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* <a href="/wiki/8._November" title="8. November">8. November</a> <a href="/wiki/1946" title="1946">1946</a> in <a href="/wiki/Moskau" title="Moskau">Moskau</a>) ist ein <a href="/wiki/Russland" title="Russland">russisch</a>-<a href="/wiki/Vereinigte_Staaten" title="Vereinigte Staaten">US-amerikanischer</a> <a href="/wiki/Biochemie" title="Biochemie">Biochemiker</a> und Professor am <a href="/wiki/California_Institute_of_Technology" title="California Institute of Technology">California Institute of Technology</a>. </p> <div id="toc" class="toc" role="navigation" aria-labelledby="mw-toc-heading"><input type="checkbox" role="button" id="toctogglecheckbox" class="toctogglecheckbox" style="display:none" /><div class="toctitle" lang="de" dir="ltr"><h2 id="mw-toc-heading">Inhaltsverzeichnis</h2><span class="toctogglespan"><label class="toctogglelabel" for="toctogglecheckbox"></label></span></div> <ul> <li class="toclevel-1 tocsection-1"><a href="#Leben"><span class="tocnumber">1</span> <span class="toctext">Leben</span></a></li> <li class="toclevel-1 tocsection-2"><a href="#Wirken"><span class="tocnumber">2</span> <span class="toctext">Wirken</span></a></li> <li class="toclevel-1 tocsection-3"><a href="#Auszeichnungen_(Auswahl)"><span class="tocnumber">3</span> <span class="toctext">Auszeichnungen (Auswahl)</span></a></li> <li class="toclevel-1 tocsection-4"><a href="#Weblinks"><span class="tocnumber">4</span> <span class="toctext">Weblinks</span></a></li> <li class="toclevel-1 tocsection-5"><a href="#Einzelnachweise"><span class="tocnumber">5</span> <span class="toctext">Einzelnachweise</span></a></li> </ul> </div> <div class="mw-heading mw-heading2"><h2 id="Leben">Leben</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Alexander_Varshavsky&veaction=edit&section=1" title="Abschnitt bearbeiten: Leben" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Alexander_Varshavsky&action=edit&section=1" title="Quellcode des Abschnitts bearbeiten: Leben"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Varshavsky erwarb 1970 einen <a href="/wiki/Bachelor" title="Bachelor">Bachelor</a> in Chemie an der <a href="/wiki/Universit%C3%A4t_Moskau" class="mw-redirect" title="Universität Moskau">Universität Moskau</a> und 1973 einen <a href="/wiki/Ph.D." title="Ph.D.">Ph.D.</a> für Biochemie am Institut für <a href="/wiki/Molekulare_Biologie" class="mw-redirect" title="Molekulare Biologie">Molekulare Biologie</a> in Moskau. Von 1973 bis 1976 war er dort Forschungsassistent. 1977 emigrierte Varshavsky in die USA, wo er im <i>Department of Biology</i> am <a href="/wiki/Massachusetts_Institute_of_Technology" title="Massachusetts Institute of Technology">Massachusetts Institute of Technology</a> zunächst eine Juniorprofessur übernahm (1977 <i>Assistant Professor</i>, 1980 <i>Associate Professor</i>) und 1986 ordentlicher Professor wurde. 1992 wechselte er an das <a href="/wiki/California_Institute_of_Technology" title="California Institute of Technology">California Institute of Technology</a>, wo er <i>Howard and Gwen Laurie Smits Professor</i> für <a href="/wiki/Zellbiologie" title="Zellbiologie">Zellbiologie</a> ist. </p> <div class="mw-heading mw-heading2"><h2 id="Wirken">Wirken</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Alexander_Varshavsky&veaction=edit&section=2" title="Abschnitt bearbeiten: Wirken" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Alexander_Varshavsky&action=edit&section=2" title="Quellcode des Abschnitts bearbeiten: Wirken"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Mittels genetischer Untersuchungen an <a href="/wiki/Hefen" title="Hefen">Hefen</a> und <a href="/wiki/Zellkultur" title="Zellkultur">Zellkulturen</a> höherer Organismen konnte Varshavsky zahlreiche Bedeutungen des <a href="/wiki/Ubiquitin" title="Ubiquitin">Ubiquitin</a>-Systems beim Abbau von <a href="/wiki/Protein" title="Protein">Proteinen</a> nach der <i><a href="/wiki/N-End_Rule" title="N-End Rule">N-End Rule</a></i> aufklären,<sup id="cite_ref-gairdner_1-0" class="reference"><a href="#cite_note-gairdner-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> die eine wichtige Rolle im <a href="/wiki/Zellzyklus" title="Zellzyklus">Zellzyklus</a>, in der <a href="/wiki/Apoptose" title="Apoptose">Apoptose</a>, in der <a href="/wiki/Maligne_Transformation" title="Maligne Transformation">malignen Transformation</a>, der <a href="/wiki/Entz%C3%BCndung" title="Entzündung">Entzündungsregulation</a> und der <a href="/wiki/Immunantwort" title="Immunantwort">Immunantwort</a> spielt.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Auszeichnungen_(Auswahl)"><span id="Auszeichnungen_.28Auswahl.29"></span>Auszeichnungen (Auswahl)</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Alexander_Varshavsky&veaction=edit&section=3" title="Abschnitt bearbeiten: Auszeichnungen (Auswahl)" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Alexander_Varshavsky&action=edit&section=3" title="Quellcode des Abschnitts bearbeiten: Auszeichnungen (Auswahl)"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <div style="column-width:30em"> <ul><li>1987 Mitgliedschaft in der <a href="/wiki/American_Academy_of_Arts_and_Sciences" title="American Academy of Arts and Sciences">American Academy of Arts and Sciences</a><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></li> <li>1995 Mitgliedschaft in der <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">National Academy of Sciences</a></li> <li>1999 <a href="/wiki/Gairdner_Foundation_International_Award" class="mw-redirect" title="Gairdner Foundation International Award">Gairdner Foundation International Award</a><sup id="cite_ref-gairdner_1-1" class="reference"><a href="#cite_note-gairdner-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></li> <li>2000 <a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award for Basic Medical Research</a></li> <li>2000 <a href="/wiki/Alfred_P._Sloan,_Jr._Prize" title="Alfred P. Sloan, Jr. Prize">Alfred P. Sloan, Jr. Prize</a></li> <li>2001 <a href="/wiki/Pasarow_Award" title="Pasarow Award">Pasarow Award</a></li> <li>2001 <a href="/wiki/Max-Planck-Forschungspreis" title="Max-Planck-Forschungspreis">Max-Planck-Forschungspreis</a></li> <li>2001 <a href="/wiki/Louisa-Gross-Horwitz-Preis" title="Louisa-Gross-Horwitz-Preis">Louisa-Gross-Horwitz-Preis</a></li> <li>2001 Mitgliedschaft in der <a href="/wiki/American_Philosophical_Society" title="American Philosophical Society">American Philosophical Society</a></li> <li>2001 <a href="/wiki/Wolf-Preis" title="Wolf-Preis">Wolf-Preis</a> in Medizin</li> <li>2001 <a href="/wiki/Massry-Preis" title="Massry-Preis">Massry-Preis</a></li> <li>2001 Ausländisches Mitglied der <a href="/wiki/European_Molecular_Biology_Organization" title="European Molecular Biology Organization">European Molecular Biology Organization</a></li> <li>2002 <a href="/wiki/E._B._Wilson_Medal" title="E. B. Wilson Medal">E. B. Wilson Medal</a></li> <li>2002 Mitgliedschaft in der <a href="/wiki/American_Association_for_the_Advancement_of_Science" title="American Association for the Advancement of Science">American Association for the Advancement of Science</a></li> <li>2005 Auswärtiges Mitglied der <a href="/wiki/Academia_Europaea" title="Academia Europaea">Academia Europaea</a><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></li> <li>2006 <a href="/wiki/L%C3%A9opold_Griffuel-Preis" class="mw-redirect" title="Léopold Griffuel-Preis">Léopold Griffuel-Preis</a></li> <li>2006 <a href="/wiki/March_of_Dimes_Prize_in_Developmental_Biology" class="mw-redirect" title="March of Dimes Prize in Developmental Biology">March of Dimes Prize in Developmental Biology</a></li> <li>2007 <a href="/wiki/Schleiden-Medaille" title="Schleiden-Medaille">Schleiden-Medaille</a></li> <li>2011 <a href="/wiki/BBVA_Foundation_Frontiers_of_Knowledge_Awards" title="BBVA Foundation Frontiers of Knowledge Awards">BBVA Foundation Frontiers of Knowledge Awards</a></li> <li>2012 <a href="/wiki/Otto-Warburg-Medaille" title="Otto-Warburg-Medaille">Otto-Warburg-Medaille</a></li> <li>2012 <a href="/wiki/K%C3%B6nig-Faisal-Preis" class="mw-redirect" title="König-Faisal-Preis">König-Faisal-Preis</a> für Wissenschaft</li> <li>2014 <a href="/wiki/Breakthrough_Prize_in_Life_Sciences" title="Breakthrough Prize in Life Sciences">Breakthrough Prize in Life Sciences</a></li> <li>2014 <a href="/wiki/Albany_Medical_Center_Prize" title="Albany Medical Center Prize">Albany Medical Center Prize</a></li> <li>2016 <a href="/wiki/Grande_m%C3%A9daille_de_l%E2%80%99Acad%C3%A9mie_des_sciences" title="Grande médaille de l’Académie des sciences">Grande médaille de l’Académie des sciences</a></li> <li>2017 <a href="/wiki/Heinrich-Wieland-Preis" title="Heinrich-Wieland-Preis">Heinrich-Wieland-Preis</a></li> <li>2024 <a href="/wiki/Dr._Paul_Janssen_Award_for_Biomedical_Research" title="Dr. Paul Janssen Award for Biomedical Research">Dr. Paul Janssen Award for Biomedical Research</a></li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="Weblinks">Weblinks</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Alexander_Varshavsky&veaction=edit&section=4" title="Abschnitt bearbeiten: Weblinks" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Alexander_Varshavsky&action=edit&section=4" title="Quellcode des Abschnitts bearbeiten: Weblinks"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="https://books.google.de/books?id=bDHwCgAAQBAJ&pg=PA311#v=onepage"><i>Alexander Varshavsky.</i> Interview mit Istvan Hargittai. Imperial College Press (2006)</a> in der Google-Buchsuche</li> <li><a rel="nofollow" class="external text" href="http://s3-us-west-1.amazonaws.com/bbe-prod-storage.cloud.caltech.edu/CVs/CV-Varshavsky.pdf">Lebenslauf</a> (PDF; 121 kB) auf biology.caltech.edu; abgerufen am 23. Januar 2016.</li> <li><a rel="nofollow" class="external text" href="https://academictree.org/chemistry/peopleinfo.php?pid=57112">Informationen</a> zu und <a rel="nofollow" class="external text" href="https://academictree.org/chemistry/tree.php?pid=57112">akademischer Stammbaum</a> von <i>Alexander Varshavsky</i> bei academictree.org<span class="editoronly" style="display:none;"></span></li></ul> <div class="mw-heading mw-heading2"><h2 id="Einzelnachweise">Einzelnachweise</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Alexander_Varshavsky&veaction=edit&section=5" title="Abschnitt bearbeiten: Einzelnachweise" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Alexander_Varshavsky&action=edit&section=5" title="Quellcode des Abschnitts bearbeiten: Einzelnachweise"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <ol class="references"> <li id="cite_note-gairdner-1"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-gairdner_1-0">a</a></sup> <sup><a href="#cite_ref-gairdner_1-1">b</a></sup></span> <span class="reference-text"><span class="cite"><a rel="nofollow" class="external text" href="https://gairdner.org/award_winners/alexander-j-varshavsky/"><i>Alexander J. Varshavsky – Gairdner Foundation.</i></a> In: <i>gairdner.org.</i><span class="Abrufdatum"> Abgerufen am 29. März 2018</span> (englisch).</span><span style="display: none;" class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adc&rfr_id=info%3Asid%2Fde.wikipedia.org%3AAlexander+Varshavsky&rft.title=Alexander+J.+Varshavsky+%E2%80%93+Gairdner+Foundation&rft.description=Alexander+J.+Varshavsky+%E2%80%93+Gairdner+Foundation&rft.identifier=https%3A%2F%2Fgairdner.org%2Faward_winners%2Falexander-j-varshavsky%2F&rft.date=&rft.language=en"> </span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><span class="cite">Mark Hofschneider: <a rel="nofollow" class="external text" href="https://laskerfoundation.org/winners/ubiquitin-system-for-regulated-protein-degradation/"><i>Ubiquitin system for regulated protein degradation.</i></a> In: <i>laskerfoundation.org.</i><span class="Abrufdatum"> Abgerufen am 1. März 2023</span> (englisch).</span><span style="display: none;" class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adc&rfr_id=info%3Asid%2Fde.wikipedia.org%3AAlexander+Varshavsky&rft.title=Ubiquitin+system+for+regulated+protein+degradation&rft.description=Ubiquitin+system+for+regulated+protein+degradation&rft.identifier=https%3A%2F%2Flaskerfoundation.org%2Fwinners%2Fubiquitin-system-for-regulated-protein-degradation%2F&rft.creator=Mark+Hofschneider&rft.language=en"> </span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text"><span class="cite"><a rel="nofollow" class="external text" href="https://www.amacad.org/multimedia/pdfs/publications/bookofmembers/ChapterV.pdf"><i>Book of Members 1780–present, Chapter V.</i></a> (PDF; 281 kB) In: <i>amacad.org.</i> <a href="/wiki/American_Academy_of_Arts_and_Sciences" title="American Academy of Arts and Sciences">American Academy of Arts and Sciences</a>,<span class="Abrufdatum"> abgerufen am 29. März 2018</span> (englisch).</span><span style="display: none;" class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adc&rfr_id=info%3Asid%2Fde.wikipedia.org%3AAlexander+Varshavsky&rft.title=Book+of+Members+1780%E2%80%93present%2C+Chapter+V&rft.description=Book+of+Members+1780%E2%80%93present%2C+Chapter+V&rft.identifier=https%3A%2F%2Fwww.amacad.org%2Fmultimedia%2Fpdfs%2Fpublications%2Fbookofmembers%2FChapterV.pdf&rft.publisher=%5B%5BAmerican+Academy+of+Arts+and+Sciences%5D%5D&rft.language=en"> </span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.ae-info.org/ae/Member/Varshavsky_Alexander">Eintrag</a> auf der Internetseite der Academia Europaea</span> </li> </ol> <style data-mw-deduplicate="TemplateStyles:r248673343">.mw-parser-output div.NavFrame{border-width:1px;border-style:solid;border-left-color:var(--dewiki-rahmenfarbe1);border-right-color:var(--dewiki-rahmenfarbe1);border-top-color:var(--dewiki-rahmenfarbe1);border-bottom-color:var(--dewiki-rahmenfarbe1);clear:both;font-size:95%;margin-top:1.5em;min-height:0;padding:2px;text-align:center}.mw-parser-output div.NavPic{float:left;padding:2px}.mw-parser-output div.NavHead{background-color:var(--dewiki-hintergrundfarbe5);font-weight:bold}.mw-parser-output div.NavFrame:after{clear:both;content:"";display:block}.mw-parser-output div.NavFrame+div.NavFrame,.mw-parser-output div.NavFrame+link+div.NavFrame,.mw-parser-output div.NavFrame+style+div.NavFrame{margin-top:-1px}.mw-parser-output .NavToggle{float:right;font-size:x-small}@media screen{html.skin-theme-clientpref-night .mw-parser-output .NavPic span[typeof="mw:File"] img{background-color:#c8ccd1}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .NavPic span[typeof="mw:File"] img{background-color:#c8ccd1}}</style><div class="NavFrame navigation-not-searchable" role="navigation"> <div class="NavHead">Träger des <a href="/wiki/Wolf-Preis" title="Wolf-Preis">Wolf-Preises</a> in Medizin</div> <div class="NavContent"> <p>1978: <a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Davis Snell</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a>, <a href="/wiki/Jon_van_Rood" title="Jon van Rood">Jon van Rood</a> | 1979: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Roger Sperry</a>, <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a>, <a href="/wiki/Oleh_Hornykiewicz" title="Oleh Hornykiewicz">Oleh Hornykiewicz</a> | 1980: <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a>, <a href="/wiki/Leo_Sachs" title="Leo Sachs">Leo Sachs</a>, <a href="/wiki/James_L._Gowans" title="James L. Gowans">James L. Gowans</a> | 1981: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a>, <a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a> | 1982: <a href="/wiki/Jean-Pierre_Changeux" title="Jean-Pierre Changeux">Jean-Pierre Changeux</a>, <a href="/wiki/Solomon_H._Snyder" title="Solomon H. Snyder">Solomon H. Snyder</a>, <a href="/wiki/James_Whyte_Black" title="James Whyte Black">James Whyte Black</a> | 1983/4: <i>nicht vergeben</i> | 1984/5: <a href="/wiki/Donald_F._Steiner" title="Donald F. Steiner">Donald F. Steiner</a> | 1986: <a href="/wiki/Hayaishi_Osamu" title="Hayaishi Osamu">Hayaishi Osamu</a> | 1987: <a href="/wiki/Pedro_Cuatrecasas" title="Pedro Cuatrecasas">Pedro Cuatrecasas</a>, <a href="/wiki/Meir_Wilchek" title="Meir Wilchek">Meir Wilchek</a> | 1988: <a href="/wiki/Henri-G%C3%A9ry_Hers" title="Henri-Géry Hers">Henri-Géry Hers</a>, <a href="/wiki/Elizabeth_F._Neufeld" title="Elizabeth F. Neufeld">Elizabeth F. Neufeld</a> | 1989: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a>, <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> | 1990: <a href="/wiki/Maclyn_McCarty" title="Maclyn McCarty">Maclyn McCarty</a> | 1991: <a href="/wiki/Seymour_Benzer" title="Seymour Benzer">Seymour Benzer</a> | 1992: <a href="/wiki/Judah_Folkman" title="Judah Folkman">Judah Folkman</a> | 1993: <i>nicht vergeben</i> | 1994/5: <a href="/wiki/Michael_Berridge" title="Michael Berridge">Michael Berridge</a>, <a href="/wiki/Yasutomi_Nishizuka" title="Yasutomi Nishizuka">Yasutomi Nishizuka</a> | 1995/6: <a href="/wiki/Stanley_Prusiner" title="Stanley Prusiner">Stanley Prusiner</a> | 1996/7: <a href="/wiki/Mary_Frances_Lyon" title="Mary Frances Lyon">Mary Frances Lyon</a> | 1998: <a href="/wiki/Michael_Sela" title="Michael Sela">Michael Sela</a>, <a href="/wiki/Ruth_Arnon" title="Ruth Arnon">Ruth Arnon</a> | 1999: <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a> | 2000: <i>nicht vergeben</i> | 2001: <a href="/wiki/Avram_Hershko" title="Avram Hershko">Avram Hershko</a>, <a class="mw-selflink selflink">Alexander Varshavsky</a> | 2002/3: <a href="/wiki/Ralph_L._Brinster" title="Ralph L. Brinster">Ralph L. Brinster</a>, <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a> | 2004: <a href="/wiki/Robert_Allan_Weinberg" title="Robert Allan Weinberg">Robert Allan Weinberg</a>, <a href="/wiki/Roger_Tsien" title="Roger Tsien">Roger Tsien</a> | 2005: <a href="/wiki/Tony_Hunter" title="Tony Hunter">Tony Hunter</a>, <a href="/wiki/Anthony_Pawson" title="Anthony Pawson">Anthony Pawson</a>, <a href="/wiki/Alexander_Levitzki" title="Alexander Levitzki">Alexander Levitzki</a> | 2006/7: <i>nicht vergeben</i> | 2008: <a href="/wiki/Howard_Cedar" title="Howard Cedar">Howard Cedar</a>, <a href="/wiki/Aharon_Razin" title="Aharon Razin">Aharon Razin</a> | 2009: <i>nicht vergeben</i> | 2010: <a href="/wiki/Axel_Ullrich" title="Axel Ullrich">Axel Ullrich</a> | 2011: <a href="/wiki/Shin%E2%80%99ya_Yamanaka" title="Shin’ya Yamanaka">Shin’ya Yamanaka</a>, <a href="/wiki/Rudolf_Jaenisch" title="Rudolf Jaenisch">Rudolf Jaenisch</a> | 2012: <a href="/wiki/Ronald_M._Evans" title="Ronald M. Evans">Ronald M. Evans</a> | 2013: <i>nicht vergeben</i> | 2014: <a href="/wiki/Nahum_Sonenberg" title="Nahum Sonenberg">Nahum Sonenberg</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a>, <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> | 2015: <a href="/wiki/John_W._Kappler" title="John W. Kappler">John W. Kappler</a>, <a href="/wiki/Philippa_Marrack" title="Philippa Marrack">Philippa Marrack</a>, <a href="/wiki/Jeffrey_Ravetch" title="Jeffrey Ravetch">Jeffrey Ravetch</a> | 2016: <a href="/wiki/C._Ronald_Kahn" title="C. Ronald Kahn">C. Ronald Kahn</a>, <a href="/wiki/Lewis_C._Cantley" title="Lewis C. Cantley">Lewis C. Cantley</a> | 2017: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> | 2018: <i>nicht vergeben</i> | 2019: <a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey M. Friedman</a> | 2020: <a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a>, <a href="/wiki/Jennifer_A._Doudna" title="Jennifer A. Doudna">Jennifer A. Doudna</a> | 2021: <a href="/wiki/Joan_A._Steitz" title="Joan A. Steitz">Joan A. Steitz</a>, <a href="/wiki/Lynne_E._Maquat" title="Lynne E. Maquat">Lynne E. Maquat</a>, <a href="/wiki/Adrian_R._Krainer" title="Adrian R. Krainer">Adrian R. Krainer</a> | 2022: <i>nicht vergeben</i> | 2023: <a href="/wiki/Daniel_J._Drucker" title="Daniel J. Drucker">Daniel J. Drucker</a> | 2024: <a href="/wiki/Botond_Roska" title="Botond Roska">Botond Roska</a>, <a href="/wiki/Jos%C3%A9-Alain_Sahel" title="José-Alain Sahel">José-Alain Sahel</a> </p> </div> </div> <div class="hintergrundfarbe1 rahmenfarbe1 navigation-not-searchable normdaten-typ-p" style="border-style: solid; border-width: 1px; clear: left; margin-bottom:1em; margin-top:1em; padding: 0.25em; overflow: hidden; word-break: break-word; word-wrap: break-word;" id="normdaten"> <div style="display: table-cell; vertical-align: middle; width: 100%;"> <div> Normdaten (Person): <a href="/wiki/Gemeinsame_Normdatei" title="Gemeinsame Normdatei">GND</a>: <span class="plainlinks-print"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/1355574730">1355574730</a></span> <span class="noprint">(<a rel="nofollow" class="external text" href="https://lobid.org/gnd/1355574730">lobid</a>, <a rel="nofollow" class="external text" href="https://swb.bsz-bw.de/DB=2.104/SET=1/TTL=1/CMD?retrace=0&trm_old=&ACT=SRCHA&IKT=2999&SRT=RLV&TRM=1355574730">OGND</a><span class="metadata">, <a rel="nofollow" class="external text" href="https://prometheus.lmu.de/gnd/1355574730">AKS</a></span>)</span> | <a href="/wiki/Library_of_Congress_Control_Number" title="Library of Congress Control Number">LCCN</a>: <span class="plainlinks-print"><a rel="nofollow" class="external text" href="https://lccn.loc.gov/no2009052096">no2009052096</a></span> | <a href="/wiki/Virtual_International_Authority_File" title="Virtual International Authority File">VIAF</a>: <span class="plainlinks-print"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/311250147/">311250147</a></span> | <span class="noprint"><a class="external text" href="https://persondata.toolforge.org/p/Alexander_Varshavsky">Wikipedia-Personensuche</a></span><span class="metadata"></span></div> </div></div> <table class="metadata rahmenfarbe1" id="Vorlage_Personendaten" style="border-style: solid; margin-top: 20px;"> <tbody><tr> <th colspan="2"><a href="/wiki/Hilfe:Personendaten" title="Hilfe:Personendaten">Personendaten</a> </th></tr> <tr> <td style="color: #aaa;">NAME </td> <td style="font-weight: bold;">Varshavsky, Alexander </td></tr> <tr> <td style="color: #aaa;">ALTERNATIVNAMEN </td> <td>Варшавский, Александр Яковлевич (russisch); Varshavsky, Alexander J. </td></tr> <tr> <td style="color: #aaa;">KURZBESCHREIBUNG </td> <td>russisch-amerikanischer Biochemiker </td></tr> <tr> <td style="color: #aaa;">GEBURTSDATUM </td> <td>8. November 1946 </td></tr> <tr> <td style="color: #aaa;">GEBURTSORT </td> <td><a href="/wiki/Moskau" title="Moskau">Moskau</a> </td></tr> </tbody></table></div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?useformat=desktop&type=1x1&usesul3=0" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Abgerufen von „<a dir="ltr" href="https://de.wikipedia.org/w/index.php?title=Alexander_Varshavsky&oldid=253066379">https://de.wikipedia.org/w/index.php?title=Alexander_Varshavsky&oldid=253066379</a>“</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Wikipedia:Kategorien" title="Wikipedia:Kategorien">Kategorien</a>: <ul><li><a href="/wiki/Kategorie:Biochemiker" title="Kategorie:Biochemiker">Biochemiker</a></li><li><a href="/wiki/Kategorie:Hochschullehrer_(Cambridge,_Massachusetts)" 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hreflang="ar" data-title="إلكسندر فارشافسكي" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%A7%D9%84%D9%83%D8%B3%D9%86%D8%AF%D8%B1_%D9%81%D8%A7%D8%B1%D8%B4%D8%A7%D9%81%D8%B3%D9%83%D9%89" title="الكسندر فارشافسكى – Ägyptisches Arabisch" lang="arz" hreflang="arz" data-title="الكسندر فارشافسكى" data-language-autonym="مصرى" data-language-local-name="Ägyptisches Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%90%D0%BB%D1%8F%D0%BA%D1%81%D0%B0%D0%BD%D0%B4%D1%80_%D0%AF%D0%BA%D0%B0%D1%9E%D0%BB%D0%B5%D0%B2%D1%96%D1%87_%D0%92%D0%B0%D1%80%D1%88%D0%B0%D1%9E%D1%81%D0%BA%D1%96" title="Аляксандр Якаўлевіч Варшаўскі – Belarussisch" lang="be" hreflang="be" data-title="Аляксандр Якаўлевіч Варшаўскі" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Alexander_Varshavsky" title="Alexander Varshavsky – Englisch" lang="en" hreflang="en" data-title="Alexander Varshavsky" data-language-autonym="English" data-language-local-name="Englisch" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Alexander_Varshavsky" title="Alexander Varshavsky – Französisch" lang="fr" hreflang="fr" data-title="Alexander Varshavsky" data-language-autonym="Français" data-language-local-name="Französisch" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Alexander_Varshavsky" title="Alexander Varshavsky – Italienisch" lang="it" hreflang="it" data-title="Alexander Varshavsky" data-language-autonym="Italiano" data-language-local-name="Italienisch" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%A2%E3%83%AC%E3%82%AF%E3%82%B5%E3%83%B3%E3%83%80%E3%83%BC%E3%83%BB%E3%83%90%E3%83%BC%E3%82%B7%E3%83%A3%E3%83%95%E3%82%B9%E3%82%AD%E3%83%BC" title="アレクサンダー・バーシャフスキー – Japanisch" lang="ja" hreflang="ja" data-title="アレクサンダー・バーシャフスキー" data-language-autonym="日本語" data-language-local-name="Japanisch" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%85%E0%B4%B2%E0%B4%95%E0%B5%8D%E0%B4%B8%E0%B4%BE%E0%B4%A3%E0%B5%8D%E0%B4%9F%E0%B5%BC_%E0%B4%B5%E0%B5%BC%E0%B4%B7%E0%B4%B5%E0%B5%8D%E0%B4%B8%E0%B5%8D%E0%B4%95%E0%B4%BF" title="അലക്സാണ്ടർ വർഷവ്സ്കി – Malayalam" lang="ml" hreflang="ml" data-title="അലക്സാണ്ടർ വർഷവ്സ്കി" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Alexander_Varshavsky" title="Alexander Varshavsky – Portugiesisch" lang="pt" hreflang="pt" data-title="Alexander Varshavsky" data-language-autonym="Português" data-language-local-name="Portugiesisch" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%92%D0%B0%D1%80%D1%88%D0%B0%D0%B2%D1%81%D0%BA%D0%B8%D0%B9,_%D0%90%D0%BB%D0%B5%D0%BA%D1%81%D0%B0%D0%BD%D0%B4%D1%80_%D0%AF%D0%BA%D0%BE%D0%B2%D0%BB%D0%B5%D0%B2%D0%B8%D1%87" title="Варшавский, Александр Яковлевич – Russisch" lang="ru" hreflang="ru" data-title="Варшавский, Александр Яковлевич" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Alexander_Varshavsky" title="Alexander Varshavsky – einfaches Englisch" lang="en-simple" hreflang="en-simple" data-title="Alexander Varshavsky" data-language-autonym="Simple English" data-language-local-name="einfaches Englisch" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E4%BA%9A%E5%8E%86%E5%B1%B1%E5%A4%A7%C2%B7%E7%93%A6%E5%B0%94%E6%B2%99%E5%A4%AB%E6%96%AF%E5%9F%BA" title="亚历山大·瓦尔沙夫斯基 – Chinesisch" lang="zh" hreflang="zh" data-title="亚历山大·瓦尔沙夫斯基" data-language-autonym="中文" data-language-local-name="Chinesisch" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a 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