as scale bars, can also be tagged so they are not confused with he track analysis but can be used to provide track dimensions. A, theropod track preserved in the Lower Jurassic Navajo Sandstone near Moab, Utah (Lockley et al., 2021), scale 25 cm. During the tagging process a length may be added to the scale ag. B, orthoimage map of an ankylosaurian track preserved on he underside of a Lower Cretaceous Plainview Sandstone bed near Canon City, Colorado (Kurtz et. al., 2001), scale 22.6 cm. " class="figure-slide-image" src="https://figures.academia-assets.com/117665830/figure_003.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/6289394/table-1-dinosaur-track-examples-pictured-in-with"><img alt="TABLE 1. Dinosaur track examples pictured in Figure | with ichnotaxonomic identification and general locality data. " class="figure-slide-image" src="https://figures.academia-assets.com/117665830/table_001.jpg" /></a></figure></div><div class="next-slide-container js-next-button-container"><button aria-label="Next" class="carousel-navigation-button js-profile-work-123167319-figures-next"><span class="material-symbols-outlined" style="font-size: 24px" translate="no">arrow_forward_ios</span></button></div></div></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="de01bb92cb255a4c7e4e29b3796f8097" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":117665830,"asset_id":123167319,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/117665830/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="123167319"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="123167319"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 123167319; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=123167319]").text(description); $(".js-view-count[data-work-id=123167319]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 123167319; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='123167319']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "de01bb92cb255a4c7e4e29b3796f8097" } } $('.js-work-strip[data-work-id=123167319]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":123167319,"title":"TRACK RECOGNITION VIA ARTIFICIAL COGNITION (TRAC): PRELIMINARY REPORT ON THE APPLICATION OF MACHINE LEARNING TO IDENITFY DINOSAUR TRACKS","translated_title":"","metadata":{"abstract":"Despite the utilization of new technologies to objectively record and document dinosaur tracks in the field, the categorization and identification of dinosaur tracks remains a largely subjective process. 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Chin","title":"TRACK RECOGNITION VIA ARTIFICIAL COGNITION (TRAC): PRELIMINARY REPORT ON THE APPLICATION OF MACHINE LEARNING TO IDENITFY DINOSAUR TRACKS"},{"id":42269596,"work_id":123167319,"tagging_user_id":52822449,"tagged_user_id":517737,"co_author_invite_id":null,"email":"i***u@hotmail.com","display_order":9,"name":"Ignacio Díaz-Martínez","title":"TRACK RECOGNITION VIA ARTIFICIAL COGNITION (TRAC): PRELIMINARY REPORT ON THE APPLICATION OF MACHINE LEARNING TO IDENITFY DINOSAUR TRACKS"},{"id":42269597,"work_id":123167319,"tagging_user_id":52822449,"tagged_user_id":703973,"co_author_invite_id":null,"email":"x***a@gmail.com","affiliation":"China University of Geosciences(Beijing)","display_order":10,"name":"Lida Xing","title":"TRACK RECOGNITION VIA ARTIFICIAL COGNITION (TRAC): PRELIMINARY REPORT ON THE APPLICATION OF MACHINE LEARNING TO IDENITFY DINOSAUR 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TRACKS"}],"downloadable_attachments":[{"id":117665830,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/117665830/thumbnails/1.jpg","file_name":"26_Murphey_et_al_02.pdf","download_url":"https://www.academia.edu/attachments/117665830/download_file","bulk_download_file_name":"TRACK_RECOGNITION_VIA_ARTIFICIAL_COGNITI.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/117665830/26_Murphey_et_al_02-libre.pdf?1724469923=\u0026response-content-disposition=attachment%3B+filename%3DTRACK_RECOGNITION_VIA_ARTIFICIAL_COGNITI.pdf\u0026Expires=1743195464\u0026Signature=dZFjPEC1mIOVU~UN2HmJ-QdobCSL3wR~ceC2Ex6GrcE3eh~c-M4WZtaLp3tLdzF9sHhyMuL2-9VblQ6XKlqmWBvadg53Ns7Gj3c6ANkBHBkIq4ngSONAydZIMP6HDCludO7m2hl3Q4aS4y36Df0wtrMwEH~iS4yR3iIOfDd2AAVjky20WDMCSlj5eBi~6e5fD8qYkiNSpjR5YUmUVAdajePrMJqwH7azXQ12YzOuLzypR13RmBoo02CIUa09-MPDpJu8LuOTBXs3fcw8rvaK5tNXsDdIgQ8gsgZ-Sf42QtEpvfw8fveyrmGHD94iA9h72L5dyATwfYqghbrwN6vd0A__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"TRACK_RECOGNITION_VIA_ARTIFICIAL_COGNITION_TRAC_PRELIMINARY_REPORT_ON_THE_APPLICATION_OF_MACHINE_LEARNING_TO_IDENITFY_DINOSAUR_TRACKS","translated_slug":"","page_count":8,"language":"en","content_type":"Work","summary":"Despite the utilization of new technologies to objectively record and document dinosaur tracks in the field, the categorization and identification of dinosaur tracks remains a largely subjective process. Track Recognition via Artificial Cognition (TRAC) is a project that seeks to provide an objective method for categorizing and identifying dinosaur tracks using track surface photographs. TRAC will employ a combination of different iterations of supervised and unsupervised deep machine learning (ML) approaches to categorize track images drawing from a data set of over 1,000 digital photographs of dinosaur footprint images from around the world. Iterations and adjustments to the ML algorithms will identify natural clusters in the image data set, and our ultimate goal is to build a model capable of assigning an ichnotaxonomic identification to new images that TRAC analyzes. If successful, TRAC can be deployed as an application that can identify dinosaur footprints to ichnotaxon for use by professional paleontologists, citizen scientists, and land managers, with the potential of expanding the scope of the project to include more types of fossils. The TRAC Project is divided into two phases. This paper presents a preliminary report on Phase I, the preparation of the image data set.","owner":{"id":52822449,"first_name":"Andrew","middle_initials":null,"last_name":"Milner","page_name":"AMilner","domain_name":"dinotrax","created_at":"2016-09-01T09:11:54.071-07:00","display_name":"Andrew Milner","url":"https://dinotrax.academia.edu/AMilner"},"attachments":[{"id":117665830,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/117665830/thumbnails/1.jpg","file_name":"26_Murphey_et_al_02.pdf","download_url":"https://www.academia.edu/attachments/117665830/download_file","bulk_download_file_name":"TRACK_RECOGNITION_VIA_ARTIFICIAL_COGNITI.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/117665830/26_Murphey_et_al_02-libre.pdf?1724469923=\u0026response-content-disposition=attachment%3B+filename%3DTRACK_RECOGNITION_VIA_ARTIFICIAL_COGNITI.pdf\u0026Expires=1743195464\u0026Signature=dZFjPEC1mIOVU~UN2HmJ-QdobCSL3wR~ceC2Ex6GrcE3eh~c-M4WZtaLp3tLdzF9sHhyMuL2-9VblQ6XKlqmWBvadg53Ns7Gj3c6ANkBHBkIq4ngSONAydZIMP6HDCludO7m2hl3Q4aS4y36Df0wtrMwEH~iS4yR3iIOfDd2AAVjky20WDMCSlj5eBi~6e5fD8qYkiNSpjR5YUmUVAdajePrMJqwH7azXQ12YzOuLzypR13RmBoo02CIUa09-MPDpJu8LuOTBXs3fcw8rvaK5tNXsDdIgQ8gsgZ-Sf42QtEpvfw8fveyrmGHD94iA9h72L5dyATwfYqghbrwN6vd0A__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":417,"name":"Paleontology","url":"https://www.academia.edu/Documents/in/Paleontology"},{"id":2588,"name":"Ichnology","url":"https://www.academia.edu/Documents/in/Ichnology"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (true) { Aedu.setUpFigureCarousel('profile-work-123167319-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="65567150"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/65567150/Reconstruction_of_the_motion_and_hindfoot_posture_of_Rhoetosaurus_brownei_Longman_1926_Sauropoda_Gravisauria_"><img alt="Research paper thumbnail of Reconstruction of the motion and hindfoot posture of Rhoetosaurus brownei Longman, 1926 (Sauropoda, Gravisauria)" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title">Reconstruction of the motion and hindfoot posture of Rhoetosaurus brownei Longman, 1926 (Sauropoda, Gravisauria)</div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">The Middle Jurassic (Late Bathonian–Middle Callovian) basal sauropod Rhoetosaurus brownei is the ...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">The Middle Jurassic (Late Bathonian–Middle Callovian) basal sauropod Rhoetosaurus brownei is the largest known pre-Cretaceous terrestrial Australian vertebrate. Represented as a near-complete right hind limb, key aspects of the palaeobiology of Rhoetosaurus remain unresolved. The current study combines three-dimensional modelling and photogrammetry to reconstruct the range of motion (ROM) of the foot of Rhoetosaurus to elicit pedal flexibility and posture. Our findings show that Rhoetosaurus had a high degree of digit mobility at all the metatarsophalangeal joints (total excursion angle &amp;gt;80°) and in the sagittal, transverse and frontal planes at the distal-most interphalangeal joints (excursion angle &amp;gt;100°). The position of the autopodial articular surfaces suggests that the metarsophalangeal joints were permanently elevated in life, an indicator of skeletal digitigrady. This joint may have been ventrally connected with the plantar aspect of the foot by a compliant fibrous pad, consistent with sauropod track data, and therefore likely to have been functional plantigrade, as has been previously suggested. These results provide new insights into the simulated ROM and posture of the hind foot of Rhoetosaurus; we recognize that additional parameters (including loading regimes and ichnological data) provide future modelling aspects to further our understanding of Rhoetosaurus pedal biomechanics.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="65567150"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="65567150"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 65567150; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=65567150]").text(description); $(".js-view-count[data-work-id=65567150]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 65567150; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='65567150']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=65567150]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":65567150,"title":"Reconstruction of the motion and hindfoot posture of Rhoetosaurus brownei Longman, 1926 (Sauropoda, Gravisauria)","translated_title":"","metadata":{"abstract":"The Middle Jurassic (Late Bathonian–Middle Callovian) basal sauropod Rhoetosaurus brownei is the largest known pre-Cretaceous terrestrial Australian vertebrate. Represented as a near-complete right hind limb, key aspects of the palaeobiology of Rhoetosaurus remain unresolved. The current study combines three-dimensional modelling and photogrammetry to reconstruct the range of motion (ROM) of the foot of Rhoetosaurus to elicit pedal flexibility and posture. Our findings show that Rhoetosaurus had a high degree of digit mobility at all the metatarsophalangeal joints (total excursion angle \u0026amp;gt;80°) and in the sagittal, transverse and frontal planes at the distal-most interphalangeal joints (excursion angle \u0026amp;gt;100°). The position of the autopodial articular surfaces suggests that the metarsophalangeal joints were permanently elevated in life, an indicator of skeletal digitigrady. This joint may have been ventrally connected with the plantar aspect of the foot by a compliant fibrous pad, consistent with sauropod track data, and therefore likely to have been functional plantigrade, as has been previously suggested. These results provide new insights into the simulated ROM and posture of the hind foot of Rhoetosaurus; we recognize that additional parameters (including loading regimes and ichnological data) provide future modelling aspects to further our understanding of Rhoetosaurus pedal biomechanics.","publication_date":{"day":null,"month":null,"year":2016,"errors":{}}},"translated_abstract":"The Middle Jurassic (Late Bathonian–Middle Callovian) basal sauropod Rhoetosaurus brownei is the largest known pre-Cretaceous terrestrial Australian vertebrate. Represented as a near-complete right hind limb, key aspects of the palaeobiology of Rhoetosaurus remain unresolved. The current study combines three-dimensional modelling and photogrammetry to reconstruct the range of motion (ROM) of the foot of Rhoetosaurus to elicit pedal flexibility and posture. Our findings show that Rhoetosaurus had a high degree of digit mobility at all the metatarsophalangeal joints (total excursion angle \u0026amp;gt;80°) and in the sagittal, transverse and frontal planes at the distal-most interphalangeal joints (excursion angle \u0026amp;gt;100°). The position of the autopodial articular surfaces suggests that the metarsophalangeal joints were permanently elevated in life, an indicator of skeletal digitigrady. This joint may have been ventrally connected with the plantar aspect of the foot by a compliant fibrous pad, consistent with sauropod track data, and therefore likely to have been functional plantigrade, as has been previously suggested. These results provide new insights into the simulated ROM and posture of the hind foot of Rhoetosaurus; we recognize that additional parameters (including loading regimes and ichnological data) provide future modelling aspects to further our understanding of Rhoetosaurus pedal biomechanics.","internal_url":"https://www.academia.edu/65567150/Reconstruction_of_the_motion_and_hindfoot_posture_of_Rhoetosaurus_brownei_Longman_1926_Sauropoda_Gravisauria_","translated_internal_url":"","created_at":"2021-12-22T14:00:08.439-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[],"slug":"Reconstruction_of_the_motion_and_hindfoot_posture_of_Rhoetosaurus_brownei_Longman_1926_Sauropoda_Gravisauria_","translated_slug":"","page_count":null,"language":"en","content_type":"Work","summary":"The Middle Jurassic (Late Bathonian–Middle Callovian) basal sauropod Rhoetosaurus brownei is the largest known pre-Cretaceous terrestrial Australian vertebrate. Represented as a near-complete right hind limb, key aspects of the palaeobiology of Rhoetosaurus remain unresolved. The current study combines three-dimensional modelling and photogrammetry to reconstruct the range of motion (ROM) of the foot of Rhoetosaurus to elicit pedal flexibility and posture. Our findings show that Rhoetosaurus had a high degree of digit mobility at all the metatarsophalangeal joints (total excursion angle \u0026amp;gt;80°) and in the sagittal, transverse and frontal planes at the distal-most interphalangeal joints (excursion angle \u0026amp;gt;100°). The position of the autopodial articular surfaces suggests that the metarsophalangeal joints were permanently elevated in life, an indicator of skeletal digitigrady. This joint may have been ventrally connected with the plantar aspect of the foot by a compliant fibrous pad, consistent with sauropod track data, and therefore likely to have been functional plantigrade, as has been previously suggested. 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-65567150-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="65567149"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/65567149/Re_examination_of_the_Lark_Quarry_dinosaur_tracksite_reveals_an_absence_of_tracks_referable_to_nonavian_theropods"><img alt="Research paper thumbnail of Re-examination of the Lark Quarry dinosaur tracksite reveals an absence of tracks referable to nonavian theropods" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title">Re-examination of the Lark Quarry dinosaur tracksite reveals an absence of tracks referable to nonavian theropods</div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="65567149"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="65567149"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 65567149; 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-65567149-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="65567147"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/65567147/Dinosaur_stampede_stopped_in_its_tracks"><img alt="Research paper thumbnail of Dinosaur stampede stopped in its tracks" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title">Dinosaur stampede stopped in its tracks</div><div class="wp-workCard_item"><span>Australasian science</span><span>, 2015</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">A forensic analysis of ancient footprints has cast doubt on claims that a dinosaur stampede took ...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">A forensic analysis of ancient footprints has cast doubt on claims that a dinosaur stampede took place at Lark Quarry.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="65567147"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="65567147"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 65567147; 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-65567147-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="65567146"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/65567146/Dinosaur_tracks_from_the_Walmadany_area_James_Price_Point_Dampier_Peninsula_Western_Australia"><img alt="Research paper thumbnail of Dinosaur tracks from the Walmadany area (James Price Point), Dampier Peninsula, Western Australia" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title">Dinosaur tracks from the Walmadany area (James Price Point), Dampier Peninsula, Western Australia</div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">The Early Cretaceous (Valanginian–Barremian) Broome Sandstone of the Dampier Peninsula, Western A...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">The Early Cretaceous (Valanginian–Barremian) Broome Sandstone of the Dampier Peninsula, Western Australia, has often been touted as having one of the most diverse dinosaurian ichnofaunas in the world. Yet with the exception of tracks assigned to the theropodan ichnotaxon Megalosauropus broomensis in 1967, and more recent work on the preservational characteristics of sauropodan tracks, descriptive accounts of the types of dinosaurian tracks that occur in the Broome Sandstone have been limited, with reports ranging from 6–16 different ichnotaxa. As such, the overall scientific significance of the ichnofauna has remained enigmatic. At the request of Goolarabooloo Traditional Custodians, 300+ hours of ichnological survey work was undertaken on the 25 km stretch of coastline in the vicinity of Walmadany (James Price Point) from 2011–2013. 48 discrete tracksites were identified in this area, containing upward of 20 track types referable to theropods, sauropods, ornithopods and thyreophora...</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="65567146"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="65567146"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 65567146; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=65567146]").text(description); $(".js-view-count[data-work-id=65567146]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 65567146; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='65567146']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=65567146]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":65567146,"title":"Dinosaur tracks from the Walmadany area (James Price Point), Dampier Peninsula, Western Australia","translated_title":"","metadata":{"abstract":"The Early Cretaceous (Valanginian–Barremian) Broome Sandstone of the Dampier Peninsula, Western Australia, has often been touted as having one of the most diverse dinosaurian ichnofaunas in the world. 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Here, we describe a diminutive tridactyl track (10.2 mm in length) from the Lower Jura...","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[],"research_interests":[{"id":155,"name":"Evolutionary Biology","url":"https://www.academia.edu/Documents/in/Evolutionary_Biology"},{"id":406,"name":"Geology","url":"https://www.academia.edu/Documents/in/Geology"},{"id":9846,"name":"Ecology","url":"https://www.academia.edu/Documents/in/Ecology"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-65567145-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="65567144"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/65567144/Three_dimensional_analysis_of_the_largest_tracks_from_the_Lark_Quarry_dinosaur_tracksite_central_western_Queensland"><img alt="Research paper thumbnail of Three-dimensional analysis of the largest tracks from the Lark Quarry dinosaur tracksite, central-western Queensland" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title">Three-dimensional analysis of the largest tracks from the Lark Quarry dinosaur tracksite, central-western Queensland</div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="65567144"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="65567144"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 65567144; 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-65567144-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="65564666"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/65564666/Lower_cretaceous_avian_dominated_theropod_thyreophoran_pterosaur_and_turtle_track_assemblages_from_the_Tugulu_Group_Xinjiang_China_ichnotaxonomy_and_palaeoecology"><img alt="Research paper thumbnail of Lower cretaceous avian-dominated, theropod, thyreophoran, pterosaur and turtle track assemblages from the Tugulu Group, Xinjiang, China: ichnotaxonomy and palaeoecology" class="work-thumbnail" src="https://attachments.academia-assets.com/77104259/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/65564666/Lower_cretaceous_avian_dominated_theropod_thyreophoran_pterosaur_and_turtle_track_assemblages_from_the_Tugulu_Group_Xinjiang_China_ichnotaxonomy_and_palaeoecology">Lower cretaceous avian-dominated, theropod, thyreophoran, pterosaur and turtle track assemblages from the Tugulu Group, Xinjiang, China: ichnotaxonomy and palaeoecology</a></div><div class="wp-workCard_item"><span>PeerJ</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe District, Karamay City, Xinjiang) have been an abundant source of some of the largest Lower Cretaceous track collections from China. They originate from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, variously divided into four formations and subgroups in the northwestern margin of the Junggar Basin. The large Huangyangquan track assemblages occur in the Lower layer/Subgroup II. Similarly-composed track assemblages also occur at the smaller Asphaltite site in the Upper Layer/Subgroup III. The Huangyangquan assemblages have yielded more than 1,500 identified tracks including abundant tracks of avian and non-avian theropods, pterosaurs and turtles and less abundant tracks of stegosaurs. Previous avian track identifications have been reassessed to conclude that Moguiornipes robustus is a taphotaxon and Koreanaornis dodsoni might be better accommodated in the ichno...</span></div><div class="wp-workCard_item"><div class="carousel-container carousel-container--sm" id="profile-work-65564666-figures"><div class="prev-slide-container js-prev-button-container"><button aria-label="Previous" class="carousel-navigation-button js-profile-work-65564666-figures-prev"><span class="material-symbols-outlined" style="font-size: 24px" translate="no">arrow_back_ios</span></button></div><div class="slides-container js-slides-container"><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095424/figure-1-location-of-the-huangyangquan-track-site-in"><img alt="Figure 1 Location of the Huangyangquan track site in Xinjiang, China. (A) East Asia region; (B) Wuerhe area; (C) Huangyangquan sites (A-K). This Huangyangquan area is currently protected by the local reservoir management department. There are isolation fences around the reservoir, which protects the footprint to a certain extent. Full-size Mal DOI: 10.7717/peerj.11476/fig-1 GEOLOGICAL SETTING Tugulu group—depositional environment, stratigraphy and age " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_001.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095426/figure-2-stratigraphic-section-of-the-lower-layer-of-the"><img alt="Figure 2 Stratigraphic section of the Lower Layer of the Tugulu Group at the Wuerhe area and Huangyangquan track sites (A, C, E, F, G, I and K) (emended from Dong, 1973a). Full-size Mal DOI: 10.7717/peerj.11476/fig-2 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_002.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095429/figure-3-invertebrate-traces-from-the-huangyangquan-area"><img alt="Figure 3 Invertebrate traces from the Huangyangquan area. Scoyenia gracilis (=Sc) (A, B), Cochlichnus anguineus (C), Helminthoidichnites tenuis (=Ht) (D, E, F), Mermia carickensis (=Me) (F, G), Cruziana problematica (=Cr) (F, G), Rusophycus isp. (=Ru) (F, G), Treptichnus isp. (=Tr) (H), Planolites montanus (=P1) (H). Scale bar in (A), (B), (F), (G) and (H) = 10 cm; ruler in (C), (D), (E) scale in centimeter. Full-size Mw DOI: 10.7717/peerj.11476/fig-3 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_003.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095433/figure-4-invertebrate-traces-from-the-huangyangquan-area"><img alt="Figure 4 Invertebrate traces from the Huangyangquan area. Lockeia siliquaria (=Lo) (A, C), Archaeonassa fossulata (=Ar) (B) and Helminthopsis abeli (=He) (D). Scale bar = 10 cm Full-size ha DOI: 10.7717/peerj.11476/fig-4 Huangyangquan area " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_004.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095437/figure-5-the-interpretive-outline-drawing-of-theropod-tracks"><img alt="Figure 5 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-A-3 (A), 16 (B), 17 (C), 18 (D), 20 (E), 23 (F), 24 (G), 21 (H); HYQ-D-5 (I), 1 (J), 2 (K), 6 (L), 12 (M), (S), 3 (T), and HYQ-A-15 (U). 7 (N), 8 (O), 11 (P) 9 (Q); HYQ-L-1 (R), 2 Full-size wr" DOI: 10.7717/peerj.11476/fig-5 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_005.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095439/figure-6-the-interpretive-outline-drawing-of-theropod-tracks"><img alt="Figure 6 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-D-3 (A), 16 (B), 4 (C), 10 (D); HYQ-E-6 (E), 10 (F), 9 (G). Full-size ns DOI: 10.7717/peerj.11476/fig-6 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_006.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095444/figure-7-lower-cretaceous-avian-dominated-theropod"><img alt="" class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_007.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095445/figure-8-the-interpretive-outline-drawing-of-theropod-tracks"><img alt="Figure 8 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-F-13 (A), 14 (B), 15 (C), 16 (D); HYQ-H-3 (E); HYQ-G-1 (F); HYQ-H-1 (G); HYQ-J-1 (H); HYQ-K-1 (I), 2 9). Full-size ns DOI: 10.7717/peerj.11476/fig-8 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_008.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095448/figure-9-the-interpretive-outline-drawing-of-theropod-tracks"><img alt="Figure 9 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-H-4 (A), 2 (B). Full-size ns DOI: 10.7717/peerj.11476/fig-9 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_009.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095452/figure-10-perspective-of-hyq-left-pedal-impression-dorsal"><img alt="Figure 10 Perspective of HYQ-E-11 (left pedal impression). (A) Dorsal view; (B) Ventral view; (C) Anterior view; (D) Posterior view; (E) Medial view; and (F) Lateral view. The track surface images are shown as orthophotographs, ambient occlusion images, and elevation images. Full-size | DOI: 10.7717/peerj.11476/fig-10 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_010.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095457/figure-11-the-interpretive-outline-drawing-of-theropod"><img alt="Figure 11 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-D-13. Full-size Mal DOI: 10.7717/peerj.11476/fig-11 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_011.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095462/figure-12-jialingpus-reflects-small-to-medium-sized-theropod"><img alt="Jialingpus reflects a small to medium-sized theropod trackmaker which registered grallatorid tracks akin to Grallator (sensu lato). However, a distinctive, centrally placed, swollen metatarsophalangeal area is a distinctive and diagnostic feature of well- preserved Jialingpus, making it different from Eubrontes—Grallator-Anchisauripus plexus tracks. In the best-preserved specimens of Jialingpus, a crease separates the small metatarsophalangeal pad region behind digit II and the larger region behind digit IV. This trait is clearly present in the theropod morphotype A tracks from the Huangyangquan site, which can, therefore, be assigned to Jialingpus. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_012.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095468/figure-13-the-interpretive-outline-drawing-of-theropod"><img alt="Figure 13 The interpretive outline drawing of theropod tracks at Huangyangquan sites. Specimens numbers: YLSNHM01243 (A); HYQ-13 (B); YLSNHM00978 (C), 00980 (D), 01236 (E), 00977 (E), 00979 (G), 01225 (H), 01237 (I), 01223 (J), 01226 (K), 01247 (L), 01268 (M), 01250 (N), 01269 (O), 01270 (P), 01271 (Q), 01272 (R), 01273 (S), 01274 (T), 01275 (U), 01276 (V), 01277 (W), 01279 (X), 01280 (Y), 01278 (Z). Full-size My DOI: 10.7717/peerj.11476/fig- 13 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_013.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095478/figure-14-the-interpretive-outline-drawing-of-theropod-and"><img alt="Figure 14 The interpretive outline drawing of theropod and bird tracks at Huangyangquan sites: YLSNHMO01246 (A); HYQ-21 (B); YLSNHMO01244 (C); HYQ-15 (D), 17 (E), 19 (F), 16 (G), 14 (H), 9 (1); YLSNHMO012353 (J). Full-size My DOI: 10.7717/peerj.11476/fig-14 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_014.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095488/figure-15-the-interpretive-outline-drawing-of-theropod-and"><img alt="Figure 15 The interpretive outline drawing of theropod and bird tracks at Huangyangquan sites: HYQ-E-1 (A), 2 (B), 3 (C)} HYQ-K-3 (D); HYQ-J-9 (E), 10 (F). Full-size Ry DOI: 10.7717/peerj.11476/fig-15 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_015.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095493/figure-16-asianopudus-niui-comparisons-between-holotypes-of"><img alt="Figure 16 Asianopudus niui. Comparisons between holotypes of Asianopudus and Chapus from China. (A) A. niui, (B) C. lockleyi and (C) A. robustus, showing anterior triangle and ratio of digit III/ Footprint length. Note similarities between A. niui and C. lockleyi. See text for details. Full-size My DOI: 10.7717/peerj.11476/fig-16 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_016.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095501/figure-17-photograph-of-theropod-track-ylsnhm-and-hyq-at"><img alt="Figure 17 Photograph of theropod track YLSNHM01265 and HYQ-A-18 at Huangyangquan site: Dorsal (A), Anterior (B), Lateral (C) and Medial (D) views of YLSNHMO01265; Dorsal (E) view of HYQ-A-18. The black arrow indicate the scale scratch lines along the side. r..11 2: lal TAAAT. 1A 9910 sew eet DIAM CLR +I " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_017.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095508/figure-18-photograph-of-theropod-track-hyq-and-at"><img alt="Figure 18 Photograph of theropod track HYQ-K-1 (A) and 2 (B) at Huangyangquan site. Full-size RY DOI: 10.7717/peerj.11476/fig-1 for establishing a new ichnospecies. First, Belvedere & Farlow (2016) and Marchetti et al. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_018.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095513/figure-19-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 19 Photograph (A) and interpretive outline drawing (B) of bird tracks at Huangyangquan site E: HYQ-E-2. Full-size MY DOI: 10.7717/peerj.11476/fig-19 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_019.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095521/figure-20-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 20 Photograph (A) and interpretive outline drawing (B) of bird tracks YLSNHMO01231, and the photograph (C) of bird tracks YLSNHM 01295 at Huangyangquan sites. Full-size My DOI: 10.7717/peerj.11476/fig-20 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_020.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095527/figure-21-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 21 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01203 (A), 01210 (B), 01213 (C), 01212 (D), 01234 (E). Full-size Kal DOI: 10.7717/peerj.11476/fig-21 Comparison & discussion " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_021.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095535/figure-22-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 22 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01214 (A), 01224 (B), 01215 (C), 01216 (D), 01217 (E), 01230 (F), 01245 (G). Full-size Mal DOI: 10.7717/peerj.11476/fig-22 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_022.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095542/figure-23-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 23 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01227 (A), 01238 (B), 01248 (C), 01232 (D), 01251 (E), 01254 (EF). Full-size RY DOI: 10.7717/peerj.11476/fig-23 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_023.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095548/figure-24-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 24 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01255 (A), 01281 (B), 01284 (C), 01286 (D), 01287 (E), 01283 (E), 01285 (G), 01288 (H), 01289 (I), 01290 (J), 01291 (K), 01292 (L), 01293 (M), 01294 (N). Full-size My DOI: 10.7717/peerj.11476/fig-24 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_024.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095552/figure-25-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 25 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01201 (A); HYQ-12 (B); YLSNHMO01201 (C), 01242 (D). Full-size My DOI: 10.7717/peerj.11476/fig-25 Koreanornipodidae includes the small subsymmetric functionally-tridactyl tracks ‘oreanaornis (Kim, 1969; Lockley et al., 1992) and Pullornipes (Lockley et al., 2006b). he latter differs from Koreanaornis mostly in having a small postero-medially directed allux, which is absent or very rare and inconspicuous in Koreanaornis. Features of ‘oreanaornis dodsoni that distinguish it from K. hamanensis include: smaller divaricatiot ngles between digits II and IV, greater overall track length and width, persistent absenc f digit I, and absence of digital pad impressions. Compared with the original type ‘oreanaornis hamanensis, K. dodsoni is larger, with a reported, more variable size range o .0-6.3 cm (Xing et al., 2011). The larger measurement is markedly inconsistent with th lentification as Koreanaornis. Likewise the K. dodsoni holotype (length 4.4 cm, width .7 cm) is more typical of Aquatilavipes, (ichnofamily Avipedidae according to Sarjeant & angston, 1994) and raises the question of whether K. dodsoni would be better ccommodated in Aquatilavipes dodsoni as a new combination. However, because ‘retaceous shorebird tracks need a comprehensive ichnotaxonomic revision, we avoid a AaArrataA anacatikKnrmant " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_025.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095557/figure-26-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 26 Photograph (A) and interpretive outline drawing (B)of bird tracks at Huangyangquan sites: YLSNHMO01201-4-6. Full-size My DOI: 10.7717/peerj.11476/fig-26 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_026.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095567/figure-27-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 27 Photograph (A) and interpretive outline drawing (B) of bird tracks at Huangyangquan sites: YLSNHMO01241. Full-size My DOI: 10.7717/peerj.11476/fig-27 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_027.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095572/figure-28-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 28 The interpretive outline drawing of bird tracks at Huangyangquan sites: HYQ-18 (A), 11 (B); YLSNHM01282 (C); HYQ-12 (D). Full-size MY DOI: 10.7717/peerj.11476/fig-28 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_028.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095580/figure-29-the-interpretive-outline-drawing-of-stegosaur"><img alt="Figure 29 The interpretive outline drawing of stegosaur tracks at Huangyangquan sites: YLSNHM01256 (A), 01258 (B), 01260 (C), 01263 (D); HYQ-D-14 (E), 15 (£). Full-size My DOI: 10.7717/peerj.11476/fig-29 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_029.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095584/figure-30-the-manus-prints-are-asymmetrical-digitigrade-and"><img alt="The manus prints are asymmetrical, digitigrade, and consist of three digit impression casts. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_030.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095592/figure-31-photograph-elevation-image-and-interpretive"><img alt="Figure 31 Photograph (A), elevation image (B) and interpretive outline drawing (C) of pterosaur tracks HYQ-7 at Huangyangquan sites. Full-size Mal DOI: 10.7717/peerj.11476/fig-31 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_031.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095597/figure-32-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 32 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-A-1 (A), 2 (B), 5 (C), 8 (D), 7 (E), 9 (F), 10 (G), 12 (H), 26 (I). Full-size RY DOI: 10.7717/peerj.11476/fig-32 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_032.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095602/figure-33-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 33 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-A-13 (A), 14 (B); HYQ-J-2 (C); HYQ-A-25 (D); HYQ-E-4 (E); HYQ-J-3 (F). Full-size MY DOI: 10.7717/peerj.11476/fig-33 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_033.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095603/figure-34-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 34 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-J-4 (A), 5 (B). Full-size My DOI: 10.7717/peerj.11476/fig-34 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_034.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095607/figure-35-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 35 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-J-6 (A); 7 (B), 8 (C), 11 (D), 12 (E). Full-size My DOI: 10.7717/peerj.11476/fig-35 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_035.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095610/figure-36-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 36 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-8 (A); YLSNHMO01218 (B); HYQ-A-19 (C); YLSNHMO01233 (D), 01229 (£). Full-size My DOI: 10.7717/peerj.11476/fig-36 YLSNHM01218 preserves two manus prints (5m and 6m) that show some " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_036.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095613/figure-37-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 37 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: YLSNHM01205 (A), 01202 (B), 01219 (C); HYQ-10 (D). Full-size MY DOI: 10.7717/peerj.11476/fig-37 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_037.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095616/figure-38-the-interpretive-outline-drawing-of-pterosaur-and"><img alt="Figure 38 The interpretive outline drawing of pterosaur and turtle tracks at Huangyangquan sites: HYQ-6 (A); YLSNHM01297 (B); HYQ-4 (C), 3 (D); YLSNHMO01266 (E); HYQ-2 (F), 1 (G). Full-size MY DOI: 10.7717/peerj.11476/fig-38 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_038.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095619/figure-39-lower-cretaceous-avian-dominated-theropod"><img alt="" class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_039.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095621/figure-40-lower-cretaceous-avian-dominated-theropod"><img alt="" class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_040.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095624/figure-41-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 41 Photograph (A) and interpretive outline drawing (B) of turtle tracks YLSNHM01206 at Huangyangquan sites. Full-size Mal DOI: 10.7717/peerj.11476/fig-41 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_041.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095628/figure-42-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 42 Photograph (A) and interpretive outline drawing (B) of turtle tracks HYQ-A-6 at Huangyangquan sites. Full-size Mal DOI: 10.7717/peerj.11476/fig-42 Morphotype A is best represented by HYQ-3 (Fig. 38). HYQ-3 is 5.3cm in length, with a length/width ratio of 0.9. HYQ-3 is typically digitigrade, and has four elongated ungual marks; the marks are roughly parallel and face anteriorly. The middle ungual marks (digits II and III) are longer than the lateral two (digits I and IV). The similar track, HYQ-A-11-2 (Fig. 39), is 1.2 cm in length, with a length/width ratio of 0.7, and only has three ungual marks - probably missing a relatively shallow impression of the lateral digit. The medial digit is the longest, while the lateral digits are subequal. The similar track assemblage YLSNHM01206 (Fig. 41) indicates a mean length/width ratio of 1.1, range from 0.3-1.9. three ungual marks — probably missing a relatively shallow impression of the lateral digit. Morphotype B is represented by HYQ-4 (Fig. 38), which is 8.9 cm in length, with a length/width ratio of 2.3. HYQ-4 is typically digitigrade, or semiplantigrade and has elongated ungual marks. Morphotype B most significantly differs from morphotype A in the higher length/width ratio and the number of digits. HYQ-4 has three ungual marks; " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_042.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095631/figure-43-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 43 Photograph (A) and interpretive outline drawing of theropod tracks As-TI1 (B), 2 (C), 3 (D) at Asphaltite sites. Full-size My DOI: 10.7717/peerj.11476/fig-43 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_043.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095635/table-1-comparison-of-the-type-material-of-the-monospecific"><img alt="Table 1 Comparison of the type material of the monospecific ichnogenus Chapus and the three ichnospecies of Asianopdus. “ Indicates the number of trackmakers, such as “2 (1)”, a set of pes and manus traces left by one trackmaker. (To), Total; (Is), Isolated; (Tw), Trackway; (Tm), Trackmaker; (Ra), Ratio. Numbers under Li, Jiang & Wang, 2020 refer to data from this paper; numbers under “Museum” refer to museum specimens. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/table_001.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095642/table-2-comparison-of-the-type-material-of-the-monospecific"><img alt="Table 2 Comparison of the type material of the monospecific ichnogenus Chapus and the three ichnospecies of Asianopdus. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/table_002.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095646/table-3-comparison-of-bone-records-and-track-records-of"><img alt="Table 3 Comparison of bone records and track records of Wuerhe area, Xinjiang, China. The Huangyangquan sites from the Lower Layer, and the Asphaltite site from the Upper Layer. Black squares indicate the existence of records. 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They originate from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, variously divided into four formations and subgroups in the northwestern margin of the Junggar Basin. The large Huangyangquan track assemblages occur in the Lower layer/Subgroup II. Similarly-composed track assemblages also occur at the smaller Asphaltite site in the Upper Layer/Subgroup III. The Huangyangquan assemblages have yielded more than 1,500 identified tracks including abundant tracks of avian and non-avian theropods, pterosaurs and turtles and less abundant tracks of stegosaurs. Previous avian track identifications have been reassessed to conclude that Moguiornipes robustus is a taphotaxon and Koreanaornis dodsoni might be better accommodated in the ichno...","publisher":"PeerJ","ai_title_tag":"Lower Cretaceous Track Assemblages in Xinjiang, China","publication_name":"PeerJ"},"translated_abstract":"Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe District, Karamay City, Xinjiang) have been an abundant source of some of the largest Lower Cretaceous track collections from China. They originate from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, variously divided into four formations and subgroups in the northwestern margin of the Junggar Basin. The large Huangyangquan track assemblages occur in the Lower layer/Subgroup II. Similarly-composed track assemblages also occur at the smaller Asphaltite site in the Upper Layer/Subgroup III. The Huangyangquan assemblages have yielded more than 1,500 identified tracks including abundant tracks of avian and non-avian theropods, pterosaurs and turtles and less abundant tracks of stegosaurs. Previous avian track identifications have been reassessed to conclude that Moguiornipes robustus is a taphotaxon and Koreanaornis dodsoni might be better accommodated in the ichno...","internal_url":"https://www.academia.edu/65564666/Lower_cretaceous_avian_dominated_theropod_thyreophoran_pterosaur_and_turtle_track_assemblages_from_the_Tugulu_Group_Xinjiang_China_ichnotaxonomy_and_palaeoecology","translated_internal_url":"","created_at":"2021-12-22T13:37:40.152-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":77104259,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/77104259/thumbnails/1.jpg","file_name":"11476.pdf","download_url":"https://www.academia.edu/attachments/77104259/download_file","bulk_download_file_name":"Lower_cretaceous_avian_dominated_theropo.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/77104259/11476-libre.pdf?1640210502=\u0026response-content-disposition=attachment%3B+filename%3DLower_cretaceous_avian_dominated_theropo.pdf\u0026Expires=1743195464\u0026Signature=DLLoVhUjvzKSt~4PJEXdT-YgSdK~psi-ENE-ZCslxUSKix9kI4Exe61Lz5gjCBFHW6OA2Rd3IRe-pUuRMuwJe94ZxjKNnBRtYl2Qp81Gzoru5snTxF-900-bqrdpiou1VlBGpPrx4K97JRLFEnHVxo5~VdhVTxfm8XQiQNGAIQJ1j2rCOeT~-dIBaXNPfkOUMARTkyeY7-0kvm3~~5NbXXJx5rcutEgWTkpf-gDJGXa6mg483iQtToTpaSIdOfX4OiSPs1sr--EvkHCijPIF5XxYT~Tw9cMKoMTY4C21K9I1sWbts1cwznaq2hnP0AoZ~Xnd1VXWE8EquLLiapiavg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Lower_cretaceous_avian_dominated_theropod_thyreophoran_pterosaur_and_turtle_track_assemblages_from_the_Tugulu_Group_Xinjiang_China_ichnotaxonomy_and_palaeoecology","translated_slug":"","page_count":73,"language":"en","content_type":"Work","summary":"Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe District, Karamay City, Xinjiang) have been an abundant source of some of the largest Lower Cretaceous track collections from China. They originate from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, variously divided into four formations and subgroups in the northwestern margin of the Junggar Basin. The large Huangyangquan track assemblages occur in the Lower layer/Subgroup II. Similarly-composed track assemblages also occur at the smaller Asphaltite site in the Upper Layer/Subgroup III. The Huangyangquan assemblages have yielded more than 1,500 identified tracks including abundant tracks of avian and non-avian theropods, pterosaurs and turtles and less abundant tracks of stegosaurs. Previous avian track identifications have been reassessed to conclude that Moguiornipes robustus is a taphotaxon and Koreanaornis dodsoni might be better accommodated in the ichno...","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[{"id":77104259,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/77104259/thumbnails/1.jpg","file_name":"11476.pdf","download_url":"https://www.academia.edu/attachments/77104259/download_file","bulk_download_file_name":"Lower_cretaceous_avian_dominated_theropo.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/77104259/11476-libre.pdf?1640210502=\u0026response-content-disposition=attachment%3B+filename%3DLower_cretaceous_avian_dominated_theropo.pdf\u0026Expires=1743195464\u0026Signature=DLLoVhUjvzKSt~4PJEXdT-YgSdK~psi-ENE-ZCslxUSKix9kI4Exe61Lz5gjCBFHW6OA2Rd3IRe-pUuRMuwJe94ZxjKNnBRtYl2Qp81Gzoru5snTxF-900-bqrdpiou1VlBGpPrx4K97JRLFEnHVxo5~VdhVTxfm8XQiQNGAIQJ1j2rCOeT~-dIBaXNPfkOUMARTkyeY7-0kvm3~~5NbXXJx5rcutEgWTkpf-gDJGXa6mg483iQtToTpaSIdOfX4OiSPs1sr--EvkHCijPIF5XxYT~Tw9cMKoMTY4C21K9I1sWbts1cwznaq2hnP0AoZ~Xnd1VXWE8EquLLiapiavg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"},{"id":77104261,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/77104261/thumbnails/1.jpg","file_name":"11476.pdf","download_url":"https://www.academia.edu/attachments/77104261/download_file","bulk_download_file_name":"Lower_cretaceous_avian_dominated_theropo.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/77104261/11476-libre.pdf?1640210502=\u0026response-content-disposition=attachment%3B+filename%3DLower_cretaceous_avian_dominated_theropo.pdf\u0026Expires=1743195464\u0026Signature=drcsrZnw1ZELMQ2ipaEPYBIOPaAq7MjtXe-47HbjNyUbtwWpLfPaSBZAD9sQ9wg1fCXcCg-NQRWJSUnbMj6-jiMHzgzkELdnR5hhlIxKfOXk6J~xF4y2anE12Tas46cVIlqyiHxOl6ldQivjxD-tJVhU4Im4gqaY9mNZVHTnu~0Ize712Y9HpRZarh6m4ldOThte2RHq8ojkFnBLwnzbQ31oRrsUjgg7W69MCzW6akr~hVr9RNitIzaQD~xah2cFIg5LT4IgkD1BhfJSZoS8K-jAylowtcW1bhN7EbuoaBVzkF9MukYutTjRxvounkDZ~KUDKgcSzlK0B2haCQZC3w__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[],"urls":[{"id":15552836,"url":"https://peerj.com/articles/11476.pdf"}]}, dispatcherData: dispatcherData }); 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While histor...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Fossil footprints provide the only evidence of Triassic-aged dinosaurs in Australia. While historically several Triassic dinosaur tracksites are reported, only one specimen exists for direct study. Evidence for the Rhondda colliery (Ipswich) trackmaker, Upper Triassic Blackstone Formation as a very large-bodied Triassic predatory dinosaur is not supported by the present study because of overestimated footprint length measurements, and we question earlier interpretations that these tracks are Eubrontes-like. Instead, we recognise shared characteristics with the sauropodomorph ichnogenus Evazoum. The track morphology resembles that of Evazoum on account of the distinct shape and position of the impression of the metatarsophalangeal pad for digit IV. The mesaxony of the Blackstone Formation track is also low, but possibly slightly higher than in Evazoum, if the proportions of the digital impressions are correctly interpreted. Finally, the trackway pattern shows constant track inward rotation, a feature observed in typical Evazoum and mostly absent in Eubrontes. Therefore, we assign these Australian tracks as Evazoum ichnosp. indet. If our assessment of the Blackstone Formation tracks is correct, they provide the first and only evidence of basal sauropodomorph dinosaurs from Australia. In addition to this record, we give an overview of other reported (but lost) Australian Triassic dinosaur footprints.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="8f8663ea3430136d6335274efb2f65f1" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":73844038,"asset_id":60365280,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/73844038/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="60365280"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="60365280"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 60365280; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=60365280]").text(description); $(".js-view-count[data-work-id=60365280]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 60365280; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='60365280']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "8f8663ea3430136d6335274efb2f65f1" } } $('.js-work-strip[data-work-id=60365280]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":60365280,"title":"Saurischian dinosaur tracks from the Upper Triassic of southern Queensland: possible evidence for Australia’s earliest sauropodomorph trackmaker","translated_title":"","metadata":{"publisher":"Informa UK Limited","grobid_abstract":"Fossil footprints provide the only evidence of Triassic-aged dinosaurs in Australia. 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Therefore, we assign these Australian tracks as Evazoum ichnosp. indet. If our assessment of the Blackstone Formation tracks is correct, they provide the first and only evidence of basal sauropodomorph dinosaurs from Australia. In addition to this record, we give an overview of other reported (but lost) Australian Triassic dinosaur footprints.","publication_name":"Historical Biology","grobid_abstract_attachment_id":73844039},"translated_abstract":null,"internal_url":"https://www.academia.edu/60365280/Saurischian_dinosaur_tracks_from_the_Upper_Triassic_of_southern_Queensland_possible_evidence_for_Australia_s_earliest_sauropodomorph_trackmaker","translated_internal_url":"","created_at":"2021-10-29T18:29:52.977-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":73844038,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/73844038/thumbnails/1.jpg","file_name":"08912963.2021.pdf","download_url":"https://www.academia.edu/attachments/73844038/download_file","bulk_download_file_name":"Saurischian_dinosaur_tracks_from_the_Upp.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/73844038/08912963.2021-libre.pdf?1635558942=\u0026response-content-disposition=attachment%3B+filename%3DSaurischian_dinosaur_tracks_from_the_Upp.pdf\u0026Expires=1743195464\u0026Signature=Dzn-KIFH8rOlO4eQ3u5vu0-axhFi9WJtppZY~QxhXaMc-Fj6CzqoBZaSrXsDM9M8c-CE3G1OCCeMHWdvPlfsQCWYPX9DI7be7hyhbI6mjv5LnzhyEB2lIt3DnYNcQML22aG~WCWKnV429K-HBD-oZLLXcnzI9TfbjON7mgGo-lr3WqFuFa9CGO5fMphfEHctBc0LXn-dioaM-PORB~r5ocjDjJr2UwFFR5oux7SdFbjlngTxJ3Mr6jy7E-NzgWL-Xbxx28j1HzA1Mat9Mj8eVwPoMv2wKUIFadC1WzaCvN5heBKiWfQg40UEs1nOcipd-5F4HemmRMgjX1IvJYIopg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Saurischian_dinosaur_tracks_from_the_Upper_Triassic_of_southern_Queensland_possible_evidence_for_Australia_s_earliest_sauropodomorph_trackmaker","translated_slug":"","page_count":11,"language":"en","content_type":"Work","summary":"Fossil footprints provide the only evidence of Triassic-aged dinosaurs in Australia. 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To date, a moderate diversity of six theropod ichnogenera has been reported, but none of these have been identified at the ichnospecies level. Thus, many morphotypes have common “generic” labels such as Grallator, Eubrontes, cf. Eubrontes or even “Eubrontes-Megalosauripus” morphotype. These morphotypes are generally more typical of the Jurassic, whereas other more distinctive theropod tracks (Minisauripus and Velociraptorichnus) are restricted to the Cretaceous. The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very well-preserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="7f6ceb987b81e668b15cc40aac58719a" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":72499007,"asset_id":57753746,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/72499007/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="57753746"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="57753746"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 57753746; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=57753746]").text(description); $(".js-view-count[data-work-id=57753746]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 57753746; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='57753746']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "7f6ceb987b81e668b15cc40aac58719a" } } $('.js-work-strip[data-work-id=57753746]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":57753746,"title":"The new ichnotaxon Eubrontes nobitai ichnosp. nov. and other saurischian tracks from the Lower Cretaceous of Sichuan Province and a review of Chinese Eubrontes-type tracks","translated_title":"","metadata":{"abstract":"The Jiaguan Formation and the underlying Feitianshan Formation (Lower Cretaceous) in Sichuan Province yield multiple saurischian (theropod–sauropod) dominated ichnofaunas. 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The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very well-preserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.","publisher":"Springer Science and Business Media LLC","ai_title_tag":"New Cretaceous Eubrontes nobitai from Sichuan","publication_name":"Journal of Palaeogeography"},"translated_abstract":"The Jiaguan Formation and the underlying Feitianshan Formation (Lower Cretaceous) in Sichuan Province yield multiple saurischian (theropod–sauropod) dominated ichnofaunas. To date, a moderate diversity of six theropod ichnogenera has been reported, but none of these have been identified at the ichnospecies level. Thus, many morphotypes have common “generic” labels such as Grallator, Eubrontes, cf. Eubrontes or even “Eubrontes-Megalosauripus” morphotype. These morphotypes are generally more typical of the Jurassic, whereas other more distinctive theropod tracks (Minisauripus and Velociraptorichnus) are restricted to the Cretaceous. The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very well-preserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.","internal_url":"https://www.academia.edu/57753746/The_new_ichnotaxon_Eubrontes_nobitai_ichnosp_nov_and_other_saurischian_tracks_from_the_Lower_Cretaceous_of_Sichuan_Province_and_a_review_of_Chinese_Eubrontes_type_tracks","translated_internal_url":"","created_at":"2021-10-14T02:57:49.746-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":72499007,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/72499007/thumbnails/1.jpg","file_name":"s42501-021-00096-y.pdf","download_url":"https://www.academia.edu/attachments/72499007/download_file","bulk_download_file_name":"The_new_ichnotaxon_Eubrontes_nobitai_ich.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/72499007/s42501-021-00096-y-libre.pdf?1634206350=\u0026response-content-disposition=attachment%3B+filename%3DThe_new_ichnotaxon_Eubrontes_nobitai_ich.pdf\u0026Expires=1743195464\u0026Signature=GYT9zF5lyHOIY7TodYYadD4kVkneNPPcK7TMXEd256lwOKRJG69Y1YAaAHajsVuWC0A065Mqbn5xJ7eeilrvcXyaICrnyPTCoB~Ao7gQ0fClCsqD1DiOSFqW3KTXpwVfzPSY8JHfBfJVXynW~F0zZO6V8MJEmFShxElUcqIyXBvY2H5YsH2X0iz8-2EBF3sbHGzyjv2E2EhBhJHGDVNJ0Hm3Q554dh7LaNjXP3V1ntWYWt77ZUF~fqklgyFgxbNKLfGfUqT-eHEJr-giMGcKyXr~lZJ9T3KHM0jDnI~qeqQuDuXvcaeaM5D5FkBoM8Ri3pUS2UeIgNHMzydJQm7oFA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"The_new_ichnotaxon_Eubrontes_nobitai_ichnosp_nov_and_other_saurischian_tracks_from_the_Lower_Cretaceous_of_Sichuan_Province_and_a_review_of_Chinese_Eubrontes_type_tracks","translated_slug":"","page_count":19,"language":"en","content_type":"Work","summary":"The Jiaguan Formation and the underlying Feitianshan Formation (Lower Cretaceous) in Sichuan Province yield multiple saurischian (theropod–sauropod) dominated ichnofaunas. To date, a moderate diversity of six theropod ichnogenera has been reported, but none of these have been identified at the ichnospecies level. Thus, many morphotypes have common “generic” labels such as Grallator, Eubrontes, cf. Eubrontes or even “Eubrontes-Megalosauripus” morphotype. These morphotypes are generally more typical of the Jurassic, whereas other more distinctive theropod tracks (Minisauripus and Velociraptorichnus) are restricted to the Cretaceous. The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very well-preserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[{"id":72499007,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/72499007/thumbnails/1.jpg","file_name":"s42501-021-00096-y.pdf","download_url":"https://www.academia.edu/attachments/72499007/download_file","bulk_download_file_name":"The_new_ichnotaxon_Eubrontes_nobitai_ich.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/72499007/s42501-021-00096-y-libre.pdf?1634206350=\u0026response-content-disposition=attachment%3B+filename%3DThe_new_ichnotaxon_Eubrontes_nobitai_ich.pdf\u0026Expires=1743195464\u0026Signature=GYT9zF5lyHOIY7TodYYadD4kVkneNPPcK7TMXEd256lwOKRJG69Y1YAaAHajsVuWC0A065Mqbn5xJ7eeilrvcXyaICrnyPTCoB~Ao7gQ0fClCsqD1DiOSFqW3KTXpwVfzPSY8JHfBfJVXynW~F0zZO6V8MJEmFShxElUcqIyXBvY2H5YsH2X0iz8-2EBF3sbHGzyjv2E2EhBhJHGDVNJ0Hm3Q554dh7LaNjXP3V1ntWYWt77ZUF~fqklgyFgxbNKLfGfUqT-eHEJr-giMGcKyXr~lZJ9T3KHM0jDnI~qeqQuDuXvcaeaM5D5FkBoM8Ri3pUS2UeIgNHMzydJQm7oFA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":289,"name":"Palaeogeography","url":"https://www.academia.edu/Documents/in/Palaeogeography"}],"urls":[{"id":12973464,"url":"https://link.springer.com/content/pdf/10.1186/s42501-021-00096-y.pdf"}]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-57753746-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="49372061"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/49372061/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea"><img alt="Research paper thumbnail of Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea" class="work-thumbnail" src="https://attachments.academia-assets.com/67745462/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/49372061/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea">Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea</a></div><div class="wp-workCard_item"><span>Scientific Reports</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation o...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation of South Korea, represent the well-known crocodylomorph ichnogenus Batrachopus. the Korean sample includes multiple, narrow-gauge, pes-only trackways with footprint lengths (FL) 18-24 cm, indicating trackmaker body lengths up to ~3.0 m. Surprisingly, the consistent absence of manus tracks in trackways, with well-preserved digital pad and skin traces, argues for bipedal trackmakers, here assigned to Batrachopus grandis ichnosp. nov. No definitive evidence, either from pes-onmanus overprinting or poor track preservation, suggests the trackways where made by quadrupeds that only appear bipedal. this interpretation helps solve previous confusion over interpretation of enigmatic tracks of bipeds from younger (? Albian) Haman formation sites by showing they are not pterosaurian as previously inferred. Rather, they support the strong consensus that pterosaurs were obligate quadrupeds, not bipeds. Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. Surprisingly the trackways never include manus imprints and therefore appear to indicate exclusively</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="7a1839d499c9aae9969f4ff422038635" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":67745462,"asset_id":49372061,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/67745462/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="49372061"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="49372061"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 49372061; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=49372061]").text(description); $(".js-view-count[data-work-id=49372061]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 49372061; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='49372061']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "7a1839d499c9aae9969f4ff422038635" } } $('.js-work-strip[data-work-id=49372061]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":49372061,"title":"Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea","translated_title":"","metadata":{"publisher":"Springer Science and Business Media LLC","ai_title_tag":"Bipedal Crocodylomorph Tracks from Cretaceous Korea","grobid_abstract":"Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation of South Korea, represent the well-known crocodylomorph ichnogenus Batrachopus. the Korean sample includes multiple, narrow-gauge, pes-only trackways with footprint lengths (FL) 18-24 cm, indicating trackmaker body lengths up to ~3.0 m. Surprisingly, the consistent absence of manus tracks in trackways, with well-preserved digital pad and skin traces, argues for bipedal trackmakers, here assigned to Batrachopus grandis ichnosp. nov. No definitive evidence, either from pes-onmanus overprinting or poor track preservation, suggests the trackways where made by quadrupeds that only appear bipedal. this interpretation helps solve previous confusion over interpretation of enigmatic tracks of bipeds from younger (? Albian) Haman formation sites by showing they are not pterosaurian as previously inferred. Rather, they support the strong consensus that pterosaurs were obligate quadrupeds, not bipeds. Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. Surprisingly the trackways never include manus imprints and therefore appear to indicate exclusively","publication_name":"Scientific Reports","grobid_abstract_attachment_id":67745463},"translated_abstract":null,"internal_url":"https://www.academia.edu/49372061/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea","translated_internal_url":"","created_at":"2021-06-25T15:09:57.067-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":67745462,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/67745462/thumbnails/1.jpg","file_name":"s41598-020-66008-7.pdf","download_url":"https://www.academia.edu/attachments/67745462/download_file","bulk_download_file_name":"Trackway_evidence_for_large_bipedal_croc.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/67745462/s41598-020-66008-7-libre.pdf?1624664665=\u0026response-content-disposition=attachment%3B+filename%3DTrackway_evidence_for_large_bipedal_croc.pdf\u0026Expires=1743195464\u0026Signature=L5fTCCNNG0NotDC3AB3fINC1CBXi9Xp8tsbn5TLYInGkqxz6HufeLbPmbs0egC~dSoDg2cTuggfLMcE8ZEAOgz7Dr06Pu79rAooHDvjueWNju6rUJkvYIyKw56LfGHWnfmCTt2zNNI0o7xq2Mkp7eGsMamFsbmB692ABmlO5sMtR9gp8O4GZj92QtdVZxA-qyWwB6i2OKSou9Lhyj4tpwyrMwkbSGOQ0xM0hqhmeGQrjlxNwyIdu4SKoMZLv5XFP6~VzbPdD4AY2ebDMpqyYd3QUbkf1rGkWy-eIaMWy9RxdG84HIX6KS8l8RWuirIi8pQcbNFTh0Eit1WJPjmg3Hg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea","translated_slug":"","page_count":13,"language":"en","content_type":"Work","summary":"Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation of South Korea, represent the well-known crocodylomorph ichnogenus Batrachopus. the Korean sample includes multiple, narrow-gauge, pes-only trackways with footprint lengths (FL) 18-24 cm, indicating trackmaker body lengths up to ~3.0 m. Surprisingly, the consistent absence of manus tracks in trackways, with well-preserved digital pad and skin traces, argues for bipedal trackmakers, here assigned to Batrachopus grandis ichnosp. nov. No definitive evidence, either from pes-onmanus overprinting or poor track preservation, suggests the trackways where made by quadrupeds that only appear bipedal. this interpretation helps solve previous confusion over interpretation of enigmatic tracks of bipeds from younger (? Albian) Haman formation sites by showing they are not pterosaurian as previously inferred. Rather, they support the strong consensus that pterosaurs were obligate quadrupeds, not bipeds. Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. Surprisingly the trackways never include manus imprints and therefore appear to indicate exclusively","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[{"id":67745462,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/67745462/thumbnails/1.jpg","file_name":"s41598-020-66008-7.pdf","download_url":"https://www.academia.edu/attachments/67745462/download_file","bulk_download_file_name":"Trackway_evidence_for_large_bipedal_croc.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/67745462/s41598-020-66008-7-libre.pdf?1624664665=\u0026response-content-disposition=attachment%3B+filename%3DTrackway_evidence_for_large_bipedal_croc.pdf\u0026Expires=1743195464\u0026Signature=L5fTCCNNG0NotDC3AB3fINC1CBXi9Xp8tsbn5TLYInGkqxz6HufeLbPmbs0egC~dSoDg2cTuggfLMcE8ZEAOgz7Dr06Pu79rAooHDvjueWNju6rUJkvYIyKw56LfGHWnfmCTt2zNNI0o7xq2Mkp7eGsMamFsbmB692ABmlO5sMtR9gp8O4GZj92QtdVZxA-qyWwB6i2OKSou9Lhyj4tpwyrMwkbSGOQ0xM0hqhmeGQrjlxNwyIdu4SKoMZLv5XFP6~VzbPdD4AY2ebDMpqyYd3QUbkf1rGkWy-eIaMWy9RxdG84HIX6KS8l8RWuirIi8pQcbNFTh0Eit1WJPjmg3Hg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"},{"id":67745463,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/67745463/thumbnails/1.jpg","file_name":"s41598-020-66008-7.pdf","download_url":"https://www.academia.edu/attachments/67745463/download_file","bulk_download_file_name":"Trackway_evidence_for_large_bipedal_croc.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/67745463/s41598-020-66008-7-libre.pdf?1624664664=\u0026response-content-disposition=attachment%3B+filename%3DTrackway_evidence_for_large_bipedal_croc.pdf\u0026Expires=1743195464\u0026Signature=YhU9iYWcaLxuyI0LyUNuSSGY~rK4XCOIv9lKEp-TrwYfI1ZvSZ1~qNFX0mWiDT35J6Nv8GWKu58mODa~zFtbIeeMCRj5wU8SN0KPj-U2CW7ThcaNqsL9~gr-SjTNk7WTmJiOtHhyPVJHvwu-jpvJn6WFskRDHwNhi4LE75gRmIQro29ArUR7tQzw2wI8oYIU11TZWmsg6-WkRFQOPI1FHQAV6iLuQH9FaTk~pvL-E8A-~GdLp59JWODghYA9OxMUsUo4m4UlpArJhxjh7uaz8kKVT09Ikf9GvcxmOM9bOZq25fFioIn8PANPvEZbCJ4Vzu2eaghGnJQwZtMQ4ZUUNw__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[],"urls":[{"id":10298754,"url":"http://www.nature.com/articles/s41598-020-66008-7.pdf"}]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-49372061-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="49372060"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/49372060/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea"><img alt="Research paper thumbnail of Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea" class="work-thumbnail" src="https://attachments.academia-assets.com/67745459/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/49372060/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea">Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea</a></div><div class="wp-workCard_item"><span>Scientific Reports</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation o...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation of South Korea, represent the well-known crocodylomorph ichnogenus Batrachopus. the Korean sample includes multiple, narrow-gauge, pes-only trackways with footprint lengths (FL) 18-24 cm, indicating trackmaker body lengths up to ~3.0 m. Surprisingly, the consistent absence of manus tracks in trackways, with well-preserved digital pad and skin traces, argues for bipedal trackmakers, here assigned to Batrachopus grandis ichnosp. nov. No definitive evidence, either from pes-onmanus overprinting or poor track preservation, suggests the trackways where made by quadrupeds that only appear bipedal. this interpretation helps solve previous confusion over interpretation of enigmatic tracks of bipeds from younger (? Albian) Haman formation sites by showing they are not pterosaurian as previously inferred. Rather, they support the strong consensus that pterosaurs were obligate quadrupeds, not bipeds. Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. Surprisingly the trackways never include manus imprints and therefore appear to indicate exclusively</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="1d0a928b94c38bf61a452e87d80ae5a5" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":67745459,"asset_id":49372060,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/67745459/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="49372060"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="49372060"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 49372060; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=49372060]").text(description); $(".js-view-count[data-work-id=49372060]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 49372060; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='49372060']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "1d0a928b94c38bf61a452e87d80ae5a5" } } $('.js-work-strip[data-work-id=49372060]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":49372060,"title":"Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea","translated_title":"","metadata":{"publisher":"Springer Science and Business Media LLC","ai_title_tag":"Bipedal Crocodylomorph Trackways from Lower Cretaceous Korea","grobid_abstract":"Large well-preserved crocodylomorph tracks from the Lower cretaceous (? 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Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. 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Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. 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The seven dinosaurian eggs reported clearly belong to the Elongatoolithidae and burrow traces were most likely made by small crustaceans. This association prompts the question as to whether invertebrate activity had relations with the buried eggs. This may be just an occasional case or the eggs may have organically increased the content of organic matter in soil which attracted the crustaceans.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="30020f316cb135ea5539b83f4704bfb9" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":66381013,"asset_id":47118777,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/66381013/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="47118777"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="47118777"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 47118777; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=47118777]").text(description); $(".js-view-count[data-work-id=47118777]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 47118777; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='47118777']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "30020f316cb135ea5539b83f4704bfb9" } } $('.js-work-strip[data-work-id=47118777]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":47118777,"title":"Dinosaur Eggs Associated with Crustacean Trace Fossils from the Upper Cretaceous of Jiangxi, China: Evidence for Foraging Behavior?","translated_title":"","metadata":{"abstract":"We report the discovery of concentrated invertebrate inchnofossils in close association with a dinosaur nest from the Hekou Formation in Jiangxi Province, China. 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This may be just an occasional case or the eggs may have organically increased the content of organic matter in soil which attracted the crustaceans.","internal_url":"https://www.academia.edu/47118777/Dinosaur_Eggs_Associated_with_Crustacean_Trace_Fossils_from_the_Upper_Cretaceous_of_Jiangxi_China_Evidence_for_Foraging_Behavior","translated_internal_url":"","created_at":"2021-04-20T21:33:48.521-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":66381013,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/66381013/thumbnails/1.jpg","file_name":"128.pdf","download_url":"https://www.academia.edu/attachments/66381013/download_file","bulk_download_file_name":"Dinosaur_Eggs_Associated_with_Crustacean.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/66381013/128-libre.pdf?1618981370=\u0026response-content-disposition=attachment%3B+filename%3DDinosaur_Eggs_Associated_with_Crustacean.pdf\u0026Expires=1743195464\u0026Signature=JvI-lCvsPp~t3DRsPpng22xEbw4cfCkKN3VgNblwZZHt4l~zsHNFtYW5xmmkWswxQVkzaJcqAF6M2vFPvKqcPu1Xsv7pks2iAKgDDS2rVf42Q04TAyM-cIXhnxM2px42~zqLkVDi8DC0XKZ~EWMdEJDUNigJMbt9j5J6Mj8fEeSLLzS5-R2yz4ztaaR1oeRN5OFddaG8PbjFlqhOOmjpWlGz4L18iGhol-fh-6tVHjxrhP0w1Me-JxjZ4nsFZ-xNXY7xl4XT8dtguL7SWVpLmNMpNvCSS1xg~RMJc8bNX34iwxO4UWgnMra78npR-Qm0tUYUDLO6aE7f43INPAM1JQ__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Dinosaur_Eggs_Associated_with_Crustacean_Trace_Fossils_from_the_Upper_Cretaceous_of_Jiangxi_China_Evidence_for_Foraging_Behavior","translated_slug":"","page_count":6,"language":"en","content_type":"Work","summary":"We report the discovery of concentrated invertebrate inchnofossils in close association with a dinosaur nest from the Hekou Formation in Jiangxi Province, China. 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-126000529-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="123167319"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/123167319/TRACK_RECOGNITION_VIA_ARTIFICIAL_COGNITION_TRAC_PRELIMINARY_REPORT_ON_THE_APPLICATION_OF_MACHINE_LEARNING_TO_IDENITFY_DINOSAUR_TRACKS"><img alt="Research paper thumbnail of TRACK RECOGNITION VIA ARTIFICIAL COGNITION (TRAC): PRELIMINARY REPORT ON THE APPLICATION OF MACHINE LEARNING TO IDENITFY DINOSAUR TRACKS" class="work-thumbnail" src="https://attachments.academia-assets.com/117665830/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/123167319/TRACK_RECOGNITION_VIA_ARTIFICIAL_COGNITION_TRAC_PRELIMINARY_REPORT_ON_THE_APPLICATION_OF_MACHINE_LEARNING_TO_IDENITFY_DINOSAUR_TRACKS">TRACK RECOGNITION VIA ARTIFICIAL COGNITION (TRAC): PRELIMINARY REPORT ON THE APPLICATION OF MACHINE LEARNING TO IDENITFY DINOSAUR TRACKS</a></div><div class="wp-workCard_item wp-workCard--coauthors"><span>by </span><span><a class="" data-click-track="profile-work-strip-authors" href="https://dinotrax.academia.edu/AMilner">Andrew Milner</a>, <a class="" data-click-track="profile-work-strip-authors" href="https://uq.academia.edu/AnthonyRomilio">Anthony Romilio</a>, and <a class="" data-click-track="profile-work-strip-authors" href="https://independent.academia.edu/HouckKaren">Karen Houck</a></span></div><div class="wp-workCard_item"><span>New Mexico Museum of Natural History and Science Bulletin</span><span>, 2024</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Despite the utilization of new technologies to objectively record and document dinosaur tracks in...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Despite the utilization of new technologies to objectively record and document dinosaur tracks in the field, the categorization and identification of dinosaur tracks remains a largely subjective process. Track Recognition via Artificial Cognition (TRAC) is a project that seeks to provide an objective method for categorizing and identifying dinosaur tracks using track surface photographs. TRAC will employ a combination of different iterations of supervised and unsupervised deep machine learning (ML) approaches to categorize track images drawing from a data set of over 1,000 digital photographs of dinosaur footprint images from around the world. Iterations and adjustments to the ML algorithms will identify natural clusters in the image data set, and our ultimate goal is to build a model capable of assigning an ichnotaxonomic identification to new images that TRAC analyzes. If successful, TRAC can be deployed as an application that can identify dinosaur footprints to ichnotaxon for use by professional paleontologists, citizen scientists, and land managers, with the potential of expanding the scope of the project to include more types of fossils. The TRAC Project is divided into two phases. This paper presents a preliminary report on Phase I, the preparation of the image data set.</span></div><div class="wp-workCard_item"><div class="carousel-container carousel-container--sm" id="profile-work-123167319-figures"><div class="prev-slide-container js-prev-button-container"><button aria-label="Previous" class="carousel-navigation-button js-profile-work-123167319-figures-prev"><span class="material-symbols-outlined" style="font-size: 24px" translate="no">arrow_back_ios</span></button></div><div class="slides-container js-slides-container"><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/6289364/figure-1-examples-of-dinosaur-footprint-images-in-the-trac"><img alt="FIGURE 1. A-M, Examples of dinosaur footprint images in the TRAC image dataset. Locality and ichnotaxonomic data are provided in Table 1. Specimens F and M are replicas of the originals. " class="figure-slide-image" src="https://figures.academia-assets.com/117665830/figure_001.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/6289374/figure-2-track-recognition-via-artificial-cognition-trac"><img alt="" class="figure-slide-image" src="https://figures.academia-assets.com/117665830/figure_002.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/6289382/figure-3-during-the-preliminary-supervised-learning-phase-he"><img alt="FIGURE 3. During the preliminary supervised learning phase, he location of the track on the image should be tagged using an application such as CVAT.ai. Additional image features, such as scale bars, can also be tagged so they are not confused with he track analysis but can be used to provide track dimensions. A, theropod track preserved in the Lower Jurassic Navajo Sandstone near Moab, Utah (Lockley et al., 2021), scale 25 cm. During the tagging process a length may be added to the scale ag. B, orthoimage map of an ankylosaurian track preserved on he underside of a Lower Cretaceous Plainview Sandstone bed near Canon City, Colorado (Kurtz et. al., 2001), scale 22.6 cm. " class="figure-slide-image" src="https://figures.academia-assets.com/117665830/figure_003.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/6289394/table-1-dinosaur-track-examples-pictured-in-with"><img alt="TABLE 1. Dinosaur track examples pictured in Figure | with ichnotaxonomic identification and general locality data. " class="figure-slide-image" src="https://figures.academia-assets.com/117665830/table_001.jpg" /></a></figure></div><div class="next-slide-container js-next-button-container"><button aria-label="Next" class="carousel-navigation-button js-profile-work-123167319-figures-next"><span class="material-symbols-outlined" style="font-size: 24px" translate="no">arrow_forward_ios</span></button></div></div></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="de01bb92cb255a4c7e4e29b3796f8097" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":117665830,"asset_id":123167319,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/117665830/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="123167319"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="123167319"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 123167319; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=123167319]").text(description); $(".js-view-count[data-work-id=123167319]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 123167319; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='123167319']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "de01bb92cb255a4c7e4e29b3796f8097" } } $('.js-work-strip[data-work-id=123167319]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":123167319,"title":"TRACK RECOGNITION VIA ARTIFICIAL COGNITION (TRAC): PRELIMINARY REPORT ON THE APPLICATION OF MACHINE LEARNING TO IDENITFY DINOSAUR TRACKS","translated_title":"","metadata":{"abstract":"Despite the utilization of new technologies to objectively record and document dinosaur tracks in the field, the categorization and identification of dinosaur tracks remains a largely subjective process. 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the utilization of new technologies to objectively record and document dinosaur tracks in the field, the categorization and identification of dinosaur tracks remains a largely subjective process. Track Recognition via Artificial Cognition (TRAC) is a project that seeks to provide an objective method for categorizing and identifying dinosaur tracks using track surface photographs. TRAC will employ a combination of different iterations of supervised and unsupervised deep machine learning (ML) approaches to categorize track images drawing from a data set of over 1,000 digital photographs of dinosaur footprint images from around the world. Iterations and adjustments to the ML algorithms will identify natural clusters in the image data set, and our ultimate goal is to build a model capable of assigning an ichnotaxonomic identification to new images that TRAC analyzes. If successful, TRAC can be deployed as an application that can identify dinosaur footprints to ichnotaxon for use by professional paleontologists, citizen scientists, and land managers, with the potential of expanding the scope of the project to include more types of fossils. 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Represented as a near-complete right hind limb, key aspects of the palaeobiology of Rhoetosaurus remain unresolved. The current study combines three-dimensional modelling and photogrammetry to reconstruct the range of motion (ROM) of the foot of Rhoetosaurus to elicit pedal flexibility and posture. Our findings show that Rhoetosaurus had a high degree of digit mobility at all the metatarsophalangeal joints (total excursion angle &amp;gt;80°) and in the sagittal, transverse and frontal planes at the distal-most interphalangeal joints (excursion angle &amp;gt;100°). The position of the autopodial articular surfaces suggests that the metarsophalangeal joints were permanently elevated in life, an indicator of skeletal digitigrady. This joint may have been ventrally connected with the plantar aspect of the foot by a compliant fibrous pad, consistent with sauropod track data, and therefore likely to have been functional plantigrade, as has been previously suggested. These results provide new insights into the simulated ROM and posture of the hind foot of Rhoetosaurus; we recognize that additional parameters (including loading regimes and ichnological data) provide future modelling aspects to further our understanding of Rhoetosaurus pedal biomechanics.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="65567150"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="65567150"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 65567150; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=65567150]").text(description); $(".js-view-count[data-work-id=65567150]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 65567150; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='65567150']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=65567150]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":65567150,"title":"Reconstruction of the motion and hindfoot posture of Rhoetosaurus brownei Longman, 1926 (Sauropoda, Gravisauria)","translated_title":"","metadata":{"abstract":"The Middle Jurassic (Late Bathonian–Middle Callovian) basal sauropod Rhoetosaurus brownei is the largest known pre-Cretaceous terrestrial Australian vertebrate. 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These results provide new insights into the simulated ROM and posture of the hind foot of Rhoetosaurus; we recognize that additional parameters (including loading regimes and ichnological data) provide future modelling aspects to further our understanding of Rhoetosaurus pedal biomechanics.","internal_url":"https://www.academia.edu/65567150/Reconstruction_of_the_motion_and_hindfoot_posture_of_Rhoetosaurus_brownei_Longman_1926_Sauropoda_Gravisauria_","translated_internal_url":"","created_at":"2021-12-22T14:00:08.439-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[],"slug":"Reconstruction_of_the_motion_and_hindfoot_posture_of_Rhoetosaurus_brownei_Longman_1926_Sauropoda_Gravisauria_","translated_slug":"","page_count":null,"language":"en","content_type":"Work","summary":"The Middle Jurassic (Late Bathonian–Middle Callovian) basal sauropod Rhoetosaurus brownei is the largest known pre-Cretaceous terrestrial Australian vertebrate. 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-65567149-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="65567147"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/65567147/Dinosaur_stampede_stopped_in_its_tracks"><img alt="Research paper thumbnail of Dinosaur stampede stopped in its tracks" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title">Dinosaur stampede stopped in its tracks</div><div class="wp-workCard_item"><span>Australasian science</span><span>, 2015</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">A forensic analysis of ancient footprints has cast doubt on claims that a dinosaur stampede took ...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">A forensic analysis of ancient footprints has cast doubt on claims that a dinosaur stampede took place at Lark Quarry.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="65567147"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="65567147"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 65567147; 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-65567147-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="65567146"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/65567146/Dinosaur_tracks_from_the_Walmadany_area_James_Price_Point_Dampier_Peninsula_Western_Australia"><img alt="Research paper thumbnail of Dinosaur tracks from the Walmadany area (James Price Point), Dampier Peninsula, Western Australia" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title">Dinosaur tracks from the Walmadany area (James Price Point), Dampier Peninsula, Western Australia</div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">The Early Cretaceous (Valanginian–Barremian) Broome Sandstone of the Dampier Peninsula, Western A...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">The Early Cretaceous (Valanginian–Barremian) Broome Sandstone of the Dampier Peninsula, Western Australia, has often been touted as having one of the most diverse dinosaurian ichnofaunas in the world. Yet with the exception of tracks assigned to the theropodan ichnotaxon Megalosauropus broomensis in 1967, and more recent work on the preservational characteristics of sauropodan tracks, descriptive accounts of the types of dinosaurian tracks that occur in the Broome Sandstone have been limited, with reports ranging from 6–16 different ichnotaxa. As such, the overall scientific significance of the ichnofauna has remained enigmatic. At the request of Goolarabooloo Traditional Custodians, 300+ hours of ichnological survey work was undertaken on the 25 km stretch of coastline in the vicinity of Walmadany (James Price Point) from 2011–2013. 48 discrete tracksites were identified in this area, containing upward of 20 track types referable to theropods, sauropods, ornithopods and thyreophora...</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="65567146"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="65567146"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 65567146; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=65567146]").text(description); $(".js-view-count[data-work-id=65567146]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 65567146; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='65567146']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=65567146]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":65567146,"title":"Dinosaur tracks from the Walmadany area (James Price Point), Dampier Peninsula, Western Australia","translated_title":"","metadata":{"abstract":"The Early Cretaceous (Valanginian–Barremian) Broome Sandstone of the Dampier Peninsula, Western Australia, has often been touted as having one of the most diverse dinosaurian ichnofaunas in the world. 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Here, we describe a diminutive tridactyl track (10.2 mm in length) from the Lower Jura...","internal_url":"https://www.academia.edu/65567145/The_smallest_non_avian_dinosaur_track_in_China_Lower_Jurassic_Sichuan_Province_","translated_internal_url":"","created_at":"2021-12-22T14:00:08.004-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[],"slug":"The_smallest_non_avian_dinosaur_track_in_China_Lower_Jurassic_Sichuan_Province_","translated_slug":"","page_count":null,"language":"en","content_type":"Work","summary":"The smallest non-avian theropods have often attracted attention of palaeobiological and ichnological research. 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-65567144-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="65564666"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/65564666/Lower_cretaceous_avian_dominated_theropod_thyreophoran_pterosaur_and_turtle_track_assemblages_from_the_Tugulu_Group_Xinjiang_China_ichnotaxonomy_and_palaeoecology"><img alt="Research paper thumbnail of Lower cretaceous avian-dominated, theropod, thyreophoran, pterosaur and turtle track assemblages from the Tugulu Group, Xinjiang, China: ichnotaxonomy and palaeoecology" class="work-thumbnail" src="https://attachments.academia-assets.com/77104259/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/65564666/Lower_cretaceous_avian_dominated_theropod_thyreophoran_pterosaur_and_turtle_track_assemblages_from_the_Tugulu_Group_Xinjiang_China_ichnotaxonomy_and_palaeoecology">Lower cretaceous avian-dominated, theropod, thyreophoran, pterosaur and turtle track assemblages from the Tugulu Group, Xinjiang, China: ichnotaxonomy and palaeoecology</a></div><div class="wp-workCard_item"><span>PeerJ</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe District, Karamay City, Xinjiang) have been an abundant source of some of the largest Lower Cretaceous track collections from China. They originate from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, variously divided into four formations and subgroups in the northwestern margin of the Junggar Basin. The large Huangyangquan track assemblages occur in the Lower layer/Subgroup II. Similarly-composed track assemblages also occur at the smaller Asphaltite site in the Upper Layer/Subgroup III. The Huangyangquan assemblages have yielded more than 1,500 identified tracks including abundant tracks of avian and non-avian theropods, pterosaurs and turtles and less abundant tracks of stegosaurs. Previous avian track identifications have been reassessed to conclude that Moguiornipes robustus is a taphotaxon and Koreanaornis dodsoni might be better accommodated in the ichno...</span></div><div class="wp-workCard_item"><div class="carousel-container carousel-container--sm" id="profile-work-65564666-figures"><div class="prev-slide-container js-prev-button-container"><button aria-label="Previous" class="carousel-navigation-button js-profile-work-65564666-figures-prev"><span class="material-symbols-outlined" style="font-size: 24px" translate="no">arrow_back_ios</span></button></div><div class="slides-container js-slides-container"><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095424/figure-1-location-of-the-huangyangquan-track-site-in"><img alt="Figure 1 Location of the Huangyangquan track site in Xinjiang, China. (A) East Asia region; (B) Wuerhe area; (C) Huangyangquan sites (A-K). This Huangyangquan area is currently protected by the local reservoir management department. There are isolation fences around the reservoir, which protects the footprint to a certain extent. Full-size Mal DOI: 10.7717/peerj.11476/fig-1 GEOLOGICAL SETTING Tugulu group—depositional environment, stratigraphy and age " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_001.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095426/figure-2-stratigraphic-section-of-the-lower-layer-of-the"><img alt="Figure 2 Stratigraphic section of the Lower Layer of the Tugulu Group at the Wuerhe area and Huangyangquan track sites (A, C, E, F, G, I and K) (emended from Dong, 1973a). Full-size Mal DOI: 10.7717/peerj.11476/fig-2 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_002.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095429/figure-3-invertebrate-traces-from-the-huangyangquan-area"><img alt="Figure 3 Invertebrate traces from the Huangyangquan area. Scoyenia gracilis (=Sc) (A, B), Cochlichnus anguineus (C), Helminthoidichnites tenuis (=Ht) (D, E, F), Mermia carickensis (=Me) (F, G), Cruziana problematica (=Cr) (F, G), Rusophycus isp. (=Ru) (F, G), Treptichnus isp. (=Tr) (H), Planolites montanus (=P1) (H). Scale bar in (A), (B), (F), (G) and (H) = 10 cm; ruler in (C), (D), (E) scale in centimeter. Full-size Mw DOI: 10.7717/peerj.11476/fig-3 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_003.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095433/figure-4-invertebrate-traces-from-the-huangyangquan-area"><img alt="Figure 4 Invertebrate traces from the Huangyangquan area. Lockeia siliquaria (=Lo) (A, C), Archaeonassa fossulata (=Ar) (B) and Helminthopsis abeli (=He) (D). Scale bar = 10 cm Full-size ha DOI: 10.7717/peerj.11476/fig-4 Huangyangquan area " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_004.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095437/figure-5-the-interpretive-outline-drawing-of-theropod-tracks"><img alt="Figure 5 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-A-3 (A), 16 (B), 17 (C), 18 (D), 20 (E), 23 (F), 24 (G), 21 (H); HYQ-D-5 (I), 1 (J), 2 (K), 6 (L), 12 (M), (S), 3 (T), and HYQ-A-15 (U). 7 (N), 8 (O), 11 (P) 9 (Q); HYQ-L-1 (R), 2 Full-size wr" DOI: 10.7717/peerj.11476/fig-5 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_005.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095439/figure-6-the-interpretive-outline-drawing-of-theropod-tracks"><img alt="Figure 6 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-D-3 (A), 16 (B), 4 (C), 10 (D); HYQ-E-6 (E), 10 (F), 9 (G). Full-size ns DOI: 10.7717/peerj.11476/fig-6 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_006.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095444/figure-7-lower-cretaceous-avian-dominated-theropod"><img alt="" class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_007.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095445/figure-8-the-interpretive-outline-drawing-of-theropod-tracks"><img alt="Figure 8 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-F-13 (A), 14 (B), 15 (C), 16 (D); HYQ-H-3 (E); HYQ-G-1 (F); HYQ-H-1 (G); HYQ-J-1 (H); HYQ-K-1 (I), 2 9). Full-size ns DOI: 10.7717/peerj.11476/fig-8 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_008.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095448/figure-9-the-interpretive-outline-drawing-of-theropod-tracks"><img alt="Figure 9 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-H-4 (A), 2 (B). Full-size ns DOI: 10.7717/peerj.11476/fig-9 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_009.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095452/figure-10-perspective-of-hyq-left-pedal-impression-dorsal"><img alt="Figure 10 Perspective of HYQ-E-11 (left pedal impression). (A) Dorsal view; (B) Ventral view; (C) Anterior view; (D) Posterior view; (E) Medial view; and (F) Lateral view. The track surface images are shown as orthophotographs, ambient occlusion images, and elevation images. Full-size | DOI: 10.7717/peerj.11476/fig-10 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_010.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095457/figure-11-the-interpretive-outline-drawing-of-theropod"><img alt="Figure 11 The interpretive outline drawing of theropod tracks at Huangyangquan sites: HYQ-D-13. Full-size Mal DOI: 10.7717/peerj.11476/fig-11 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_011.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095462/figure-12-jialingpus-reflects-small-to-medium-sized-theropod"><img alt="Jialingpus reflects a small to medium-sized theropod trackmaker which registered grallatorid tracks akin to Grallator (sensu lato). However, a distinctive, centrally placed, swollen metatarsophalangeal area is a distinctive and diagnostic feature of well- preserved Jialingpus, making it different from Eubrontes—Grallator-Anchisauripus plexus tracks. In the best-preserved specimens of Jialingpus, a crease separates the small metatarsophalangeal pad region behind digit II and the larger region behind digit IV. This trait is clearly present in the theropod morphotype A tracks from the Huangyangquan site, which can, therefore, be assigned to Jialingpus. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_012.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095468/figure-13-the-interpretive-outline-drawing-of-theropod"><img alt="Figure 13 The interpretive outline drawing of theropod tracks at Huangyangquan sites. Specimens numbers: YLSNHM01243 (A); HYQ-13 (B); YLSNHM00978 (C), 00980 (D), 01236 (E), 00977 (E), 00979 (G), 01225 (H), 01237 (I), 01223 (J), 01226 (K), 01247 (L), 01268 (M), 01250 (N), 01269 (O), 01270 (P), 01271 (Q), 01272 (R), 01273 (S), 01274 (T), 01275 (U), 01276 (V), 01277 (W), 01279 (X), 01280 (Y), 01278 (Z). Full-size My DOI: 10.7717/peerj.11476/fig- 13 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_013.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095478/figure-14-the-interpretive-outline-drawing-of-theropod-and"><img alt="Figure 14 The interpretive outline drawing of theropod and bird tracks at Huangyangquan sites: YLSNHMO01246 (A); HYQ-21 (B); YLSNHMO01244 (C); HYQ-15 (D), 17 (E), 19 (F), 16 (G), 14 (H), 9 (1); YLSNHMO012353 (J). Full-size My DOI: 10.7717/peerj.11476/fig-14 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_014.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095488/figure-15-the-interpretive-outline-drawing-of-theropod-and"><img alt="Figure 15 The interpretive outline drawing of theropod and bird tracks at Huangyangquan sites: HYQ-E-1 (A), 2 (B), 3 (C)} HYQ-K-3 (D); HYQ-J-9 (E), 10 (F). Full-size Ry DOI: 10.7717/peerj.11476/fig-15 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_015.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095493/figure-16-asianopudus-niui-comparisons-between-holotypes-of"><img alt="Figure 16 Asianopudus niui. Comparisons between holotypes of Asianopudus and Chapus from China. (A) A. niui, (B) C. lockleyi and (C) A. robustus, showing anterior triangle and ratio of digit III/ Footprint length. Note similarities between A. niui and C. lockleyi. See text for details. Full-size My DOI: 10.7717/peerj.11476/fig-16 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_016.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095501/figure-17-photograph-of-theropod-track-ylsnhm-and-hyq-at"><img alt="Figure 17 Photograph of theropod track YLSNHM01265 and HYQ-A-18 at Huangyangquan site: Dorsal (A), Anterior (B), Lateral (C) and Medial (D) views of YLSNHMO01265; Dorsal (E) view of HYQ-A-18. The black arrow indicate the scale scratch lines along the side. r..11 2: lal TAAAT. 1A 9910 sew eet DIAM CLR +I " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_017.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095508/figure-18-photograph-of-theropod-track-hyq-and-at"><img alt="Figure 18 Photograph of theropod track HYQ-K-1 (A) and 2 (B) at Huangyangquan site. Full-size RY DOI: 10.7717/peerj.11476/fig-1 for establishing a new ichnospecies. First, Belvedere & Farlow (2016) and Marchetti et al. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_018.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095513/figure-19-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 19 Photograph (A) and interpretive outline drawing (B) of bird tracks at Huangyangquan site E: HYQ-E-2. Full-size MY DOI: 10.7717/peerj.11476/fig-19 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_019.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095521/figure-20-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 20 Photograph (A) and interpretive outline drawing (B) of bird tracks YLSNHMO01231, and the photograph (C) of bird tracks YLSNHM 01295 at Huangyangquan sites. Full-size My DOI: 10.7717/peerj.11476/fig-20 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_020.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095527/figure-21-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 21 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01203 (A), 01210 (B), 01213 (C), 01212 (D), 01234 (E). Full-size Kal DOI: 10.7717/peerj.11476/fig-21 Comparison & discussion " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_021.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095535/figure-22-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 22 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01214 (A), 01224 (B), 01215 (C), 01216 (D), 01217 (E), 01230 (F), 01245 (G). Full-size Mal DOI: 10.7717/peerj.11476/fig-22 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_022.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095542/figure-23-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 23 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01227 (A), 01238 (B), 01248 (C), 01232 (D), 01251 (E), 01254 (EF). Full-size RY DOI: 10.7717/peerj.11476/fig-23 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_023.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095548/figure-24-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 24 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01255 (A), 01281 (B), 01284 (C), 01286 (D), 01287 (E), 01283 (E), 01285 (G), 01288 (H), 01289 (I), 01290 (J), 01291 (K), 01292 (L), 01293 (M), 01294 (N). Full-size My DOI: 10.7717/peerj.11476/fig-24 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_024.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095552/figure-25-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 25 The interpretive outline drawing of bird tracks at Huangyangquan sites: YLSNHM01201 (A); HYQ-12 (B); YLSNHMO01201 (C), 01242 (D). Full-size My DOI: 10.7717/peerj.11476/fig-25 Koreanornipodidae includes the small subsymmetric functionally-tridactyl tracks ‘oreanaornis (Kim, 1969; Lockley et al., 1992) and Pullornipes (Lockley et al., 2006b). he latter differs from Koreanaornis mostly in having a small postero-medially directed allux, which is absent or very rare and inconspicuous in Koreanaornis. Features of ‘oreanaornis dodsoni that distinguish it from K. hamanensis include: smaller divaricatiot ngles between digits II and IV, greater overall track length and width, persistent absenc f digit I, and absence of digital pad impressions. Compared with the original type ‘oreanaornis hamanensis, K. dodsoni is larger, with a reported, more variable size range o .0-6.3 cm (Xing et al., 2011). The larger measurement is markedly inconsistent with th lentification as Koreanaornis. Likewise the K. dodsoni holotype (length 4.4 cm, width .7 cm) is more typical of Aquatilavipes, (ichnofamily Avipedidae according to Sarjeant & angston, 1994) and raises the question of whether K. dodsoni would be better ccommodated in Aquatilavipes dodsoni as a new combination. However, because ‘retaceous shorebird tracks need a comprehensive ichnotaxonomic revision, we avoid a AaArrataA anacatikKnrmant " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_025.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095557/figure-26-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 26 Photograph (A) and interpretive outline drawing (B)of bird tracks at Huangyangquan sites: YLSNHMO01201-4-6. Full-size My DOI: 10.7717/peerj.11476/fig-26 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_026.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095567/figure-27-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 27 Photograph (A) and interpretive outline drawing (B) of bird tracks at Huangyangquan sites: YLSNHMO01241. Full-size My DOI: 10.7717/peerj.11476/fig-27 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_027.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095572/figure-28-the-interpretive-outline-drawing-of-bird-tracks-at"><img alt="Figure 28 The interpretive outline drawing of bird tracks at Huangyangquan sites: HYQ-18 (A), 11 (B); YLSNHM01282 (C); HYQ-12 (D). Full-size MY DOI: 10.7717/peerj.11476/fig-28 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_028.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095580/figure-29-the-interpretive-outline-drawing-of-stegosaur"><img alt="Figure 29 The interpretive outline drawing of stegosaur tracks at Huangyangquan sites: YLSNHM01256 (A), 01258 (B), 01260 (C), 01263 (D); HYQ-D-14 (E), 15 (£). Full-size My DOI: 10.7717/peerj.11476/fig-29 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_029.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095584/figure-30-the-manus-prints-are-asymmetrical-digitigrade-and"><img alt="The manus prints are asymmetrical, digitigrade, and consist of three digit impression casts. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_030.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095592/figure-31-photograph-elevation-image-and-interpretive"><img alt="Figure 31 Photograph (A), elevation image (B) and interpretive outline drawing (C) of pterosaur tracks HYQ-7 at Huangyangquan sites. Full-size Mal DOI: 10.7717/peerj.11476/fig-31 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_031.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095597/figure-32-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 32 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-A-1 (A), 2 (B), 5 (C), 8 (D), 7 (E), 9 (F), 10 (G), 12 (H), 26 (I). Full-size RY DOI: 10.7717/peerj.11476/fig-32 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_032.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095602/figure-33-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 33 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-A-13 (A), 14 (B); HYQ-J-2 (C); HYQ-A-25 (D); HYQ-E-4 (E); HYQ-J-3 (F). Full-size MY DOI: 10.7717/peerj.11476/fig-33 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_033.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095603/figure-34-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 34 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-J-4 (A), 5 (B). Full-size My DOI: 10.7717/peerj.11476/fig-34 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_034.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095607/figure-35-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 35 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-J-6 (A); 7 (B), 8 (C), 11 (D), 12 (E). Full-size My DOI: 10.7717/peerj.11476/fig-35 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_035.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095610/figure-36-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 36 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: HYQ-8 (A); YLSNHMO01218 (B); HYQ-A-19 (C); YLSNHMO01233 (D), 01229 (£). Full-size My DOI: 10.7717/peerj.11476/fig-36 YLSNHM01218 preserves two manus prints (5m and 6m) that show some " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_036.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095613/figure-37-the-interpretive-outline-drawing-of-pterosaur"><img alt="Figure 37 The interpretive outline drawing of pterosaur tracks at Huangyangquan sites: YLSNHM01205 (A), 01202 (B), 01219 (C); HYQ-10 (D). Full-size MY DOI: 10.7717/peerj.11476/fig-37 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_037.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095616/figure-38-the-interpretive-outline-drawing-of-pterosaur-and"><img alt="Figure 38 The interpretive outline drawing of pterosaur and turtle tracks at Huangyangquan sites: HYQ-6 (A); YLSNHM01297 (B); HYQ-4 (C), 3 (D); YLSNHMO01266 (E); HYQ-2 (F), 1 (G). Full-size MY DOI: 10.7717/peerj.11476/fig-38 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_038.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095619/figure-39-lower-cretaceous-avian-dominated-theropod"><img alt="" class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_039.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095621/figure-40-lower-cretaceous-avian-dominated-theropod"><img alt="" class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_040.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095624/figure-41-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 41 Photograph (A) and interpretive outline drawing (B) of turtle tracks YLSNHM01206 at Huangyangquan sites. Full-size Mal DOI: 10.7717/peerj.11476/fig-41 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_041.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095628/figure-42-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 42 Photograph (A) and interpretive outline drawing (B) of turtle tracks HYQ-A-6 at Huangyangquan sites. Full-size Mal DOI: 10.7717/peerj.11476/fig-42 Morphotype A is best represented by HYQ-3 (Fig. 38). HYQ-3 is 5.3cm in length, with a length/width ratio of 0.9. HYQ-3 is typically digitigrade, and has four elongated ungual marks; the marks are roughly parallel and face anteriorly. The middle ungual marks (digits II and III) are longer than the lateral two (digits I and IV). The similar track, HYQ-A-11-2 (Fig. 39), is 1.2 cm in length, with a length/width ratio of 0.7, and only has three ungual marks - probably missing a relatively shallow impression of the lateral digit. The medial digit is the longest, while the lateral digits are subequal. The similar track assemblage YLSNHM01206 (Fig. 41) indicates a mean length/width ratio of 1.1, range from 0.3-1.9. three ungual marks — probably missing a relatively shallow impression of the lateral digit. Morphotype B is represented by HYQ-4 (Fig. 38), which is 8.9 cm in length, with a length/width ratio of 2.3. HYQ-4 is typically digitigrade, or semiplantigrade and has elongated ungual marks. Morphotype B most significantly differs from morphotype A in the higher length/width ratio and the number of digits. HYQ-4 has three ungual marks; " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_042.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095631/figure-43-photograph-and-interpretive-outline-drawing-of"><img alt="Figure 43 Photograph (A) and interpretive outline drawing of theropod tracks As-TI1 (B), 2 (C), 3 (D) at Asphaltite sites. Full-size My DOI: 10.7717/peerj.11476/fig-43 " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/figure_043.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095635/table-1-comparison-of-the-type-material-of-the-monospecific"><img alt="Table 1 Comparison of the type material of the monospecific ichnogenus Chapus and the three ichnospecies of Asianopdus. “ Indicates the number of trackmakers, such as “2 (1)”, a set of pes and manus traces left by one trackmaker. (To), Total; (Is), Isolated; (Tw), Trackway; (Tm), Trackmaker; (Ra), Ratio. Numbers under Li, Jiang & Wang, 2020 refer to data from this paper; numbers under “Museum” refer to museum specimens. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/table_001.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095642/table-2-comparison-of-the-type-material-of-the-monospecific"><img alt="Table 2 Comparison of the type material of the monospecific ichnogenus Chapus and the three ichnospecies of Asianopdus. " class="figure-slide-image" src="https://figures.academia-assets.com/77104259/table_002.jpg" /></a></figure><figure class="figure-slide-container"><a href="https://www.academia.edu/figures/33095646/table-3-comparison-of-bone-records-and-track-records-of"><img alt="Table 3 Comparison of bone records and track records of Wuerhe area, Xinjiang, China. The Huangyangquan sites from the Lower Layer, and the Asphaltite site from the Upper Layer. Black squares indicate the existence of records. 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They originate from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, variously divided into four formations and subgroups in the northwestern margin of the Junggar Basin. The large Huangyangquan track assemblages occur in the Lower layer/Subgroup II. Similarly-composed track assemblages also occur at the smaller Asphaltite site in the Upper Layer/Subgroup III. The Huangyangquan assemblages have yielded more than 1,500 identified tracks including abundant tracks of avian and non-avian theropods, pterosaurs and turtles and less abundant tracks of stegosaurs. Previous avian track identifications have been reassessed to conclude that Moguiornipes robustus is a taphotaxon and Koreanaornis dodsoni might be better accommodated in the ichno...","publisher":"PeerJ","ai_title_tag":"Lower Cretaceous Track Assemblages in Xinjiang, China","publication_name":"PeerJ"},"translated_abstract":"Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe District, Karamay City, Xinjiang) have been an abundant source of some of the largest Lower Cretaceous track collections from China. They originate from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, variously divided into four formations and subgroups in the northwestern margin of the Junggar Basin. The large Huangyangquan track assemblages occur in the Lower layer/Subgroup II. Similarly-composed track assemblages also occur at the smaller Asphaltite site in the Upper Layer/Subgroup III. The Huangyangquan assemblages have yielded more than 1,500 identified tracks including abundant tracks of avian and non-avian theropods, pterosaurs and turtles and less abundant tracks of stegosaurs. Previous avian track identifications have been reassessed to conclude that Moguiornipes robustus is a taphotaxon and Koreanaornis dodsoni might be better accommodated in the ichno...","internal_url":"https://www.academia.edu/65564666/Lower_cretaceous_avian_dominated_theropod_thyreophoran_pterosaur_and_turtle_track_assemblages_from_the_Tugulu_Group_Xinjiang_China_ichnotaxonomy_and_palaeoecology","translated_internal_url":"","created_at":"2021-12-22T13:37:40.152-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":77104259,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/77104259/thumbnails/1.jpg","file_name":"11476.pdf","download_url":"https://www.academia.edu/attachments/77104259/download_file","bulk_download_file_name":"Lower_cretaceous_avian_dominated_theropo.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/77104259/11476-libre.pdf?1640210502=\u0026response-content-disposition=attachment%3B+filename%3DLower_cretaceous_avian_dominated_theropo.pdf\u0026Expires=1743195464\u0026Signature=DLLoVhUjvzKSt~4PJEXdT-YgSdK~psi-ENE-ZCslxUSKix9kI4Exe61Lz5gjCBFHW6OA2Rd3IRe-pUuRMuwJe94ZxjKNnBRtYl2Qp81Gzoru5snTxF-900-bqrdpiou1VlBGpPrx4K97JRLFEnHVxo5~VdhVTxfm8XQiQNGAIQJ1j2rCOeT~-dIBaXNPfkOUMARTkyeY7-0kvm3~~5NbXXJx5rcutEgWTkpf-gDJGXa6mg483iQtToTpaSIdOfX4OiSPs1sr--EvkHCijPIF5XxYT~Tw9cMKoMTY4C21K9I1sWbts1cwznaq2hnP0AoZ~Xnd1VXWE8EquLLiapiavg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Lower_cretaceous_avian_dominated_theropod_thyreophoran_pterosaur_and_turtle_track_assemblages_from_the_Tugulu_Group_Xinjiang_China_ichnotaxonomy_and_palaeoecology","translated_slug":"","page_count":73,"language":"en","content_type":"Work","summary":"Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe District, Karamay City, Xinjiang) have been an abundant source of some of the largest Lower Cretaceous track collections from China. They originate from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, variously divided into four formations and subgroups in the northwestern margin of the Junggar Basin. The large Huangyangquan track assemblages occur in the Lower layer/Subgroup II. Similarly-composed track assemblages also occur at the smaller Asphaltite site in the Upper Layer/Subgroup III. The Huangyangquan assemblages have yielded more than 1,500 identified tracks including abundant tracks of avian and non-avian theropods, pterosaurs and turtles and less abundant tracks of stegosaurs. Previous avian track identifications have been reassessed to conclude that Moguiornipes robustus is a taphotaxon and Koreanaornis dodsoni might be better accommodated in the ichno...","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[{"id":77104259,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/77104259/thumbnails/1.jpg","file_name":"11476.pdf","download_url":"https://www.academia.edu/attachments/77104259/download_file","bulk_download_file_name":"Lower_cretaceous_avian_dominated_theropo.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/77104259/11476-libre.pdf?1640210502=\u0026response-content-disposition=attachment%3B+filename%3DLower_cretaceous_avian_dominated_theropo.pdf\u0026Expires=1743195464\u0026Signature=DLLoVhUjvzKSt~4PJEXdT-YgSdK~psi-ENE-ZCslxUSKix9kI4Exe61Lz5gjCBFHW6OA2Rd3IRe-pUuRMuwJe94ZxjKNnBRtYl2Qp81Gzoru5snTxF-900-bqrdpiou1VlBGpPrx4K97JRLFEnHVxo5~VdhVTxfm8XQiQNGAIQJ1j2rCOeT~-dIBaXNPfkOUMARTkyeY7-0kvm3~~5NbXXJx5rcutEgWTkpf-gDJGXa6mg483iQtToTpaSIdOfX4OiSPs1sr--EvkHCijPIF5XxYT~Tw9cMKoMTY4C21K9I1sWbts1cwznaq2hnP0AoZ~Xnd1VXWE8EquLLiapiavg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"},{"id":77104261,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/77104261/thumbnails/1.jpg","file_name":"11476.pdf","download_url":"https://www.academia.edu/attachments/77104261/download_file","bulk_download_file_name":"Lower_cretaceous_avian_dominated_theropo.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/77104261/11476-libre.pdf?1640210502=\u0026response-content-disposition=attachment%3B+filename%3DLower_cretaceous_avian_dominated_theropo.pdf\u0026Expires=1743195464\u0026Signature=drcsrZnw1ZELMQ2ipaEPYBIOPaAq7MjtXe-47HbjNyUbtwWpLfPaSBZAD9sQ9wg1fCXcCg-NQRWJSUnbMj6-jiMHzgzkELdnR5hhlIxKfOXk6J~xF4y2anE12Tas46cVIlqyiHxOl6ldQivjxD-tJVhU4Im4gqaY9mNZVHTnu~0Ize712Y9HpRZarh6m4ldOThte2RHq8ojkFnBLwnzbQ31oRrsUjgg7W69MCzW6akr~hVr9RNitIzaQD~xah2cFIg5LT4IgkD1BhfJSZoS8K-jAylowtcW1bhN7EbuoaBVzkF9MukYutTjRxvounkDZ~KUDKgcSzlK0B2haCQZC3w__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[],"urls":[{"id":15552836,"url":"https://peerj.com/articles/11476.pdf"}]}, dispatcherData: dispatcherData }); 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While histor...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Fossil footprints provide the only evidence of Triassic-aged dinosaurs in Australia. While historically several Triassic dinosaur tracksites are reported, only one specimen exists for direct study. Evidence for the Rhondda colliery (Ipswich) trackmaker, Upper Triassic Blackstone Formation as a very large-bodied Triassic predatory dinosaur is not supported by the present study because of overestimated footprint length measurements, and we question earlier interpretations that these tracks are Eubrontes-like. Instead, we recognise shared characteristics with the sauropodomorph ichnogenus Evazoum. The track morphology resembles that of Evazoum on account of the distinct shape and position of the impression of the metatarsophalangeal pad for digit IV. The mesaxony of the Blackstone Formation track is also low, but possibly slightly higher than in Evazoum, if the proportions of the digital impressions are correctly interpreted. Finally, the trackway pattern shows constant track inward rotation, a feature observed in typical Evazoum and mostly absent in Eubrontes. Therefore, we assign these Australian tracks as Evazoum ichnosp. indet. If our assessment of the Blackstone Formation tracks is correct, they provide the first and only evidence of basal sauropodomorph dinosaurs from Australia. In addition to this record, we give an overview of other reported (but lost) Australian Triassic dinosaur footprints.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="8f8663ea3430136d6335274efb2f65f1" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":73844038,"asset_id":60365280,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/73844038/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="60365280"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="60365280"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 60365280; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=60365280]").text(description); $(".js-view-count[data-work-id=60365280]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 60365280; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='60365280']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "8f8663ea3430136d6335274efb2f65f1" } } $('.js-work-strip[data-work-id=60365280]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":60365280,"title":"Saurischian dinosaur tracks from the Upper Triassic of southern Queensland: possible evidence for Australia’s earliest sauropodomorph trackmaker","translated_title":"","metadata":{"publisher":"Informa UK Limited","grobid_abstract":"Fossil footprints provide the only evidence of Triassic-aged dinosaurs in Australia. 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Therefore, we assign these Australian tracks as Evazoum ichnosp. indet. If our assessment of the Blackstone Formation tracks is correct, they provide the first and only evidence of basal sauropodomorph dinosaurs from Australia. In addition to this record, we give an overview of other reported (but lost) Australian Triassic dinosaur footprints.","publication_name":"Historical Biology","grobid_abstract_attachment_id":73844039},"translated_abstract":null,"internal_url":"https://www.academia.edu/60365280/Saurischian_dinosaur_tracks_from_the_Upper_Triassic_of_southern_Queensland_possible_evidence_for_Australia_s_earliest_sauropodomorph_trackmaker","translated_internal_url":"","created_at":"2021-10-29T18:29:52.977-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":73844038,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/73844038/thumbnails/1.jpg","file_name":"08912963.2021.pdf","download_url":"https://www.academia.edu/attachments/73844038/download_file","bulk_download_file_name":"Saurischian_dinosaur_tracks_from_the_Upp.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/73844038/08912963.2021-libre.pdf?1635558942=\u0026response-content-disposition=attachment%3B+filename%3DSaurischian_dinosaur_tracks_from_the_Upp.pdf\u0026Expires=1743195464\u0026Signature=Dzn-KIFH8rOlO4eQ3u5vu0-axhFi9WJtppZY~QxhXaMc-Fj6CzqoBZaSrXsDM9M8c-CE3G1OCCeMHWdvPlfsQCWYPX9DI7be7hyhbI6mjv5LnzhyEB2lIt3DnYNcQML22aG~WCWKnV429K-HBD-oZLLXcnzI9TfbjON7mgGo-lr3WqFuFa9CGO5fMphfEHctBc0LXn-dioaM-PORB~r5ocjDjJr2UwFFR5oux7SdFbjlngTxJ3Mr6jy7E-NzgWL-Xbxx28j1HzA1Mat9Mj8eVwPoMv2wKUIFadC1WzaCvN5heBKiWfQg40UEs1nOcipd-5F4HemmRMgjX1IvJYIopg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Saurischian_dinosaur_tracks_from_the_Upper_Triassic_of_southern_Queensland_possible_evidence_for_Australia_s_earliest_sauropodomorph_trackmaker","translated_slug":"","page_count":11,"language":"en","content_type":"Work","summary":"Fossil footprints provide the only evidence of Triassic-aged dinosaurs in Australia. 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To date, a moderate diversity of six theropod ichnogenera has been reported, but none of these have been identified at the ichnospecies level. Thus, many morphotypes have common “generic” labels such as Grallator, Eubrontes, cf. Eubrontes or even “Eubrontes-Megalosauripus” morphotype. These morphotypes are generally more typical of the Jurassic, whereas other more distinctive theropod tracks (Minisauripus and Velociraptorichnus) are restricted to the Cretaceous. The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very well-preserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="7f6ceb987b81e668b15cc40aac58719a" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":72499007,"asset_id":57753746,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/72499007/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="57753746"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="57753746"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 57753746; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=57753746]").text(description); $(".js-view-count[data-work-id=57753746]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 57753746; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='57753746']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "7f6ceb987b81e668b15cc40aac58719a" } } $('.js-work-strip[data-work-id=57753746]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":57753746,"title":"The new ichnotaxon Eubrontes nobitai ichnosp. nov. and other saurischian tracks from the Lower Cretaceous of Sichuan Province and a review of Chinese Eubrontes-type tracks","translated_title":"","metadata":{"abstract":"The Jiaguan Formation and the underlying Feitianshan Formation (Lower Cretaceous) in Sichuan Province yield multiple saurischian (theropod–sauropod) dominated ichnofaunas. 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The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very well-preserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.","publisher":"Springer Science and Business Media LLC","ai_title_tag":"New Cretaceous Eubrontes nobitai from Sichuan","publication_name":"Journal of Palaeogeography"},"translated_abstract":"The Jiaguan Formation and the underlying Feitianshan Formation (Lower Cretaceous) in Sichuan Province yield multiple saurischian (theropod–sauropod) dominated ichnofaunas. To date, a moderate diversity of six theropod ichnogenera has been reported, but none of these have been identified at the ichnospecies level. Thus, many morphotypes have common “generic” labels such as Grallator, Eubrontes, cf. Eubrontes or even “Eubrontes-Megalosauripus” morphotype. These morphotypes are generally more typical of the Jurassic, whereas other more distinctive theropod tracks (Minisauripus and Velociraptorichnus) are restricted to the Cretaceous. The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very well-preserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.","internal_url":"https://www.academia.edu/57753746/The_new_ichnotaxon_Eubrontes_nobitai_ichnosp_nov_and_other_saurischian_tracks_from_the_Lower_Cretaceous_of_Sichuan_Province_and_a_review_of_Chinese_Eubrontes_type_tracks","translated_internal_url":"","created_at":"2021-10-14T02:57:49.746-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":72499007,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/72499007/thumbnails/1.jpg","file_name":"s42501-021-00096-y.pdf","download_url":"https://www.academia.edu/attachments/72499007/download_file","bulk_download_file_name":"The_new_ichnotaxon_Eubrontes_nobitai_ich.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/72499007/s42501-021-00096-y-libre.pdf?1634206350=\u0026response-content-disposition=attachment%3B+filename%3DThe_new_ichnotaxon_Eubrontes_nobitai_ich.pdf\u0026Expires=1743195464\u0026Signature=GYT9zF5lyHOIY7TodYYadD4kVkneNPPcK7TMXEd256lwOKRJG69Y1YAaAHajsVuWC0A065Mqbn5xJ7eeilrvcXyaICrnyPTCoB~Ao7gQ0fClCsqD1DiOSFqW3KTXpwVfzPSY8JHfBfJVXynW~F0zZO6V8MJEmFShxElUcqIyXBvY2H5YsH2X0iz8-2EBF3sbHGzyjv2E2EhBhJHGDVNJ0Hm3Q554dh7LaNjXP3V1ntWYWt77ZUF~fqklgyFgxbNKLfGfUqT-eHEJr-giMGcKyXr~lZJ9T3KHM0jDnI~qeqQuDuXvcaeaM5D5FkBoM8Ri3pUS2UeIgNHMzydJQm7oFA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"The_new_ichnotaxon_Eubrontes_nobitai_ichnosp_nov_and_other_saurischian_tracks_from_the_Lower_Cretaceous_of_Sichuan_Province_and_a_review_of_Chinese_Eubrontes_type_tracks","translated_slug":"","page_count":19,"language":"en","content_type":"Work","summary":"The Jiaguan Formation and the underlying Feitianshan Formation (Lower Cretaceous) in Sichuan Province yield multiple saurischian (theropod–sauropod) dominated ichnofaunas. To date, a moderate diversity of six theropod ichnogenera has been reported, but none of these have been identified at the ichnospecies level. Thus, many morphotypes have common “generic” labels such as Grallator, Eubrontes, cf. Eubrontes or even “Eubrontes-Megalosauripus” morphotype. These morphotypes are generally more typical of the Jurassic, whereas other more distinctive theropod tracks (Minisauripus and Velociraptorichnus) are restricted to the Cretaceous. The new ichnospecies Eubrontes nobitai ichnosp nov. is distinguished from Jurassic morphotypes based on a very well-preserved trackway and represents the first-named Eubrontes ichnospecies from the Cretaceous of Asia.","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[{"id":72499007,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/72499007/thumbnails/1.jpg","file_name":"s42501-021-00096-y.pdf","download_url":"https://www.academia.edu/attachments/72499007/download_file","bulk_download_file_name":"The_new_ichnotaxon_Eubrontes_nobitai_ich.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/72499007/s42501-021-00096-y-libre.pdf?1634206350=\u0026response-content-disposition=attachment%3B+filename%3DThe_new_ichnotaxon_Eubrontes_nobitai_ich.pdf\u0026Expires=1743195464\u0026Signature=GYT9zF5lyHOIY7TodYYadD4kVkneNPPcK7TMXEd256lwOKRJG69Y1YAaAHajsVuWC0A065Mqbn5xJ7eeilrvcXyaICrnyPTCoB~Ao7gQ0fClCsqD1DiOSFqW3KTXpwVfzPSY8JHfBfJVXynW~F0zZO6V8MJEmFShxElUcqIyXBvY2H5YsH2X0iz8-2EBF3sbHGzyjv2E2EhBhJHGDVNJ0Hm3Q554dh7LaNjXP3V1ntWYWt77ZUF~fqklgyFgxbNKLfGfUqT-eHEJr-giMGcKyXr~lZJ9T3KHM0jDnI~qeqQuDuXvcaeaM5D5FkBoM8Ri3pUS2UeIgNHMzydJQm7oFA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":289,"name":"Palaeogeography","url":"https://www.academia.edu/Documents/in/Palaeogeography"}],"urls":[{"id":12973464,"url":"https://link.springer.com/content/pdf/10.1186/s42501-021-00096-y.pdf"}]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-57753746-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="49372061"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/49372061/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea"><img alt="Research paper thumbnail of Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea" class="work-thumbnail" src="https://attachments.academia-assets.com/67745462/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/49372061/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea">Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea</a></div><div class="wp-workCard_item"><span>Scientific Reports</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation o...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation of South Korea, represent the well-known crocodylomorph ichnogenus Batrachopus. the Korean sample includes multiple, narrow-gauge, pes-only trackways with footprint lengths (FL) 18-24 cm, indicating trackmaker body lengths up to ~3.0 m. Surprisingly, the consistent absence of manus tracks in trackways, with well-preserved digital pad and skin traces, argues for bipedal trackmakers, here assigned to Batrachopus grandis ichnosp. nov. No definitive evidence, either from pes-onmanus overprinting or poor track preservation, suggests the trackways where made by quadrupeds that only appear bipedal. this interpretation helps solve previous confusion over interpretation of enigmatic tracks of bipeds from younger (? Albian) Haman formation sites by showing they are not pterosaurian as previously inferred. Rather, they support the strong consensus that pterosaurs were obligate quadrupeds, not bipeds. Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. Surprisingly the trackways never include manus imprints and therefore appear to indicate exclusively</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="7a1839d499c9aae9969f4ff422038635" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":67745462,"asset_id":49372061,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/67745462/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="49372061"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="49372061"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 49372061; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=49372061]").text(description); $(".js-view-count[data-work-id=49372061]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 49372061; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='49372061']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "7a1839d499c9aae9969f4ff422038635" } } $('.js-work-strip[data-work-id=49372061]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":49372061,"title":"Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea","translated_title":"","metadata":{"publisher":"Springer Science and Business Media LLC","ai_title_tag":"Bipedal Crocodylomorph Tracks from Cretaceous Korea","grobid_abstract":"Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation of South Korea, represent the well-known crocodylomorph ichnogenus Batrachopus. the Korean sample includes multiple, narrow-gauge, pes-only trackways with footprint lengths (FL) 18-24 cm, indicating trackmaker body lengths up to ~3.0 m. Surprisingly, the consistent absence of manus tracks in trackways, with well-preserved digital pad and skin traces, argues for bipedal trackmakers, here assigned to Batrachopus grandis ichnosp. nov. No definitive evidence, either from pes-onmanus overprinting or poor track preservation, suggests the trackways where made by quadrupeds that only appear bipedal. this interpretation helps solve previous confusion over interpretation of enigmatic tracks of bipeds from younger (? Albian) Haman formation sites by showing they are not pterosaurian as previously inferred. Rather, they support the strong consensus that pterosaurs were obligate quadrupeds, not bipeds. Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. Surprisingly the trackways never include manus imprints and therefore appear to indicate exclusively","publication_name":"Scientific Reports","grobid_abstract_attachment_id":67745463},"translated_abstract":null,"internal_url":"https://www.academia.edu/49372061/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea","translated_internal_url":"","created_at":"2021-06-25T15:09:57.067-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":67745462,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/67745462/thumbnails/1.jpg","file_name":"s41598-020-66008-7.pdf","download_url":"https://www.academia.edu/attachments/67745462/download_file","bulk_download_file_name":"Trackway_evidence_for_large_bipedal_croc.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/67745462/s41598-020-66008-7-libre.pdf?1624664665=\u0026response-content-disposition=attachment%3B+filename%3DTrackway_evidence_for_large_bipedal_croc.pdf\u0026Expires=1743195464\u0026Signature=L5fTCCNNG0NotDC3AB3fINC1CBXi9Xp8tsbn5TLYInGkqxz6HufeLbPmbs0egC~dSoDg2cTuggfLMcE8ZEAOgz7Dr06Pu79rAooHDvjueWNju6rUJkvYIyKw56LfGHWnfmCTt2zNNI0o7xq2Mkp7eGsMamFsbmB692ABmlO5sMtR9gp8O4GZj92QtdVZxA-qyWwB6i2OKSou9Lhyj4tpwyrMwkbSGOQ0xM0hqhmeGQrjlxNwyIdu4SKoMZLv5XFP6~VzbPdD4AY2ebDMpqyYd3QUbkf1rGkWy-eIaMWy9RxdG84HIX6KS8l8RWuirIi8pQcbNFTh0Eit1WJPjmg3Hg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea","translated_slug":"","page_count":13,"language":"en","content_type":"Work","summary":"Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation of South Korea, represent the well-known crocodylomorph ichnogenus Batrachopus. the Korean sample includes multiple, narrow-gauge, pes-only trackways with footprint lengths (FL) 18-24 cm, indicating trackmaker body lengths up to ~3.0 m. Surprisingly, the consistent absence of manus tracks in trackways, with well-preserved digital pad and skin traces, argues for bipedal trackmakers, here assigned to Batrachopus grandis ichnosp. nov. No definitive evidence, either from pes-onmanus overprinting or poor track preservation, suggests the trackways where made by quadrupeds that only appear bipedal. this interpretation helps solve previous confusion over interpretation of enigmatic tracks of bipeds from younger (? Albian) Haman formation sites by showing they are not pterosaurian as previously inferred. Rather, they support the strong consensus that pterosaurs were obligate quadrupeds, not bipeds. Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. Surprisingly the trackways never include manus imprints and therefore appear to indicate exclusively","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[{"id":67745462,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/67745462/thumbnails/1.jpg","file_name":"s41598-020-66008-7.pdf","download_url":"https://www.academia.edu/attachments/67745462/download_file","bulk_download_file_name":"Trackway_evidence_for_large_bipedal_croc.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/67745462/s41598-020-66008-7-libre.pdf?1624664665=\u0026response-content-disposition=attachment%3B+filename%3DTrackway_evidence_for_large_bipedal_croc.pdf\u0026Expires=1743195464\u0026Signature=L5fTCCNNG0NotDC3AB3fINC1CBXi9Xp8tsbn5TLYInGkqxz6HufeLbPmbs0egC~dSoDg2cTuggfLMcE8ZEAOgz7Dr06Pu79rAooHDvjueWNju6rUJkvYIyKw56LfGHWnfmCTt2zNNI0o7xq2Mkp7eGsMamFsbmB692ABmlO5sMtR9gp8O4GZj92QtdVZxA-qyWwB6i2OKSou9Lhyj4tpwyrMwkbSGOQ0xM0hqhmeGQrjlxNwyIdu4SKoMZLv5XFP6~VzbPdD4AY2ebDMpqyYd3QUbkf1rGkWy-eIaMWy9RxdG84HIX6KS8l8RWuirIi8pQcbNFTh0Eit1WJPjmg3Hg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"},{"id":67745463,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/67745463/thumbnails/1.jpg","file_name":"s41598-020-66008-7.pdf","download_url":"https://www.academia.edu/attachments/67745463/download_file","bulk_download_file_name":"Trackway_evidence_for_large_bipedal_croc.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/67745463/s41598-020-66008-7-libre.pdf?1624664664=\u0026response-content-disposition=attachment%3B+filename%3DTrackway_evidence_for_large_bipedal_croc.pdf\u0026Expires=1743195464\u0026Signature=YhU9iYWcaLxuyI0LyUNuSSGY~rK4XCOIv9lKEp-TrwYfI1ZvSZ1~qNFX0mWiDT35J6Nv8GWKu58mODa~zFtbIeeMCRj5wU8SN0KPj-U2CW7ThcaNqsL9~gr-SjTNk7WTmJiOtHhyPVJHvwu-jpvJn6WFskRDHwNhi4LE75gRmIQro29ArUR7tQzw2wI8oYIU11TZWmsg6-WkRFQOPI1FHQAV6iLuQH9FaTk~pvL-E8A-~GdLp59JWODghYA9OxMUsUo4m4UlpArJhxjh7uaz8kKVT09Ikf9GvcxmOM9bOZq25fFioIn8PANPvEZbCJ4Vzu2eaghGnJQwZtMQ4ZUUNw__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[],"urls":[{"id":10298754,"url":"http://www.nature.com/articles/s41598-020-66008-7.pdf"}]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-49372061-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="49372060"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/49372060/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea"><img alt="Research paper thumbnail of Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea" class="work-thumbnail" src="https://attachments.academia-assets.com/67745459/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/49372060/Trackway_evidence_for_large_bipedal_crocodylomorphs_from_the_Cretaceous_of_Korea">Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea</a></div><div class="wp-workCard_item"><span>Scientific Reports</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation o...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Large well-preserved crocodylomorph tracks from the Lower cretaceous (? Aptian) Jinju formation of South Korea, represent the well-known crocodylomorph ichnogenus Batrachopus. the Korean sample includes multiple, narrow-gauge, pes-only trackways with footprint lengths (FL) 18-24 cm, indicating trackmaker body lengths up to ~3.0 m. Surprisingly, the consistent absence of manus tracks in trackways, with well-preserved digital pad and skin traces, argues for bipedal trackmakers, here assigned to Batrachopus grandis ichnosp. nov. No definitive evidence, either from pes-onmanus overprinting or poor track preservation, suggests the trackways where made by quadrupeds that only appear bipedal. this interpretation helps solve previous confusion over interpretation of enigmatic tracks of bipeds from younger (? Albian) Haman formation sites by showing they are not pterosaurian as previously inferred. Rather, they support the strong consensus that pterosaurs were obligate quadrupeds, not bipeds. Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. Surprisingly the trackways never include manus imprints and therefore appear to indicate exclusively</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="1d0a928b94c38bf61a452e87d80ae5a5" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":67745459,"asset_id":49372060,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/67745459/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="49372060"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="49372060"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 49372060; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=49372060]").text(description); $(".js-view-count[data-work-id=49372060]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 49372060; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='49372060']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "1d0a928b94c38bf61a452e87d80ae5a5" } } $('.js-work-strip[data-work-id=49372060]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":49372060,"title":"Trackway evidence for large bipedal crocodylomorphs from the Cretaceous of Korea","translated_title":"","metadata":{"publisher":"Springer Science and Business Media LLC","ai_title_tag":"Bipedal Crocodylomorph Trackways from Lower Cretaceous Korea","grobid_abstract":"Large well-preserved crocodylomorph tracks from the Lower cretaceous (? 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Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. 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Lower Jurassic Batrachopus with foot lengths (FL) in the 2-8 cm range, and cretaceous Crocodylopodus (FL up to ~9.0 cm) known only from Korea and Spain registered narrow gauge trackways indicating semi-terrestrial/terrestrial quadrupedal gaits. Both ichnogenera, from ichnofamily Batrachopodidae, have been attributed to Protosuchus-like semi-terrestrial crocodylomorphs. the occurrence of bipedal B. grandis ichnosp. nov. is evidence of such adaptations in the Korean cretaceous. Crocodylomorph tracks are generally rare in the Mesozoic of Asia. It has been suggested that this is in part due to the lack of sedimentary facies representing suitable habitats for this group of ostensibly aquatic trackmakers 1. However, the crocodylomorph ichnofamily Batrachopodidae 2,3 appears to represent more terrestrially-adapted forms 3,4. As currently defined, the batrachopodids 5 include Batrachopus and Crocodylopodus, the former mostly known from small tracks (~2.0-8.0 cm long) from the Lower Jurassic of North America 4 , Europe 5,6 and Africa 7 , the latter primarily from the Cretaceous of Europe 8,9. The ichnofamily also includes Antipus known only from one well-described Lower Jurassic trackway from North America which is considered a synonym of Batrachopus by some 3 , but not all ichnologists 10. Recently Crocodylopodus was reported from the Lower Cretaceous (?Aptian) 11 Jinju Formation of Korea 12,13 , where it represents the first Asian occurrence, and adds to the extraordinarily rich Jinju Formation, ichnofauna described as a Konservat-Lagerstätten 14-19. Spanish 8,9 and Korean 12,13 Crocodylopodus represent relatively small animals (footprint lengths less than ~9.0-10.0 cm) with slender digit traces, which may reflect penetrative track preservation 20. However, the trackway configuration is diagnostically crocodilian, with elongate tetradactyl pes and outwardly rotated pentadactyl manus, not unlike those of extant crocodilians 21-23 (Supplementary Information: SI Fig. 1). Here we report on a large, newly-discovered Jinju Formation assemblage with multiple trackways of large crocodylomorph tracks with footprint lengths up to 24.0 cm, from the Sacheon Jahye-ri tracksite, near Sacheon City (Fig. 1). These tracks are more than twice as large as any previously reported batrachopodid tracks and closely resemble Batrachopus with well-preserved pes footprints with clear digital pad impressions and localized skin traces. 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The seven dinosaurian eggs reported clearly belong to the Elongatoolithidae and burrow traces were most likely made by small crustaceans. This association prompts the question as to whether invertebrate activity had relations with the buried eggs. This may be just an occasional case or the eggs may have organically increased the content of organic matter in soil which attracted the crustaceans.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="30020f316cb135ea5539b83f4704bfb9" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":66381013,"asset_id":47118777,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/66381013/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="47118777"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="47118777"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 47118777; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=47118777]").text(description); $(".js-view-count[data-work-id=47118777]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 47118777; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='47118777']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "30020f316cb135ea5539b83f4704bfb9" } } $('.js-work-strip[data-work-id=47118777]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":47118777,"title":"Dinosaur Eggs Associated with Crustacean Trace Fossils from the Upper Cretaceous of Jiangxi, China: Evidence for Foraging Behavior?","translated_title":"","metadata":{"abstract":"We report the discovery of concentrated invertebrate inchnofossils in close association with a dinosaur nest from the Hekou Formation in Jiangxi Province, China. 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BIOL2201: Studies of Dinosaur ichnites</div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="2373395"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="2373395"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 2373395; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=2373395]").text(description); $(".js-view-count[data-work-id=2373395]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 2373395; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='2373395']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=2373395]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":2373395,"title":"2011 Guest Lecturer: Second year evolution The University of Queensland. BIOL2201: Studies of Dinosaur ichnites","translated_title":"","metadata":{},"translated_abstract":null,"internal_url":"https://www.academia.edu/2373395/2011_Guest_Lecturer_Second_year_evolution_The_University_of_Queensland_BIOL2201_Studies_of_Dinosaur_ichnites","translated_internal_url":"","created_at":"2013-01-07T12:40:06.232-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"talk","co_author_tags":[],"downloadable_attachments":[],"slug":"2011_Guest_Lecturer_Second_year_evolution_The_University_of_Queensland_BIOL2201_Studies_of_Dinosaur_ichnites","translated_slug":"","page_count":null,"language":"en","content_type":"Work","summary":null,"owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[],"research_interests":[],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-2373395-figures'); } }); </script> </div><div class="profile--tab_content_container js-tab-pane tab-pane" data-section-id="348729" id="conferencepresentations"><div class="js-work-strip profile--work_container" data-work-id="11383543"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/11383543/Ichnological_evidence_for_pedal_posture_in_ornithopodan_dinosaurs"><img alt="Research paper thumbnail of Ichnological evidence for pedal posture in ornithopodan dinosaurs " class="work-thumbnail" src="https://attachments.academia-assets.com/36945826/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/11383543/Ichnological_evidence_for_pedal_posture_in_ornithopodan_dinosaurs">Ichnological evidence for pedal posture in ornithopodan dinosaurs </a></div><div class="wp-workCard_item wp-workCard--coauthors"><span>by </span><span><a class="" data-click-track="profile-work-strip-authors" href="https://uq.academia.edu/AnthonyRomilio">Anthony Romilio</a> and <a class="" data-click-track="profile-work-strip-authors" href="https://uq.academia.edu/JayNair">Jay Nair</a></span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Evolutionary transitions involving changes in the pedal posture may also lead to changes in assoc...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Evolutionary transitions involving changes in the pedal posture may also lead to changes in associated soft-tissue structures of the pes. For extinct taxa, pedal posture can be gleaned in part from osteological evidence, augmented by ichnological data. Two different pedal postures have previously been proposed for ornithopodan dinosaurs: a digitigrade posture where the metatarsodigital joint is in close proximity to the plantar surface; and a subunguligrade posture where the metatarsodigital joint is permanently elevated. These postures have been linked to plesiomorphic (non-hadrosaurid) and derived (hadrosaurid) ornithopodan body fossil taxa, respectively. However, ichnological data suggest that the majority of ornithopodan taxa were digitigrade, with tracks typically displaying prominent metatarsodigital pad impressions, including those attributed to hadrosaurids. We propose that the information from both the osteological and the ichnological evidence indicates there are distinct ‘skeletal’ and the ‘functional’ pedal postures within Ornithopoda. The likely plesiomorphic condition for ornithopods is a pedal posture that is both skeletally and functionally digitigrade, while the derived hadrosaurid posture is skeletally subunguligrade but functionally digitigrade. The hadrosaurid-like pedal posture was not limited to hadrosaurids, but may also have been present in some basal hadrosauriforms as well as in some basal Ankylopollexia. The latter possibility indicates that the hadrosauroid-like postural configuration may have evolved independently among different clades. Tracks assigned to the ichnogenus Wintonopus indicate that some small- to medium –bodied ornithopodan trackmakers had permanently elevated metatarsodigital joints but without an associated plantar (metatarsodigital) pad. The suggested pedal posture of these trackmakers is skeletally and functionally subunguligrade, a posture that was previously unrecognized for ornithopodans. By intergrating osteological data with soft-tissue data derived from tracks, we find evidence for multiple transformational pathways within Ornithopoda that mirror the attainment of derived postures in extant lineages.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="82ee15b3bc9c1cf10f1a280d086ecf63" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":36945826,"asset_id":11383543,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/36945826/download_file?s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="11383543"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="11383543"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 11383543; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=11383543]").text(description); $(".js-view-count[data-work-id=11383543]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 11383543; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='11383543']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "82ee15b3bc9c1cf10f1a280d086ecf63" } } $('.js-work-strip[data-work-id=11383543]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":11383543,"title":"Ichnological evidence for pedal posture in ornithopodan dinosaurs ","translated_title":"","metadata":{"abstract":"Evolutionary transitions involving changes in the pedal posture may also lead to changes in associated soft-tissue structures of the pes. For extinct taxa, pedal posture can be gleaned in part from osteological evidence, augmented by ichnological data. Two different pedal postures have previously been proposed for ornithopodan dinosaurs: a digitigrade posture where the metatarsodigital joint is in close proximity to the plantar surface; and a subunguligrade posture where the metatarsodigital joint is permanently elevated. These postures have been linked to plesiomorphic (non-hadrosaurid) and derived (hadrosaurid) ornithopodan body fossil taxa, respectively. However, ichnological data suggest that the majority of ornithopodan taxa were digitigrade, with tracks typically displaying prominent metatarsodigital pad impressions, including those attributed to hadrosaurids. We propose that the information from both the osteological and the ichnological evidence indicates there are distinct ‘skeletal’ and the ‘functional’ pedal postures within Ornithopoda. The likely plesiomorphic condition for ornithopods is a pedal posture that is both skeletally and functionally digitigrade, while the derived hadrosaurid posture is skeletally subunguligrade but functionally digitigrade. The hadrosaurid-like pedal posture was not limited to hadrosaurids, but may also have been present in some basal hadrosauriforms as well as in some basal Ankylopollexia. The latter possibility indicates that the hadrosauroid-like postural configuration may have evolved independently among different clades. Tracks assigned to the ichnogenus Wintonopus indicate that some small- to medium –bodied ornithopodan trackmakers had permanently elevated metatarsodigital joints but without an associated plantar (metatarsodigital) pad. The suggested pedal posture of these trackmakers is skeletally and functionally subunguligrade, a posture that was previously unrecognized for ornithopodans. By intergrating osteological data with soft-tissue data derived from tracks, we find evidence for multiple transformational pathways within Ornithopoda that mirror the attainment of derived postures in extant lineages.","ai_title_tag":"Pedal Posture in Ornithopodan Dinosaurs: Ichnological Insights"},"translated_abstract":"Evolutionary transitions involving changes in the pedal posture may also lead to changes in associated soft-tissue structures of the pes. For extinct taxa, pedal posture can be gleaned in part from osteological evidence, augmented by ichnological data. Two different pedal postures have previously been proposed for ornithopodan dinosaurs: a digitigrade posture where the metatarsodigital joint is in close proximity to the plantar surface; and a subunguligrade posture where the metatarsodigital joint is permanently elevated. These postures have been linked to plesiomorphic (non-hadrosaurid) and derived (hadrosaurid) ornithopodan body fossil taxa, respectively. However, ichnological data suggest that the majority of ornithopodan taxa were digitigrade, with tracks typically displaying prominent metatarsodigital pad impressions, including those attributed to hadrosaurids. We propose that the information from both the osteological and the ichnological evidence indicates there are distinct ‘skeletal’ and the ‘functional’ pedal postures within Ornithopoda. The likely plesiomorphic condition for ornithopods is a pedal posture that is both skeletally and functionally digitigrade, while the derived hadrosaurid posture is skeletally subunguligrade but functionally digitigrade. The hadrosaurid-like pedal posture was not limited to hadrosaurids, but may also have been present in some basal hadrosauriforms as well as in some basal Ankylopollexia. The latter possibility indicates that the hadrosauroid-like postural configuration may have evolved independently among different clades. Tracks assigned to the ichnogenus Wintonopus indicate that some small- to medium –bodied ornithopodan trackmakers had permanently elevated metatarsodigital joints but without an associated plantar (metatarsodigital) pad. The suggested pedal posture of these trackmakers is skeletally and functionally subunguligrade, a posture that was previously unrecognized for ornithopodans. By intergrating osteological data with soft-tissue data derived from tracks, we find evidence for multiple transformational pathways within Ornithopoda that mirror the attainment of derived postures in extant lineages.","internal_url":"https://www.academia.edu/11383543/Ichnological_evidence_for_pedal_posture_in_ornithopodan_dinosaurs","translated_internal_url":"","created_at":"2015-03-11T21:28:03.994-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"conference_presentation","co_author_tags":[{"id":255329,"work_id":11383543,"tagging_user_id":379100,"tagged_user_id":6652285,"co_author_invite_id":null,"email":"s***y@uq.edu.au","affiliation":"The University of Queensland, Australia","display_order":null,"name":"Steve Salisbury","title":"Ichnological evidence for pedal posture in ornithopodan dinosaurs "},{"id":255330,"work_id":11383543,"tagging_user_id":379100,"tagged_user_id":293950,"co_author_invite_id":null,"email":"j***r@gmail.com","affiliation":"The University of Queensland, Australia","display_order":null,"name":"Jay Nair","title":"Ichnological evidence for pedal posture in ornithopodan dinosaurs "}],"downloadable_attachments":[{"id":36945826,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/36945826/thumbnails/1.jpg","file_name":"Romilio__Salisbury__Nair_SVP_2014_abstract_01.pdf","download_url":"https://www.academia.edu/attachments/36945826/download_file","bulk_download_file_name":"Ichnological_evidence_for_pedal_posture.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/36945826/Romilio__Salisbury__Nair_SVP_2014_abstract_01-libre.pdf?1426134410=\u0026response-content-disposition=attachment%3B+filename%3DIchnological_evidence_for_pedal_posture.pdf\u0026Expires=1743185331\u0026Signature=I~U4MaT5zMd~u~76tZi3OmbBuWmxVG80Up32oMxkP7qHioqDt-egxFwNpx3C41jJtTHnhue3-AbrBSzlh6BB5Pma3qm9g0OddHQsvthuXIBSvNg1A-HnKLzyWjckl7UKQsCwAl9MWD~T-jIf0UXO79UJTCJ1658yXl50lIbX4Pv97V~5eplDZ7wr1tThHudfNg7vPMhWzCsHwk0OG0x77F8H3-zcmVWs-XIN7q8R9n7agdCZxvm~V~nGXIq6zhGX9ixkBpUA3Bt83Caz3Zp37ul9jda3ntDZPAvHKapCOfRkO4p79GlU8R8niiJElRvAyhzAwuZU8vlWmjXua4m8hQ__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Ichnological_evidence_for_pedal_posture_in_ornithopodan_dinosaurs","translated_slug":"","page_count":2,"language":"en","content_type":"Work","summary":"Evolutionary transitions involving changes in the pedal posture may also lead to changes in associated soft-tissue structures of the pes. For extinct taxa, pedal posture can be gleaned in part from osteological evidence, augmented by ichnological data. Two different pedal postures have previously been proposed for ornithopodan dinosaurs: a digitigrade posture where the metatarsodigital joint is in close proximity to the plantar surface; and a subunguligrade posture where the metatarsodigital joint is permanently elevated. These postures have been linked to plesiomorphic (non-hadrosaurid) and derived (hadrosaurid) ornithopodan body fossil taxa, respectively. However, ichnological data suggest that the majority of ornithopodan taxa were digitigrade, with tracks typically displaying prominent metatarsodigital pad impressions, including those attributed to hadrosaurids. We propose that the information from both the osteological and the ichnological evidence indicates there are distinct ‘skeletal’ and the ‘functional’ pedal postures within Ornithopoda. The likely plesiomorphic condition for ornithopods is a pedal posture that is both skeletally and functionally digitigrade, while the derived hadrosaurid posture is skeletally subunguligrade but functionally digitigrade. The hadrosaurid-like pedal posture was not limited to hadrosaurids, but may also have been present in some basal hadrosauriforms as well as in some basal Ankylopollexia. The latter possibility indicates that the hadrosauroid-like postural configuration may have evolved independently among different clades. Tracks assigned to the ichnogenus Wintonopus indicate that some small- to medium –bodied ornithopodan trackmakers had permanently elevated metatarsodigital joints but without an associated plantar (metatarsodigital) pad. The suggested pedal posture of these trackmakers is skeletally and functionally subunguligrade, a posture that was previously unrecognized for ornithopodans. By intergrating osteological data with soft-tissue data derived from tracks, we find evidence for multiple transformational pathways within Ornithopoda that mirror the attainment of derived postures in extant lineages.","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[{"id":36945826,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/36945826/thumbnails/1.jpg","file_name":"Romilio__Salisbury__Nair_SVP_2014_abstract_01.pdf","download_url":"https://www.academia.edu/attachments/36945826/download_file","bulk_download_file_name":"Ichnological_evidence_for_pedal_posture.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/36945826/Romilio__Salisbury__Nair_SVP_2014_abstract_01-libre.pdf?1426134410=\u0026response-content-disposition=attachment%3B+filename%3DIchnological_evidence_for_pedal_posture.pdf\u0026Expires=1743185331\u0026Signature=I~U4MaT5zMd~u~76tZi3OmbBuWmxVG80Up32oMxkP7qHioqDt-egxFwNpx3C41jJtTHnhue3-AbrBSzlh6BB5Pma3qm9g0OddHQsvthuXIBSvNg1A-HnKLzyWjckl7UKQsCwAl9MWD~T-jIf0UXO79UJTCJ1658yXl50lIbX4Pv97V~5eplDZ7wr1tThHudfNg7vPMhWzCsHwk0OG0x77F8H3-zcmVWs-XIN7q8R9n7agdCZxvm~V~nGXIq6zhGX9ixkBpUA3Bt83Caz3Zp37ul9jda3ntDZPAvHKapCOfRkO4p79GlU8R8niiJElRvAyhzAwuZU8vlWmjXua4m8hQ__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":2588,"name":"Ichnology","url":"https://www.academia.edu/Documents/in/Ichnology"},{"id":24542,"name":"Dinosaur Paleontology","url":"https://www.academia.edu/Documents/in/Dinosaur_Paleontology"},{"id":30201,"name":"Ornithopods","url":"https://www.academia.edu/Documents/in/Ornithopods"},{"id":63152,"name":"Dinosaur Tracks","url":"https://www.academia.edu/Documents/in/Dinosaur_Tracks"},{"id":91553,"name":"Dinosaur footprints and trackways","url":"https://www.academia.edu/Documents/in/Dinosaur_footprints_and_trackways"},{"id":135597,"name":"Hadrosauridae","url":"https://www.academia.edu/Documents/in/Hadrosauridae"},{"id":228356,"name":"Posture","url":"https://www.academia.edu/Documents/in/Posture"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-11383543-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="2373415"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/2373415/Re_interpretation_of_the_dinosaur_track_maker_identities_and_tracksite_scenario_at_Lark_Quarry_of_the_mid_Cretaceous_late_Albian_Cenomanian_Winton_Formation_central_western_Queensland_Australia_Romilio_A_and_S_W_Salisbury"><img alt="Research paper thumbnail of Re-interpretation of the dinosaur track-maker identities and tracksite scenario at Lark Quarry, of the mid-Cretaceous (late Albian-Cenomanian) Winton Formation, central-western Queensland, Australia. Romilio, A., and S.W. Salisbury" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title">Re-interpretation of the dinosaur track-maker identities and tracksite scenario at Lark Quarry, of the mid-Cretaceous (late Albian-Cenomanian) Winton Formation, central-western Queensland, Australia. Romilio, A., and S.W. Salisbury</div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">A re-examination of the track morphology and sedimentary structures does not appear to support th...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">A re-examination of the track morphology and sedimentary structures does not appear to support the original interpretation of the Lark Quarry tracksite preserving the movements of an Allosaurus-sized theropod in pursuit of prey, the mixed herding of small-bodied theropod and ornithopod dinosaurs, or the occurrence of a stampede. Comparative and multivariate data supports the reinterpretation of the large track-maker as an ornithopod, with the tracks assigned to Amblydactylus cf. A. gethingi. Three-dimensional track morphology shows that the small, purportedly theropod prints assigned to Skartopus australis are preservational variants of the ornithopod tracks assigned to Wintonopus latomorum. These re-evaluations remove any evidence for the presence of theropods in the Lark Quarry ichnofauna. The presence of asymmetrical ripples, vegetation drag marks, small dinosaur swimming tracks, and a footprint of a small dinosaur standing in flowing water indicate that the tracks were preserved in a fluvial environment. Superposition of tracks and other sedimentary structures suggests that the Amblydactylus cf. A. gethingi trackmaker moved across the ancient Lark Quarry site earlier than the Wintonopus trackmakers, such that the two trackmaker species probably did not interact.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="2373415"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="2373415"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 2373415; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=2373415]").text(description); $(".js-view-count[data-work-id=2373415]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 2373415; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='2373415']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=2373415]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":2373415,"title":"Re-interpretation of the dinosaur track-maker identities and tracksite scenario at Lark Quarry, of the mid-Cretaceous (late Albian-Cenomanian) Winton Formation, central-western Queensland, Australia. Romilio, A., and S.W. Salisbury","translated_title":"","metadata":{"abstract":"A re-examination of the track morphology and sedimentary structures does not appear to support the original interpretation of the Lark Quarry tracksite preserving the movements of an Allosaurus-sized theropod in pursuit of prey, the mixed herding of small-bodied theropod and ornithopod dinosaurs, or the occurrence of a stampede. Comparative and multivariate data supports the reinterpretation of the large track-maker as an ornithopod, with the tracks assigned to Amblydactylus cf. A. gethingi. Three-dimensional track morphology shows that the small, purportedly theropod prints assigned to Skartopus australis are preservational variants of the ornithopod tracks assigned to Wintonopus latomorum. These re-evaluations remove any evidence for the presence of theropods in the Lark Quarry ichnofauna. The presence of asymmetrical ripples, vegetation drag marks, small dinosaur swimming tracks, and a footprint of a small dinosaur standing in flowing water indicate that the tracks were preserved in a fluvial environment. Superposition of tracks and other sedimentary structures suggests that the Amblydactylus cf. A. gethingi trackmaker moved across the ancient Lark Quarry site earlier than the Wintonopus trackmakers, such that the two trackmaker species probably did not interact.","location":"Obernkirchen, Germany","event_date":{"day":14,"month":4,"year":2011,"errors":{}},"journal_name":"Dinosaur Track Symposium Abstracts Volume"},"translated_abstract":"A re-examination of the track morphology and sedimentary structures does not appear to support the original interpretation of the Lark Quarry tracksite preserving the movements of an Allosaurus-sized theropod in pursuit of prey, the mixed herding of small-bodied theropod and ornithopod dinosaurs, or the occurrence of a stampede. Comparative and multivariate data supports the reinterpretation of the large track-maker as an ornithopod, with the tracks assigned to Amblydactylus cf. A. gethingi. Three-dimensional track morphology shows that the small, purportedly theropod prints assigned to Skartopus australis are preservational variants of the ornithopod tracks assigned to Wintonopus latomorum. These re-evaluations remove any evidence for the presence of theropods in the Lark Quarry ichnofauna. The presence of asymmetrical ripples, vegetation drag marks, small dinosaur swimming tracks, and a footprint of a small dinosaur standing in flowing water indicate that the tracks were preserved in a fluvial environment. Superposition of tracks and other sedimentary structures suggests that the Amblydactylus cf. A. gethingi trackmaker moved across the ancient Lark Quarry site earlier than the Wintonopus trackmakers, such that the two trackmaker species probably did not interact.","internal_url":"https://www.academia.edu/2373415/Re_interpretation_of_the_dinosaur_track_maker_identities_and_tracksite_scenario_at_Lark_Quarry_of_the_mid_Cretaceous_late_Albian_Cenomanian_Winton_Formation_central_western_Queensland_Australia_Romilio_A_and_S_W_Salisbury","translated_internal_url":"","created_at":"2013-01-07T12:56:25.638-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"conference_presentation","co_author_tags":[],"downloadable_attachments":[],"slug":"Re_interpretation_of_the_dinosaur_track_maker_identities_and_tracksite_scenario_at_Lark_Quarry_of_the_mid_Cretaceous_late_Albian_Cenomanian_Winton_Formation_central_western_Queensland_Australia_Romilio_A_and_S_W_Salisbury","translated_slug":"","page_count":null,"language":"en","content_type":"Work","summary":"A re-examination of the track morphology and sedimentary structures does not appear to support the original interpretation of the Lark Quarry tracksite preserving the movements of an Allosaurus-sized theropod in pursuit of prey, the mixed herding of small-bodied theropod and ornithopod dinosaurs, or the occurrence of a stampede. Comparative and multivariate data supports the reinterpretation of the large track-maker as an ornithopod, with the tracks assigned to Amblydactylus cf. A. gethingi. Three-dimensional track morphology shows that the small, purportedly theropod prints assigned to Skartopus australis are preservational variants of the ornithopod tracks assigned to Wintonopus latomorum. These re-evaluations remove any evidence for the presence of theropods in the Lark Quarry ichnofauna. The presence of asymmetrical ripples, vegetation drag marks, small dinosaur swimming tracks, and a footprint of a small dinosaur standing in flowing water indicate that the tracks were preserved in a fluvial environment. Superposition of tracks and other sedimentary structures suggests that the Amblydactylus cf. A. gethingi trackmaker moved across the ancient Lark Quarry site earlier than the Wintonopus trackmakers, such that the two trackmaker species probably did not interact.","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[],"research_interests":[],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-2373415-figures'); } }); </script> <div class="js-work-strip profile--work_container" data-work-id="2373413"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" rel="nofollow" href="https://www.academia.edu/2373413/Re_examination_of_the_Lark_Quarry_dinosaur_tracksite_reveals_an_absence_of_tracks_referable_to_nonavian_theropods_Romilio_A_and_S_Salisbury"><img alt="Research paper thumbnail of Re-examination of the Lark Quarry dinosaur tracksite reveals an absence of tracks referable to nonavian theropods. Romilio, A., and S. Salisbury" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"> Re-examination of the Lark Quarry dinosaur tracksite reveals an absence of tracks referable to nonavian theropods. Romilio, A., and S. Salisbury</div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">"The late Albian-Cenomanian Lark Quarry dinosaur tracksite near Winton, central-western Queenslan...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">"The late Albian-Cenomanian Lark Quarry dinosaur tracksite near Winton, central-western Queensland has previously been interpreted as preserving trackways indicative of two types of theropods and a single type of ornithopod dinosaur, represented by the ichnotaxa cf. Tyranosauropus, Skartopus australis and Wintonopus latomorum, respectively. The purported occurrence of theropod prints enabled the original researchers of Lark Quarry to speculate on possible behavioural interactions between dinosaur species: the stalking of prey by a large Allosaurus-sized theropod, and the mixed herding of small theropods with small-to-medium sized ornithopod dinosaurs. <br /> <br />Recent evidence indicates a large ornithopod dinosaur akin to Muttaburrasaurus made the large tracks that were originally assigned to cf. Tyrannosauropus, with these tracks re-interpreted as Amblydactylus cf. A. gethingi. Further analysis of the smaller tracks at Lark Quarry indicates that Skartopus australis tracks can co-occur with those of Wintonopus latomorum, both within individual trackways and within singular footprints. We conclude that Skartopus australis is a junior synonym of the ornithopod ichnotaxa Wintonopus latomorum. The range of variability displayed by Wintonopus tracks can be explained in terms of the dynamic interaction between the track-maker’s foot and underlying substrate. It thus appears that theropods were absent from the Lark Quarry dinosaur tracksite. There is consequently no evidence for the occurrence of large (Allosaurus-sized or larger) theropods in Australia during the mid-Cretaceous. Nor does Lark Quarry preserve any evidence for the mixed herding of small-bodied theropods with ornithopods. Previous claims for the occurrence of a ‘stampede’ at Lark Quarry need to be reassessed in light of these results. "</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="2373413"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="2373413"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 2373413; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=2373413]").text(description); $(".js-view-count[data-work-id=2373413]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 2373413; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='2373413']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=2373413]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":2373413,"title":" Re-examination of the Lark Quarry dinosaur tracksite reveals an absence of tracks referable to nonavian theropods. Romilio, A., and S. Salisbury","translated_title":"","metadata":{"abstract":"\"The late Albian-Cenomanian Lark Quarry dinosaur tracksite near Winton, central-western Queensland has previously been interpreted as preserving trackways indicative of two types of theropods and a single type of ornithopod dinosaur, represented by the ichnotaxa cf. Tyranosauropus, Skartopus australis and Wintonopus latomorum, respectively. The purported occurrence of theropod prints enabled the original researchers of Lark Quarry to speculate on possible behavioural interactions between dinosaur species: the stalking of prey by a large Allosaurus-sized theropod, and the mixed herding of small theropods with small-to-medium sized ornithopod dinosaurs. \r\n\r\nRecent evidence indicates a large ornithopod dinosaur akin to Muttaburrasaurus made the large tracks that were originally assigned to cf. Tyrannosauropus, with these tracks re-interpreted as Amblydactylus cf. A. gethingi. Further analysis of the smaller tracks at Lark Quarry indicates that Skartopus australis tracks can co-occur with those of Wintonopus latomorum, both within individual trackways and within singular footprints. We conclude that Skartopus australis is a junior synonym of the ornithopod ichnotaxa Wintonopus latomorum. The range of variability displayed by Wintonopus tracks can be explained in terms of the dynamic interaction between the track-maker’s foot and underlying substrate. It thus appears that theropods were absent from the Lark Quarry dinosaur tracksite. There is consequently no evidence for the occurrence of large (Allosaurus-sized or larger) theropods in Australia during the mid-Cretaceous. Nor does Lark Quarry preserve any evidence for the mixed herding of small-bodied theropods with ornithopods. Previous claims for the occurrence of a ‘stampede’ at Lark Quarry need to be reassessed in light of these results. \" ","location":"Perth, Australia","event_date":{"day":28,"month":4,"year":2011,"errors":{}},"journal_name":"Geological Survey of Western Australia 2011, CAVEPS Perth 2011, 13th Conference on Australasian Vertebrate Evolution Palaeontology and Systematics"},"translated_abstract":"\"The late Albian-Cenomanian Lark Quarry dinosaur tracksite near Winton, central-western Queensland has previously been interpreted as preserving trackways indicative of two types of theropods and a single type of ornithopod dinosaur, represented by the ichnotaxa cf. Tyranosauropus, Skartopus australis and Wintonopus latomorum, respectively. The purported occurrence of theropod prints enabled the original researchers of Lark Quarry to speculate on possible behavioural interactions between dinosaur species: the stalking of prey by a large Allosaurus-sized theropod, and the mixed herding of small theropods with small-to-medium sized ornithopod dinosaurs. \r\n\r\nRecent evidence indicates a large ornithopod dinosaur akin to Muttaburrasaurus made the large tracks that were originally assigned to cf. Tyrannosauropus, with these tracks re-interpreted as Amblydactylus cf. A. gethingi. Further analysis of the smaller tracks at Lark Quarry indicates that Skartopus australis tracks can co-occur with those of Wintonopus latomorum, both within individual trackways and within singular footprints. We conclude that Skartopus australis is a junior synonym of the ornithopod ichnotaxa Wintonopus latomorum. The range of variability displayed by Wintonopus tracks can be explained in terms of the dynamic interaction between the track-maker’s foot and underlying substrate. It thus appears that theropods were absent from the Lark Quarry dinosaur tracksite. There is consequently no evidence for the occurrence of large (Allosaurus-sized or larger) theropods in Australia during the mid-Cretaceous. Nor does Lark Quarry preserve any evidence for the mixed herding of small-bodied theropods with ornithopods. Previous claims for the occurrence of a ‘stampede’ at Lark Quarry need to be reassessed in light of these results. \" ","internal_url":"https://www.academia.edu/2373413/Re_examination_of_the_Lark_Quarry_dinosaur_tracksite_reveals_an_absence_of_tracks_referable_to_nonavian_theropods_Romilio_A_and_S_Salisbury","translated_internal_url":"","created_at":"2013-01-07T12:52:43.485-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":379100,"coauthors_can_edit":true,"document_type":"conference_presentation","co_author_tags":[],"downloadable_attachments":[],"slug":"Re_examination_of_the_Lark_Quarry_dinosaur_tracksite_reveals_an_absence_of_tracks_referable_to_nonavian_theropods_Romilio_A_and_S_Salisbury","translated_slug":"","page_count":null,"language":"en","content_type":"Work","summary":"\"The late Albian-Cenomanian Lark Quarry dinosaur tracksite near Winton, central-western Queensland has previously been interpreted as preserving trackways indicative of two types of theropods and a single type of ornithopod dinosaur, represented by the ichnotaxa cf. Tyranosauropus, Skartopus australis and Wintonopus latomorum, respectively. The purported occurrence of theropod prints enabled the original researchers of Lark Quarry to speculate on possible behavioural interactions between dinosaur species: the stalking of prey by a large Allosaurus-sized theropod, and the mixed herding of small theropods with small-to-medium sized ornithopod dinosaurs. \r\n\r\nRecent evidence indicates a large ornithopod dinosaur akin to Muttaburrasaurus made the large tracks that were originally assigned to cf. Tyrannosauropus, with these tracks re-interpreted as Amblydactylus cf. A. gethingi. Further analysis of the smaller tracks at Lark Quarry indicates that Skartopus australis tracks can co-occur with those of Wintonopus latomorum, both within individual trackways and within singular footprints. We conclude that Skartopus australis is a junior synonym of the ornithopod ichnotaxa Wintonopus latomorum. The range of variability displayed by Wintonopus tracks can be explained in terms of the dynamic interaction between the track-maker’s foot and underlying substrate. It thus appears that theropods were absent from the Lark Quarry dinosaur tracksite. There is consequently no evidence for the occurrence of large (Allosaurus-sized or larger) theropods in Australia during the mid-Cretaceous. Nor does Lark Quarry preserve any evidence for the mixed herding of small-bodied theropods with ornithopods. Previous claims for the occurrence of a ‘stampede’ at Lark Quarry need to be reassessed in light of these results. \" ","owner":{"id":379100,"first_name":"Anthony","middle_initials":null,"last_name":"Romilio","page_name":"AnthonyRomilio","domain_name":"uq","created_at":"2011-03-27T21:19:32.416-07:00","display_name":"Anthony Romilio","url":"https://uq.academia.edu/AnthonyRomilio"},"attachments":[],"research_interests":[],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") if (false) { Aedu.setUpFigureCarousel('profile-work-2373413-figures'); } }); </script> </div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-a9bf3a2bc8c89fa2a77156577594264ee8a0f214d74241bc0fcd3f69f8d107ac.js","https://a.academia-assets.com/assets/google_contacts-0dfb882d836b94dbcb4a2d123d6933fc9533eda5be911641f20b4eb428429600.js"], function() { // from javascript_helper.rb $('.js-google-connect-button').click(function(e) { e.preventDefault(); 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