CINXE.COM
Systems ecology - Wikipedia
<!DOCTYPE html> <html class="client-nojs vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available" lang="en" dir="ltr"> <head> <meta charset="UTF-8"> <title>Systems ecology - Wikipedia</title> <script>(function(){var className="client-js vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available";var cookie=document.cookie.match(/(?:^|; )enwikimwclientpreferences=([^;]+)/);if(cookie){cookie[1].split('%2C').forEach(function(pref){className=className.replace(new RegExp('(^| )'+pref.replace(/-clientpref-\w+$|[^\w-]+/g,'')+'-clientpref-\\w+( |$)'),'$1'+pref+'$2');});}document.documentElement.className=className;}());RLCONF={"wgBreakFrames":false,"wgSeparatorTransformTable":["",""],"wgDigitTransformTable":["",""],"wgDefaultDateFormat":"dmy", "wgMonthNames":["","January","February","March","April","May","June","July","August","September","October","November","December"],"wgRequestId":"b6e66ce4-ad3f-4bc7-a644-7cb0e57c52ea","wgCanonicalNamespace":"","wgCanonicalSpecialPageName":false,"wgNamespaceNumber":0,"wgPageName":"Systems_ecology","wgTitle":"Systems ecology","wgCurRevisionId":1185361143,"wgRevisionId":1185361143,"wgArticleId":3253299,"wgIsArticle":true,"wgIsRedirect":false,"wgAction":"view","wgUserName":null,"wgUserGroups":["*"],"wgCategories":["CS1 maint: multiple names: authors list","CS1 maint: numeric names: authors list","Articles with short description","Short description matches Wikidata","Commons category link from Wikidata","Systems ecology","Environmental science","Environmental social science","Formal sciences","Systems theory"],"wgPageViewLanguage":"en","wgPageContentLanguage":"en","wgPageContentModel":"wikitext","wgRelevantPageName":"Systems_ecology","wgRelevantArticleId":3253299,"wgIsProbablyEditable":true, "wgRelevantPageIsProbablyEditable":true,"wgRestrictionEdit":[],"wgRestrictionMove":[],"wgNoticeProject":"wikipedia","wgCiteReferencePreviewsActive":false,"wgFlaggedRevsParams":{"tags":{"status":{"levels":1}}},"wgMediaViewerOnClick":true,"wgMediaViewerEnabledByDefault":true,"wgPopupsFlags":0,"wgVisualEditor":{"pageLanguageCode":"en","pageLanguageDir":"ltr","pageVariantFallbacks":"en"},"wgMFDisplayWikibaseDescriptions":{"search":true,"watchlist":true,"tagline":false,"nearby":true},"wgWMESchemaEditAttemptStepOversample":false,"wgWMEPageLength":10000,"wgRelatedArticlesCompat":[],"wgCentralAuthMobileDomain":false,"wgEditSubmitButtonLabelPublish":true,"wgULSPosition":"interlanguage","wgULSisCompactLinksEnabled":false,"wgVector2022LanguageInHeader":true,"wgULSisLanguageSelectorEmpty":false,"wgWikibaseItemId":"Q3076438","wgCheckUserClientHintsHeadersJsApi":["brands","architecture","bitness","fullVersionList","mobile","model","platform","platformVersion"],"GEHomepageSuggestedEditsEnableTopics": true,"wgGETopicsMatchModeEnabled":false,"wgGEStructuredTaskRejectionReasonTextInputEnabled":false,"wgGELevelingUpEnabledForUser":false};RLSTATE={"ext.globalCssJs.user.styles":"ready","site.styles":"ready","user.styles":"ready","ext.globalCssJs.user":"ready","user":"ready","user.options":"loading","ext.cite.styles":"ready","skins.vector.search.codex.styles":"ready","skins.vector.styles":"ready","skins.vector.icons":"ready","jquery.makeCollapsible.styles":"ready","ext.wikimediamessages.styles":"ready","ext.visualEditor.desktopArticleTarget.noscript":"ready","ext.uls.interlanguage":"ready","wikibase.client.init":"ready","ext.wikimediaBadges":"ready"};RLPAGEMODULES=["ext.cite.ux-enhancements","mediawiki.page.media","site","mediawiki.page.ready","jquery.makeCollapsible","mediawiki.toc","skins.vector.js","ext.centralNotice.geoIP","ext.centralNotice.startUp","ext.gadget.ReferenceTooltips","ext.gadget.switcher","ext.urlShortener.toolbar","ext.centralauth.centralautologin","mmv.bootstrap", "ext.popups","ext.visualEditor.desktopArticleTarget.init","ext.visualEditor.targetLoader","ext.echo.centralauth","ext.eventLogging","ext.wikimediaEvents","ext.navigationTiming","ext.uls.interface","ext.cx.eventlogging.campaigns","ext.cx.uls.quick.actions","wikibase.client.vector-2022","ext.checkUser.clientHints","ext.growthExperiments.SuggestedEditSession","wikibase.sidebar.tracking"];</script> <script>(RLQ=window.RLQ||[]).push(function(){mw.loader.impl(function(){return["user.options@12s5i",function($,jQuery,require,module){mw.user.tokens.set({"patrolToken":"+\\","watchToken":"+\\","csrfToken":"+\\"}); }];});});</script> <link rel="stylesheet" href="/w/load.php?lang=en&modules=ext.cite.styles%7Cext.uls.interlanguage%7Cext.visualEditor.desktopArticleTarget.noscript%7Cext.wikimediaBadges%7Cext.wikimediamessages.styles%7Cjquery.makeCollapsible.styles%7Cskins.vector.icons%2Cstyles%7Cskins.vector.search.codex.styles%7Cwikibase.client.init&only=styles&skin=vector-2022"> <script async="" src="/w/load.php?lang=en&modules=startup&only=scripts&raw=1&skin=vector-2022"></script> <meta name="ResourceLoaderDynamicStyles" content=""> <link rel="stylesheet" href="/w/load.php?lang=en&modules=site.styles&only=styles&skin=vector-2022"> <meta name="generator" content="MediaWiki 1.44.0-wmf.4"> <meta name="referrer" content="origin"> <meta name="referrer" content="origin-when-cross-origin"> <meta name="robots" content="max-image-preview:standard"> <meta name="format-detection" content="telephone=no"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/1/1c/Genomics_GTL_Program_Payoffs.jpg"> <meta property="og:image:width" content="1200"> <meta property="og:image:height" content="901"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/1/1c/Genomics_GTL_Program_Payoffs.jpg"> <meta property="og:image:width" content="800"> <meta property="og:image:height" content="601"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Genomics_GTL_Program_Payoffs.jpg/640px-Genomics_GTL_Program_Payoffs.jpg"> <meta property="og:image:width" content="640"> <meta property="og:image:height" content="480"> <meta name="viewport" content="width=1120"> <meta property="og:title" content="Systems ecology - Wikipedia"> <meta property="og:type" content="website"> <link rel="preconnect" href="//upload.wikimedia.org"> <link rel="alternate" media="only screen and (max-width: 640px)" href="//en.m.wikipedia.org/wiki/Systems_ecology"> <link rel="alternate" type="application/x-wiki" title="Edit this page" href="/w/index.php?title=Systems_ecology&action=edit"> <link rel="apple-touch-icon" href="/static/apple-touch/wikipedia.png"> <link rel="icon" href="/static/favicon/wikipedia.ico"> <link rel="search" type="application/opensearchdescription+xml" href="/w/rest.php/v1/search" title="Wikipedia (en)"> <link rel="EditURI" type="application/rsd+xml" href="//en.wikipedia.org/w/api.php?action=rsd"> <link rel="canonical" href="https://en.wikipedia.org/wiki/Systems_ecology"> <link rel="license" href="https://creativecommons.org/licenses/by-sa/4.0/deed.en"> <link rel="alternate" type="application/atom+xml" title="Wikipedia Atom feed" href="/w/index.php?title=Special:RecentChanges&feed=atom"> <link rel="dns-prefetch" href="//meta.wikimedia.org" /> <link rel="dns-prefetch" href="//login.wikimedia.org"> </head> <body class="skin--responsive skin-vector skin-vector-search-vue mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject mw-editable page-Systems_ecology rootpage-Systems_ecology skin-vector-2022 action-view"><a class="mw-jump-link" href="#bodyContent">Jump to content</a> <div class="vector-header-container"> <header class="vector-header mw-header"> <div class="vector-header-start"> <nav class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-dropdown" class="vector-dropdown vector-main-menu-dropdown vector-button-flush-left vector-button-flush-right" > <input type="checkbox" id="vector-main-menu-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-main-menu-dropdown" class="vector-dropdown-checkbox " aria-label="Main menu" > <label id="vector-main-menu-dropdown-label" for="vector-main-menu-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-menu mw-ui-icon-wikimedia-menu"></span> <span class="vector-dropdown-label-text">Main menu</span> </label> <div class="vector-dropdown-content"> <div id="vector-main-menu-unpinned-container" class="vector-unpinned-container"> <div id="vector-main-menu" class="vector-main-menu vector-pinnable-element"> <div class="vector-pinnable-header vector-main-menu-pinnable-header vector-pinnable-header-unpinned" data-feature-name="main-menu-pinned" data-pinnable-element-id="vector-main-menu" data-pinned-container-id="vector-main-menu-pinned-container" data-unpinned-container-id="vector-main-menu-unpinned-container" > <div class="vector-pinnable-header-label">Main menu</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-main-menu.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-main-menu.unpin">hide</button> </div> <div id="p-navigation" class="vector-menu mw-portlet mw-portlet-navigation" > <div class="vector-menu-heading"> Navigation </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-mainpage-description" class="mw-list-item"><a href="/wiki/Main_Page" title="Visit the main page [z]" accesskey="z"><span>Main page</span></a></li><li id="n-contents" class="mw-list-item"><a href="/wiki/Wikipedia:Contents" title="Guides to browsing Wikipedia"><span>Contents</span></a></li><li id="n-currentevents" class="mw-list-item"><a href="/wiki/Portal:Current_events" title="Articles related to current events"><span>Current events</span></a></li><li id="n-randompage" class="mw-list-item"><a href="/wiki/Special:Random" title="Visit a randomly selected article [x]" accesskey="x"><span>Random article</span></a></li><li id="n-aboutsite" class="mw-list-item"><a href="/wiki/Wikipedia:About" title="Learn about Wikipedia and how it works"><span>About Wikipedia</span></a></li><li id="n-contactpage" class="mw-list-item"><a href="//en.wikipedia.org/wiki/Wikipedia:Contact_us" title="How to contact Wikipedia"><span>Contact us</span></a></li> </ul> </div> </div> <div id="p-interaction" class="vector-menu mw-portlet mw-portlet-interaction" > <div class="vector-menu-heading"> Contribute </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-help" class="mw-list-item"><a href="/wiki/Help:Contents" title="Guidance on how to use and edit Wikipedia"><span>Help</span></a></li><li id="n-introduction" class="mw-list-item"><a href="/wiki/Help:Introduction" title="Learn how to edit Wikipedia"><span>Learn to edit</span></a></li><li id="n-portal" class="mw-list-item"><a href="/wiki/Wikipedia:Community_portal" title="The hub for editors"><span>Community portal</span></a></li><li id="n-recentchanges" class="mw-list-item"><a href="/wiki/Special:RecentChanges" title="A list of recent changes to Wikipedia [r]" accesskey="r"><span>Recent changes</span></a></li><li id="n-upload" class="mw-list-item"><a href="/wiki/Wikipedia:File_upload_wizard" title="Add images or other media for use on Wikipedia"><span>Upload file</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> <a href="/wiki/Main_Page" class="mw-logo"> <img class="mw-logo-icon" src="/static/images/icons/wikipedia.png" alt="" aria-hidden="true" height="50" width="50"> <span class="mw-logo-container skin-invert"> <img class="mw-logo-wordmark" alt="Wikipedia" src="/static/images/mobile/copyright/wikipedia-wordmark-en.svg" style="width: 7.5em; height: 1.125em;"> <img class="mw-logo-tagline" alt="The Free Encyclopedia" src="/static/images/mobile/copyright/wikipedia-tagline-en.svg" width="117" height="13" style="width: 7.3125em; height: 0.8125em;"> </span> </a> </div> <div class="vector-header-end"> <div id="p-search" role="search" class="vector-search-box-vue vector-search-box-collapses vector-search-box-show-thumbnail vector-search-box-auto-expand-width vector-search-box"> <a href="/wiki/Special:Search" class="cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only search-toggle" title="Search Wikipedia [f]" accesskey="f"><span class="vector-icon mw-ui-icon-search mw-ui-icon-wikimedia-search"></span> <span>Search</span> </a> <div class="vector-typeahead-search-container"> <div class="cdx-typeahead-search cdx-typeahead-search--show-thumbnail cdx-typeahead-search--auto-expand-width"> <form action="/w/index.php" id="searchform" class="cdx-search-input cdx-search-input--has-end-button"> <div id="simpleSearch" class="cdx-search-input__input-wrapper" data-search-loc="header-moved"> <div class="cdx-text-input cdx-text-input--has-start-icon"> <input class="cdx-text-input__input" type="search" name="search" placeholder="Search Wikipedia" aria-label="Search Wikipedia" autocapitalize="sentences" title="Search Wikipedia [f]" accesskey="f" id="searchInput" > <span class="cdx-text-input__icon cdx-text-input__start-icon"></span> </div> <input type="hidden" name="title" value="Special:Search"> </div> <button class="cdx-button cdx-search-input__end-button">Search</button> </form> </div> </div> </div> <nav class="vector-user-links vector-user-links-wide" aria-label="Personal tools"> <div class="vector-user-links-main"> <div id="p-vector-user-menu-preferences" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-userpage" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-dropdown" class="vector-dropdown " title="Change the appearance of the page's font size, width, and color" > <input type="checkbox" id="vector-appearance-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-appearance-dropdown" class="vector-dropdown-checkbox " aria-label="Appearance" > <label id="vector-appearance-dropdown-label" for="vector-appearance-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-appearance mw-ui-icon-wikimedia-appearance"></span> <span class="vector-dropdown-label-text">Appearance</span> </label> <div class="vector-dropdown-content"> <div id="vector-appearance-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <div id="p-vector-user-menu-notifications" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-overflow" class="vector-menu mw-portlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="https://donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_en.wikipedia.org&uselang=en" class=""><span>Donate</span></a> </li> <li id="pt-createaccount-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:CreateAccount&returnto=Systems+ecology" title="You are encouraged to create an account and log in; however, it is not mandatory" class=""><span>Create account</span></a> </li> <li id="pt-login-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:UserLogin&returnto=Systems+ecology" title="You're encouraged to log in; however, it's not mandatory. [o]" accesskey="o" class=""><span>Log in</span></a> </li> </ul> </div> </div> </div> <div id="vector-user-links-dropdown" class="vector-dropdown vector-user-menu vector-button-flush-right vector-user-menu-logged-out" title="Log in and more options" > <input type="checkbox" id="vector-user-links-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-user-links-dropdown" class="vector-dropdown-checkbox " aria-label="Personal tools" > <label id="vector-user-links-dropdown-label" for="vector-user-links-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-ellipsis mw-ui-icon-wikimedia-ellipsis"></span> <span class="vector-dropdown-label-text">Personal tools</span> </label> <div class="vector-dropdown-content"> <div id="p-personal" class="vector-menu mw-portlet mw-portlet-personal user-links-collapsible-item" title="User menu" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport" class="user-links-collapsible-item mw-list-item"><a href="https://donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_en.wikipedia.org&uselang=en"><span>Donate</span></a></li><li id="pt-createaccount" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:CreateAccount&returnto=Systems+ecology" title="You are encouraged to create an account and log in; however, it is not mandatory"><span class="vector-icon mw-ui-icon-userAdd mw-ui-icon-wikimedia-userAdd"></span> <span>Create account</span></a></li><li id="pt-login" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:UserLogin&returnto=Systems+ecology" title="You're encouraged to log in; however, it's not mandatory. [o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>Log in</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> Pages for logged out editors <a href="/wiki/Help:Introduction" aria-label="Learn more about editing"><span>learn more</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/Special:MyContributions" title="A list of edits made from this IP address [y]" accesskey="y"><span>Contributions</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/Special:MyTalk" title="Discussion about edits from this IP address [n]" accesskey="n"><span>Talk</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"><!-- CentralNotice --></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="Contents" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Contents</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">hide</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">(Top)</div> </a> </li> <li id="toc-Overview" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Overview"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Overview</span> </div> </a> <ul id="toc-Overview-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Closely_related_fields" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Closely_related_fields"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Closely related fields</span> </div> </a> <button aria-controls="toc-Closely_related_fields-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Closely related fields subsection</span> </button> <ul id="toc-Closely_related_fields-sublist" class="vector-toc-list"> <li id="toc-Deep_ecology" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Deep_ecology"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1</span> <span>Deep ecology</span> </div> </a> <ul id="toc-Deep_ecology-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Earth_systems_engineering_and_management" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Earth_systems_engineering_and_management"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.2</span> <span>Earth systems engineering and management</span> </div> </a> <ul id="toc-Earth_systems_engineering_and_management-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Ecological_economics" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Ecological_economics"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.3</span> <span>Ecological economics</span> </div> </a> <ul id="toc-Ecological_economics-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Ecological_energetics" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Ecological_energetics"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.4</span> <span>Ecological energetics</span> </div> </a> <ul id="toc-Ecological_energetics-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Ecological_humanities" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Ecological_humanities"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.5</span> <span>Ecological humanities</span> </div> </a> <ul id="toc-Ecological_humanities-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Ecosystem_ecology" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Ecosystem_ecology"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.6</span> <span>Ecosystem ecology</span> </div> </a> <ul id="toc-Ecosystem_ecology-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Industrial_ecology" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Industrial_ecology"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.7</span> <span>Industrial ecology</span> </div> </a> <ul id="toc-Industrial_ecology-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Bibliography" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Bibliography"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Bibliography</span> </div> </a> <ul id="toc-Bibliography-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Systems ecology</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 6 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-6" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">6 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B9%D9%84%D9%85_%D8%A8%D9%8A%D8%A6%D8%A9_%D8%A7%D9%84%D9%86%D8%B8%D9%85" title="علم بيئة النظم – Arabic" lang="ar" hreflang="ar" data-title="علم بيئة النظم" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Ecolog%C3%ADa_de_sistemas" title="Ecología de sistemas – Spanish" lang="es" hreflang="es" data-title="Ecología de sistemas" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%A8%D9%88%D9%85%E2%80%8C%D8%B4%D9%86%D8%A7%D8%B3%DB%8C_%D8%B3%DB%8C%D8%B3%D8%AA%D9%85%E2%80%8C%D9%87%D8%A7" title="بومشناسی سیستمها – Persian" lang="fa" hreflang="fa" data-title="بومشناسی سیستمها" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Systeemecologie" title="Systeemecologie – Dutch" lang="nl" hreflang="nl" data-title="Systeemecologie" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Sistemska_ekologija" title="Sistemska ekologija – Slovenian" lang="sl" hreflang="sl" data-title="Sistemska ekologija" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A1%D0%B8%D1%81%D1%82%D0%B5%D0%BC%D0%BD%D0%B0_%D0%B5%D0%BA%D0%BE%D0%BB%D0%BE%D0%B3%D1%96%D1%8F" title="Системна екологія – Ukrainian" lang="uk" hreflang="uk" data-title="Системна екологія" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q3076438#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Systems_ecology" title="View the content page [c]" accesskey="c"><span>Article</span></a></li><li id="ca-talk" class="vector-tab-noicon mw-list-item"><a href="/wiki/Talk:Systems_ecology" rel="discussion" title="Discuss improvements to the content page [t]" accesskey="t"><span>Talk</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="Change language variant" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">English</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="Views"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Systems_ecology"><span>Read</span></a></li><li id="ca-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Systems_ecology&action=edit" title="Edit this page [e]" accesskey="e"><span>Edit</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Systems_ecology&action=history" title="Past revisions of this page [h]" accesskey="h"><span>View history</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="Tools" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Tools</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">Tools</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">hide</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="More options" > <div class="vector-menu-heading"> Actions </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/Systems_ecology"><span>Read</span></a></li><li id="ca-more-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Systems_ecology&action=edit" title="Edit this page [e]" accesskey="e"><span>Edit</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Systems_ecology&action=history"><span>View history</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> General </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/Special:WhatLinksHere/Systems_ecology" title="List of all English Wikipedia pages containing links to this page [j]" accesskey="j"><span>What links here</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Special:RecentChangesLinked/Systems_ecology" rel="nofollow" title="Recent changes in pages linked from this page [k]" accesskey="k"><span>Related changes</span></a></li><li id="t-upload" class="mw-list-item"><a href="/wiki/Wikipedia:File_Upload_Wizard" title="Upload files [u]" accesskey="u"><span>Upload file</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Special:SpecialPages" title="A list of all special pages [q]" accesskey="q"><span>Special pages</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Systems_ecology&oldid=1185361143" title="Permanent link to this revision of this page"><span>Permanent link</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=Systems_ecology&action=info" title="More information about this page"><span>Page information</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=Special:CiteThisPage&page=Systems_ecology&id=1185361143&wpFormIdentifier=titleform" title="Information on how to cite this page"><span>Cite this page</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=Special:UrlShortener&url=https%3A%2F%2Fen.wikipedia.org%2Fwiki%2FSystems_ecology"><span>Get shortened URL</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=Special:QrCode&url=https%3A%2F%2Fen.wikipedia.org%2Fwiki%2FSystems_ecology"><span>Download QR code</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> Print/export </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=Special:DownloadAsPdf&page=Systems_ecology&action=show-download-screen" title="Download this page as a PDF file"><span>Download as PDF</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=Systems_ecology&printable=yes" title="Printable version of this page [p]" accesskey="p"><span>Printable version</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> In other projects </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="wb-otherproject-link wb-otherproject-commons mw-list-item"><a href="https://commons.wikimedia.org/wiki/Category:Systems_ecology" hreflang="en"><span>Wikimedia Commons</span></a></li><li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q3076438" title="Structured data on this page hosted by Wikidata [g]" accesskey="g"><span>Wikidata item</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Appearance</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">hide</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Holistic approach to the study of ecological systems</div> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Genomics_GTL_Program_Payoffs.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Genomics_GTL_Program_Payoffs.jpg/300px-Genomics_GTL_Program_Payoffs.jpg" decoding="async" width="300" height="225" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Genomics_GTL_Program_Payoffs.jpg/450px-Genomics_GTL_Program_Payoffs.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Genomics_GTL_Program_Payoffs.jpg/600px-Genomics_GTL_Program_Payoffs.jpg 2x" data-file-width="750" data-file-height="563" /></a><figcaption>Ecological analysis of CO<sub>2</sub> in an <a href="/wiki/Ecosystem" title="Ecosystem">ecosystem</a></figcaption></figure> <p><b>Systems ecology</b> is an <a href="/wiki/Interdisciplinary" class="mw-redirect" title="Interdisciplinary">interdisciplinary</a> field of <a href="/wiki/Ecology" title="Ecology">ecology</a>, a subset of <a href="/wiki/Earth_system_science" title="Earth system science">Earth system science</a>, that takes a <a href="/wiki/Holism" title="Holism">holistic</a> approach to the study of ecological systems, especially <a href="/wiki/Ecosystem" title="Ecosystem">ecosystems</a>.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Systems ecology can be seen as an application of <a href="/wiki/General_systems_theory" class="mw-redirect" title="General systems theory">general systems theory</a> to ecology. Central to the systems ecology approach is the idea that an ecosystem is a <a href="/wiki/Complex_system" title="Complex system">complex system</a> exhibiting <a href="/wiki/Emergent_properties" class="mw-redirect" title="Emergent properties">emergent properties</a>. Systems ecology focuses on interactions and transactions within and between biological and ecological systems, and is especially concerned with the way the functioning of ecosystems can be influenced by human interventions. It uses and extends concepts from <a href="/wiki/Thermodynamics" title="Thermodynamics">thermodynamics</a> and develops other macroscopic descriptions of complex systems. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Overview">Overview</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=1" title="Edit section: Overview"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Systems ecology seeks a <a href="/wiki/Holism" title="Holism">holistic</a> view of the interactions and transactions within and between biological and ecological systems. Systems ecologists realise that the function of any ecosystem can be influenced by human economics in fundamental ways. They have therefore taken an additional transdisciplinary step by including economics in the consideration of <a href="/wiki/Ecological_economics" title="Ecological economics">ecological-economic</a> systems. In the words of <a href="/w/index.php?title=R.L._Kitching&action=edit&redlink=1" class="new" title="R.L. Kitching (page does not exist)">R.L. Kitching</a>:<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <ul><li><i>Systems ecology can be defined as the approach to the study of ecology of organisms using the techniques and philosophy of systems analysis: that is, the methods and tools developed, largely in engineering, for studying, characterizing and making predictions about complex entities, that is, systems..</i></li> <li><i>In any study of an ecological system, an essential early procedure is to draw a diagram of the system of interest ... diagrams indicate the system's <b>boundaries</b> by a solid line. Within these boundaries, series of components are isolated which have been chosen to represent that portion of the world in which the systems analyst is interested ... If there are no connections across the systems' boundaries with the surrounding <b>systems environments</b>, the systems are described as </i>closed<i>. Ecological work, however, deals almost exclusively with </i>open<i> systems.</i><sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li></ul> <p>As a mode of scientific enquiry, a central feature of Systems Ecology is the general application of the <a href="/wiki/Energy_flow_(ecology)" title="Energy flow (ecology)">principles of energetics</a> to all systems at any scale. Perhaps the most notable proponent of this view was <a href="/wiki/Howard_T._Odum" title="Howard T. Odum">Howard T. Odum</a> - sometimes considered the father of ecosystems ecology. In this approach the principles of energetics constitute <a href="/wiki/Energy_flow_(ecology)" title="Energy flow (ecology)">ecosystem principles</a>. Reasoning by <a href="/w/index.php?title=Formal_analogy&action=edit&redlink=1" class="new" title="Formal analogy (page does not exist)">formal analogy</a> from one system to another enables the Systems Ecologist to see principles functioning in an analogous manner across system-scale boundaries. H.T. Odum commonly used the <a href="/wiki/Energy_Systems_Language" class="mw-redirect" title="Energy Systems Language">Energy Systems Language</a> as a tool for making systems diagrams and flow charts. </p><p>The fourth of these principles, the <a href="/wiki/Maximum_power_principle" title="Maximum power principle">principle of maximum power efficiency</a>, takes central place in the analysis and synthesis of ecological systems. The fourth principle suggests that the most evolutionarily advantageous system function occurs when the environmental load matches the internal resistance of the system. The further the environmental load is from matching the internal resistance, the further the system is away from its sustainable steady state. Therefore, the systems ecologist engages in a task of resistance and impedance matching in <a href="/wiki/Ecological_engineering" title="Ecological engineering">ecological engineering</a>, just as the <a href="/wiki/Electronic_engineer" class="mw-redirect" title="Electronic engineer">electronic engineer</a> would do. </p> <div class="mw-heading mw-heading2"><h2 id="Closely_related_fields">Closely related fields</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=2" title="Edit section: Closely related fields"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Deep_ecology">Deep ecology</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=3" title="Edit section: Deep ecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Deep_ecology" title="Deep ecology">Deep ecology</a></div> <p>Deep ecology is an ideology whose metaphysical underpinnings are deeply concerned with the science of ecology. The term was coined by <a href="/wiki/Arne_N%C3%A6ss" title="Arne Næss">Arne Naess</a>, a Norwegian philosopher, <a href="/wiki/Gandhi" class="mw-redirect" title="Gandhi">Gandhian</a> scholar, and <a href="/wiki/Environmental_activist" class="mw-redirect" title="Environmental activist">environmental activist</a>. He argues that the prevailing approach to <a href="/wiki/Environmental_management" class="mw-redirect" title="Environmental management">environmental management</a> is <a href="/wiki/Anthropocentric" class="mw-redirect" title="Anthropocentric">anthropocentric</a>, and that the natural environment is not only "more complex than we imagine, it is more complex than we can imagine."<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> Naess formulated deep ecology in 1973 at an environmental conference in Budapest. </p><p><a href="/wiki/Joanna_Macy" title="Joanna Macy">Joanna Macy</a>, <a href="/wiki/John_Seed" title="John Seed">John Seed</a>, and others developed Naess' thesis into a branch they called <i>experiential deep ecology</i>. Their efforts were motivated by a need they perceived for the development of an "<a href="/wiki/Ecological_self" title="Ecological self">ecological self</a>", which views the human ego as an integrated part of a living system that encompasses the individual. They sought to transcend <a href="/wiki/Altruism" title="Altruism">altruism</a> with a deeper self-interest based on <a href="/w/index.php?title=Biospherical_equality&action=edit&redlink=1" class="new" title="Biospherical equality (page does not exist)">biospherical equality</a> beyond human chauvinism. </p> <div class="mw-heading mw-heading3"><h3 id="Earth_systems_engineering_and_management">Earth systems engineering and management</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=4" title="Edit section: Earth systems engineering and management"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Earth_systems_engineering_and_management" title="Earth systems engineering and management">Earth systems engineering and management</a></div> <p>Earth systems engineering and management (ESEM) is a discipline used to analyze, design, engineer and manage <a href="/wiki/Complexity" title="Complexity">complex</a> <a href="/wiki/Natural_environment" title="Natural environment">environmental</a> <a href="/wiki/System" title="System">systems</a>. It entails a wide range of subject areas including anthropology, engineering, <a href="/wiki/Environmental_science" title="Environmental science">environmental science</a>, ethics and philosophy. At its core, ESEM looks to "rationally design and manage coupled human-natural systems in a highly integrated and ethical fashion" </p> <div class="mw-heading mw-heading3"><h3 id="Ecological_economics">Ecological economics</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=5" title="Edit section: Ecological economics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Ecological_economics" title="Ecological economics">Ecological economics</a></div> <p>Ecological economics is a transdisciplinary field of academic research that addresses the dynamic and spatial interdependence between human economies and natural <a href="/wiki/Ecosystem" title="Ecosystem">ecosystems</a>. Ecological economics brings together and connects different disciplines, within the natural and social sciences but especially between these broad areas. As the name suggests, the field is made up of researchers with a background in <a href="/wiki/Economics" title="Economics">economics</a> and <a href="/wiki/Ecology" title="Ecology">ecology</a>. An important motivation for the emergence of ecological economics has been criticism on the assumptions and approaches of traditional (mainstream) <a href="/wiki/Environmental_economics" title="Environmental economics">environmental and resource economics</a>. </p> <div class="mw-heading mw-heading3"><h3 id="Ecological_energetics">Ecological energetics</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=6" title="Edit section: Ecological energetics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Ecological_energetics" class="mw-redirect" title="Ecological energetics">Ecological energetics</a></div> <p>Ecological energetics is the quantitative study of the flow of energy through ecological systems. It aims to uncover the <a href="/wiki/Scientific_law" title="Scientific law">principles</a> which describe the propensity of such energy flows through the trophic, or 'energy availing' levels of ecological networks. In systems ecology the principles of ecosystem energy flows or "ecosystem laws" (i.e. principles of ecological energetics) are considered formally analogous to the principles of energetics. </p> <div class="mw-heading mw-heading3"><h3 id="Ecological_humanities">Ecological humanities</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=7" title="Edit section: Ecological humanities"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Ecological_humanities" class="mw-redirect" title="Ecological humanities">Ecological humanities</a></div> <p>Ecological humanities aims to bridge the divides between the sciences and the humanities, and between <a href="/wiki/Western_culture" title="Western culture">Western</a>, <a href="/wiki/Eastern_culture" title="Eastern culture">Eastern</a> and <a href="/wiki/Indigenous_peoples" title="Indigenous peoples">Indigenous</a> ways of knowing nature. Like ecocentric political theory, the ecological humanities are characterised by a connectivity ontology and a commitment to two fundamental axioms relating to the need to submit to <a href="/wiki/Ecology" title="Ecology">ecological</a> laws and to see humanity as part of a larger living system. </p> <div class="mw-heading mw-heading3"><h3 id="Ecosystem_ecology">Ecosystem ecology</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=8" title="Edit section: Ecosystem ecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Ecoecolfigure1.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/en/thumb/4/44/Ecoecolfigure1.jpg/220px-Ecoecolfigure1.jpg" decoding="async" width="220" height="144" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/44/Ecoecolfigure1.jpg/330px-Ecoecolfigure1.jpg 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/44/Ecoecolfigure1.jpg/440px-Ecoecolfigure1.jpg 2x" data-file-width="569" data-file-height="372" /></a><figcaption>A <a href="/wiki/Riparian_forest" title="Riparian forest">riparian forest</a> in the <a href="/wiki/White_Mountains_(New_Hampshire)" class="mw-redirect" title="White Mountains (New Hampshire)">White Mountains</a>, <a href="/wiki/New_Hampshire" title="New Hampshire">New Hampshire</a> (USA)</figcaption></figure> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Ecosystem_ecology" title="Ecosystem ecology">Ecosystem ecology</a></div> <p>Ecosystem ecology is the integrated study of <a href="/wiki/Life" title="Life">biotic</a> and <a href="/wiki/Abiotic" class="mw-redirect" title="Abiotic">abiotic</a> components of <a href="/wiki/Ecosystem" title="Ecosystem">ecosystems</a> and their interactions within an ecosystem framework. This <a href="/wiki/Science" title="Science">science</a> examines how ecosystems work and relates this to their components such as <a href="/wiki/Chemical" class="mw-redirect" title="Chemical">chemicals</a>, <a href="/wiki/Bedrock" title="Bedrock">bedrock</a>, <a href="/wiki/Soil" title="Soil">soil</a>, <a href="/wiki/Plant" title="Plant">plants</a>, and <a href="/wiki/Animal" title="Animal">animals</a>. Ecosystem ecology examines physical and biological structure and examines how these ecosystem characteristics interact. </p><p>The relationship between systems ecology and ecosystem ecology is complex. Much of systems ecology can be considered a subset of ecosystem ecology. Ecosystem ecology also utilizes methods that have little to do with the holistic approach of systems ecology. However, systems ecology more actively considers external influences such as economics that usually fall outside the bounds of ecosystem ecology. Whereas ecosystem ecology can be defined as the scientific study of ecosystems, systems ecology is more of a particular approach to the study of ecological systems and phenomena that interact with these systems. </p> <div class="mw-heading mw-heading3"><h3 id="Industrial_ecology">Industrial ecology</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=9" title="Edit section: Industrial ecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Industrial_ecology" title="Industrial ecology">Industrial ecology</a></div> <p>Industrial ecology is the study of <a href="/wiki/Industrial_process" class="mw-redirect" title="Industrial process">industrial processes</a> as linear (open loop) systems, in which resource and capital investments move through the system to become waste, to a closed loop system where wastes become inputs for new processes. </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=10" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239009302">.mw-parser-output .portalbox{padding:0;margin:0.5em 0;display:table;box-sizing:border-box;max-width:175px;list-style:none}.mw-parser-output .portalborder{border:1px solid var(--border-color-base,#a2a9b1);padding:0.1em;background:var(--background-color-neutral-subtle,#f8f9fa)}.mw-parser-output .portalbox-entry{display:table-row;font-size:85%;line-height:110%;height:1.9em;font-style:italic;font-weight:bold}.mw-parser-output .portalbox-image{display:table-cell;padding:0.2em;vertical-align:middle;text-align:center}.mw-parser-output .portalbox-link{display:table-cell;padding:0.2em 0.2em 0.2em 0.3em;vertical-align:middle}@media(min-width:720px){.mw-parser-output .portalleft{clear:left;float:left;margin:0.5em 1em 0.5em 0}.mw-parser-output .portalright{clear:right;float:right;margin:0.5em 0 0.5em 1em}}</style><ul role="navigation" aria-label="Portals" class="noprint portalbox portalborder portalright"> <li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/43/The_Earth_seen_from_Apollo_17_with_transparent_background.png/28px-The_Earth_seen_from_Apollo_17_with_transparent_background.png" decoding="async" width="28" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/43/The_Earth_seen_from_Apollo_17_with_transparent_background.png/42px-The_Earth_seen_from_Apollo_17_with_transparent_background.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/43/The_Earth_seen_from_Apollo_17_with_transparent_background.png/56px-The_Earth_seen_from_Apollo_17_with_transparent_background.png 2x" data-file-width="2790" data-file-height="2776" /></span></span></span><span class="portalbox-link"><a href="/wiki/Portal:Earth_sciences" title="Portal:Earth sciences">Earth sciences portal</a></span></li><li class="portalbox-entry"><span class="portalbox-image"><span class="mw-image-border noviewer" typeof="mw:File"><span><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6a/Earth_Day_Flag.png/32px-Earth_Day_Flag.png" decoding="async" width="32" height="21" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6a/Earth_Day_Flag.png/48px-Earth_Day_Flag.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6a/Earth_Day_Flag.png/64px-Earth_Day_Flag.png 2x" data-file-width="6000" data-file-height="4000" /></span></span></span><span class="portalbox-link"><a href="/wiki/Portal:Ecology" title="Portal:Ecology">Ecology portal</a></span></li><li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><span><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f5/Terra.png/28px-Terra.png" decoding="async" width="28" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f5/Terra.png/42px-Terra.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f5/Terra.png/56px-Terra.png 2x" data-file-width="600" data-file-height="600" /></span></span></span><span class="portalbox-link"><a href="/wiki/Portal:Geography" title="Portal:Geography">Geography portal</a></span></li><li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><span><img alt="diagram" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/00/Complex-adaptive-system.jpg/32px-Complex-adaptive-system.jpg" decoding="async" width="32" height="23" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/00/Complex-adaptive-system.jpg/48px-Complex-adaptive-system.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/00/Complex-adaptive-system.jpg/64px-Complex-adaptive-system.jpg 2x" data-file-width="2100" data-file-height="1500" /></span></span></span><span class="portalbox-link"><a href="/wiki/Portal:Systems_science" title="Portal:Systems science">Systems science portal</a></span></li></ul> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col" style="column-width: 22em;"> <ul><li><a href="/wiki/Agroecology" title="Agroecology">Agroecology</a></li> <li><a href="/wiki/Earth_system_science" title="Earth system science">Earth system science</a></li> <li><a href="/wiki/Ecosystem_ecology" title="Ecosystem ecology">Ecosystem ecology</a></li> <li><a href="/wiki/Ecological_literacy" title="Ecological literacy">Ecological literacy</a></li> <li><a href="/wiki/Emergy" title="Emergy">Emergy</a></li> <li><a href="/wiki/Energy_flow_(ecology)" title="Energy flow (ecology)">Energy flow (ecology)</a></li> <li><a href="/wiki/Energy_Systems_Language" class="mw-redirect" title="Energy Systems Language">Energy Systems Language</a></li> <li><a href="/wiki/Holism_in_science" title="Holism in science">Holism in science</a></li> <li><a href="/wiki/Holon_(philosophy)" title="Holon (philosophy)">Holon (philosophy)</a></li> <li><a href="/wiki/Holistic_management" class="mw-redirect" title="Holistic management">Holistic management</a></li> <li><a href="/wiki/Landscape_ecology" title="Landscape ecology">Landscape ecology</a></li> <li><a href="/wiki/Antireductionism" title="Antireductionism">Antireductionism</a></li> <li><a href="/wiki/Biosemiotics" title="Biosemiotics">Biosemiotics</a></li> <li><a href="/wiki/Ecosemiotics" title="Ecosemiotics">Ecosemiotics</a></li> <li><a href="/wiki/MuSIASEM" title="MuSIASEM">MuSIASEM</a></li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=11" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width reflist-columns-2"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFShugart,_Herman_H._(Herman_Henry),_1944-O'Neill,_R._V._(Robert_V.),_1940-1979" class="citation book cs1">Shugart, Herman H. (Herman Henry), 1944-; O'Neill, R. V. (Robert V.), 1940- (1979). <i>Systems ecology</i>. Dowden, Hutchinson & Ross. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-87933-347-2" title="Special:BookSources/0-87933-347-2"><bdi>0-87933-347-2</bdi></a>. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/4664585">4664585</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Systems+ecology&rft.pub=Dowden%2C+Hutchinson+%26+Ross&rft.date=1979&rft_id=info%3Aoclcnum%2F4664585&rft.isbn=0-87933-347-2&rft.au=Shugart%2C+Herman+H.+%28Herman+Henry%29%2C+1944-&rft.au=O%27Neill%2C+R.+V.+%28Robert+V.%29%2C+1940-&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASystems+ecology" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_book" title="Template:Cite book">cite book</a>}}</code>: CS1 maint: multiple names: authors list (<a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">link</a>) CS1 maint: numeric names: authors list (<a href="/wiki/Category:CS1_maint:_numeric_names:_authors_list" title="Category:CS1 maint: numeric names: authors list">link</a>)</span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFVan_Dyne,_George_M.,_1932-1966" class="citation book cs1">Van Dyne, George M., 1932- (1966). <i>Ecosystems, systems ecology, and systems ecologists</i>. Oak Ridge National Laboratory, Health Physics Divisions. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/4247138">4247138</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Ecosystems%2C+systems+ecology%2C+and+systems+ecologists.&rft.pub=Oak+Ridge+National+Laboratory%2C+Health+Physics+Divisions&rft.date=1966&rft_id=info%3Aoclcnum%2F4247138&rft.au=Van+Dyne%2C+George+M.%2C+1932-&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASystems+ecology" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_book" title="Template:Cite book">cite book</a>}}</code>: CS1 maint: multiple names: authors list (<a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">link</a>) CS1 maint: numeric names: authors list (<a href="/wiki/Category:CS1_maint:_numeric_names:_authors_list" title="Category:CS1 maint: numeric names: authors list">link</a>)</span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWilkinson,_David_M.,_1963-2006" class="citation book cs1">Wilkinson, David M., 1963- (2006). <i>Fundamental processes in ecology : an earth systems approach</i>. Oxford University Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-19-856846-0" title="Special:BookSources/0-19-856846-0"><bdi>0-19-856846-0</bdi></a>. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/67383832">67383832</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Fundamental+processes+in+ecology+%3A+an+earth+systems+approach&rft.pub=Oxford+University+Press&rft.date=2006&rft_id=info%3Aoclcnum%2F67383832&rft.isbn=0-19-856846-0&rft.au=Wilkinson%2C+David+M.%2C+1963-&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASystems+ecology" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_book" title="Template:Cite book">cite book</a>}}</code>: CS1 maint: multiple names: authors list (<a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">link</a>) CS1 maint: numeric names: authors list (<a href="/wiki/Category:CS1_maint:_numeric_names:_authors_list" title="Category:CS1 maint: numeric names: authors list">link</a>)</span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKitching,_R._L._(Roger_Laurence),_1945-1983" class="citation book cs1">Kitching, R. L. (Roger Laurence), 1945- (1983). <i>Systems ecology : an introduction to ecological modelling</i>. University of Queensland Press. p. 9. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-7022-1813-8" title="Special:BookSources/0-7022-1813-8"><bdi>0-7022-1813-8</bdi></a>. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/8845946">8845946</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Systems+ecology+%3A+an+introduction+to+ecological+modelling&rft.pages=9&rft.pub=University+of+Queensland+Press&rft.date=1983&rft_id=info%3Aoclcnum%2F8845946&rft.isbn=0-7022-1813-8&rft.au=Kitching%2C+R.+L.+%28Roger+Laurence%29%2C+1945-&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASystems+ecology" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_book" title="Template:Cite book">cite book</a>}}</code>: CS1 maint: multiple names: authors list (<a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">link</a>) CS1 maint: numeric names: authors list (<a href="/wiki/Category:CS1_maint:_numeric_names:_authors_list" title="Category:CS1 maint: numeric names: authors list">link</a>)</span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKitching,_R._L._(Roger_Laurence),_1945-1983" class="citation book cs1">Kitching, R. L. (Roger Laurence), 1945- (1983). <i>Systems ecology : an introduction to ecological modelling</i>. University of Queensland Press. p. 11. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-7022-1813-8" title="Special:BookSources/0-7022-1813-8"><bdi>0-7022-1813-8</bdi></a>. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/8845946">8845946</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Systems+ecology+%3A+an+introduction+to+ecological+modelling&rft.pages=11&rft.pub=University+of+Queensland+Press&rft.date=1983&rft_id=info%3Aoclcnum%2F8845946&rft.isbn=0-7022-1813-8&rft.au=Kitching%2C+R.+L.+%28Roger+Laurence%29%2C+1945-&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASystems+ecology" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_book" title="Template:Cite book">cite book</a>}}</code>: CS1 maint: multiple names: authors list (<a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">link</a>) CS1 maint: numeric names: authors list (<a href="/wiki/Category:CS1_maint:_numeric_names:_authors_list" title="Category:CS1 maint: numeric names: authors list">link</a>)</span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text">A statement attributed to British biologist <a href="/wiki/J.B.S._Haldane" class="mw-redirect" title="J.B.S. Haldane">J.B.S. Haldane</a></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="Bibliography">Bibliography</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=12" title="Edit section: Bibliography"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239549316">.mw-parser-output .refbegin{margin-bottom:0.5em}.mw-parser-output .refbegin-hanging-indents>ul{margin-left:0}.mw-parser-output .refbegin-hanging-indents>ul>li{margin-left:0;padding-left:3.2em;text-indent:-3.2em}.mw-parser-output .refbegin-hanging-indents ul,.mw-parser-output .refbegin-hanging-indents ul li{list-style:none}@media(max-width:720px){.mw-parser-output .refbegin-hanging-indents>ul>li{padding-left:1.6em;text-indent:-1.6em}}.mw-parser-output .refbegin-columns{margin-top:0.3em}.mw-parser-output .refbegin-columns ul{margin-top:0}.mw-parser-output .refbegin-columns li{page-break-inside:avoid;break-inside:avoid-column}@media screen{.mw-parser-output .refbegin{font-size:90%}}</style><div class="refbegin refbegin-columns references-column-width" style="column-width: 30em"> <ul><li><a href="/wiki/Gregory_Bateson" title="Gregory Bateson">Gregory Bateson</a>, <i>Steps to an Ecology of Mind</i>, 2000.</li> <li>Kenneth Edmund Ferguson, <i>Systems Analysis in Ecology</i>, WATT, 1966, 276 pp.</li> <li>Efraim Halfon, <i>Theoretical Systems Ecology: Advances and Case Studies</i>, Academic Press, 1979.</li> <li>J. W. Haefner, <i>Modeling Biological Systems: Principles and Applications</i>, London., UK, Chapman and Hall 1996, 473 pp.</li> <li>Richard F Johnston, Peter W Frank, Charles Duncan Michener, <i>Annual Review of Ecology and Systematics</i>, 1976, 307 pp.</li> <li>Jorgensen, Sven E., "Introduction to Systems Ecology", CRC Press, 2012.</li> <li>R.L. Kitching, <i>Systems ecology</i>, University of Queensland Press, 1983.</li> <li><a href="/wiki/Howard_T._Odum" title="Howard T. Odum">Howard T. Odum</a>, <i>Systems Ecology: An Introduction</i>, Wiley-Interscience, 1983.</li> <li><a href="/wiki/Howard_T._Odum" title="Howard T. Odum">Howard T. Odum</a>, <i>Ecological and General Systems: An Introduction to Systems Ecology</i>. University Press of Colorado, Niwot, CO, 1994.</li> <li>Friedrich Recknagel, <i>Applied Systems Ecology: Approach and Case Studies in Aquatic Ecology</i>, 1989.</li> <li>James. Sanderson & Larry D. Harris, <i>Landscape Ecology: A Top-down Approach</i>, 2000, 246 pp.</li> <li>Sheldon Smith, <i>Human Systems Ecology: Studies in the Integration of Political Economy</i>, 1989.</li> <li>Shugart, H.H., O’Neil, R.V. (Eds.) <i>Systems Ecology</i>, Dowden, Hutchinson & Ross, Inc., 1979.</li> <li>Van Dyne, George M., <i>Ecosystems, Systems Ecology, and Systems Ecologists</i>, ORNL- 3975. Oak Ridge National Laboratory, Oak Ridge, TN, pp. 1–40, 1966.</li> <li>Patten, Bernard C. (editor), "Systems Analysis and Simulation in Ecology", Volume 1, Academic Press, 1971.</li> <li>Patten, Bernard C. (editor), "Systems Analysis and Simulation in Ecology", Volume 2, Academic Press, 1972.</li> <li>Patten, Bernard C. (editor), "Systems Analysis and Simulation in Ecology", Volume 3, Academic Press, 1975.</li> <li>Patten, Bernard C. (editor), "Systems Analysis and Simulation in Ecology", Volume 4, Academic Press, 1976.</li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Systems_ecology&action=edit&section=13" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png" decoding="async" width="30" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/45px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/59px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></div> <div class="side-box-text plainlist">Wikimedia Commons has media related to <span style="font-weight: bold; font-style: italic;"><a href="https://commons.wikimedia.org/wiki/Category:Systems_ecology" class="extiw" title="commons:Category:Systems ecology">Systems ecology</a></span>.</div></div> </div> <dl><dd>Organisations</dd></dl> <ul><li><a rel="nofollow" class="external text" href="http://www.ecology.su.se/">Systems Ecology Department</a> at the Stockholm University.</li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20080603214448/http://www.falw.vu.nl/Onderzoeksinstituten/index.cfm/home_subsection.cfm/subsectionid/55B99586-E22B-4B5C-89C65764F35DDB54">Systems Ecology Department</a> at the University of Amsterdam.</li> <li><a rel="nofollow" class="external text" href="http://www.esf.edu/efb/hall/se/">Systems ecology Lab</a> at SUNY-ESF.</li> <li><a rel="nofollow" class="external text" href="http://www.ees.ufl.edu/homepp/brown/syseco/">Systems Ecology program</a> at the University of Florida</li> <li><a rel="nofollow" class="external text" href="http://www.dbs.umt.edu/systemsecology/">Systems Ecology program</a> at the University of Montana</li> <li><a rel="nofollow" class="external text" href="http://www.sysecol.ethz.ch/">Terrestrial Systems Ecology</a> of ETH Zürich.</li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist dt::after{content:": "}.mw-parser-output .hlist dd::after,.mw-parser-output .hlist li::after{content:" · ";font-weight:bold}.mw-parser-output .hlist dd:last-child::after,.mw-parser-output .hlist dt:last-child::after,.mw-parser-output .hlist li:last-child::after{content:none}.mw-parser-output .hlist dd dd:first-child::before,.mw-parser-output .hlist dd dt:first-child::before,.mw-parser-output .hlist dd li:first-child::before,.mw-parser-output .hlist dt dd:first-child::before,.mw-parser-output .hlist dt dt:first-child::before,.mw-parser-output .hlist dt li:first-child::before,.mw-parser-output .hlist li dd:first-child::before,.mw-parser-output .hlist li dt:first-child::before,.mw-parser-output .hlist li li:first-child::before{content:" (";font-weight:normal}.mw-parser-output .hlist dd dd:last-child::after,.mw-parser-output .hlist dd dt:last-child::after,.mw-parser-output .hlist dd li:last-child::after,.mw-parser-output .hlist dt dd:last-child::after,.mw-parser-output .hlist dt dt:last-child::after,.mw-parser-output .hlist dt li:last-child::after,.mw-parser-output .hlist li dd:last-child::after,.mw-parser-output .hlist li dt:last-child::after,.mw-parser-output .hlist li li:last-child::after{content:")";font-weight:normal}.mw-parser-output .hlist ol{counter-reset:listitem}.mw-parser-output .hlist ol>li{counter-increment:listitem}.mw-parser-output .hlist ol>li::before{content:" "counter(listitem)"\a0 "}.mw-parser-output .hlist dd ol>li:first-child::before,.mw-parser-output .hlist dt ol>li:first-child::before,.mw-parser-output .hlist li ol>li:first-child::before{content:" ("counter(listitem)"\a0 "}</style><style data-mw-deduplicate="TemplateStyles:r1236075235">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output .navbox-subgroup{width:100%}.mw-parser-output .navbox-group,.mw-parser-output .navbox-title,.mw-parser-output .navbox-abovebelow{padding:0.25em 1em;line-height:1.5em;text-align:center}.mw-parser-output .navbox-group{white-space:nowrap;text-align:right}.mw-parser-output .navbox,.mw-parser-output .navbox-subgroup{background-color:#fdfdfd}.mw-parser-output .navbox-list{line-height:1.5em;border-color:#fdfdfd}.mw-parser-output .navbox-list-with-group{text-align:left;border-left-width:2px;border-left-style:solid}.mw-parser-output tr+tr>.navbox-abovebelow,.mw-parser-output tr+tr>.navbox-group,.mw-parser-output tr+tr>.navbox-image,.mw-parser-output tr+tr>.navbox-list{border-top:2px solid #fdfdfd}.mw-parser-output .navbox-title{background-color:#ccf}.mw-parser-output .navbox-abovebelow,.mw-parser-output .navbox-group,.mw-parser-output .navbox-subgroup .navbox-title{background-color:#ddf}.mw-parser-output .navbox-subgroup .navbox-group,.mw-parser-output .navbox-subgroup .navbox-abovebelow{background-color:#e6e6ff}.mw-parser-output .navbox-even{background-color:#f7f7f7}.mw-parser-output .navbox-odd{background-color:transparent}.mw-parser-output .navbox .hlist td dl,.mw-parser-output .navbox .hlist td ol,.mw-parser-output .navbox .hlist td ul,.mw-parser-output .navbox td.hlist dl,.mw-parser-output .navbox td.hlist ol,.mw-parser-output .navbox td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Ecology:_Modelling_ecosystems:_Trophic_components" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Modelling_ecosystems" title="Template:Modelling ecosystems"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Modelling_ecosystems" title="Template talk:Modelling ecosystems"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Modelling_ecosystems" title="Special:EditPage/Template:Modelling ecosystems"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Ecology:_Modelling_ecosystems:_Trophic_components" style="font-size:114%;margin:0 4em"><a href="/wiki/Ecology" title="Ecology">Ecology</a>: <a href="/wiki/Ecosystem_model" title="Ecosystem model">Modelling ecosystems</a>: <a href="/wiki/Trophic_level" title="Trophic level">Trophic</a> components</div></th></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abiotic_component" title="Abiotic component">Abiotic component</a></li> <li><a href="/wiki/Abiotic_stress" title="Abiotic stress">Abiotic stress</a></li> <li><a href="/wiki/Behavioral_ecology" title="Behavioral ecology">Behaviour</a></li> <li><a href="/wiki/Biogeochemical_cycle" title="Biogeochemical cycle">Biogeochemical cycle</a></li> <li><a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">Biomass</a></li> <li><a href="/wiki/Biotic_component" class="mw-redirect" title="Biotic component">Biotic component</a></li> <li><a href="/wiki/Biotic_stress" title="Biotic stress">Biotic stress</a></li> <li><a href="/wiki/Carrying_capacity" title="Carrying capacity">Carrying capacity</a></li> <li><a href="/wiki/Competition_(biology)" title="Competition (biology)">Competition</a></li> <li><a href="/wiki/Ecosystem" title="Ecosystem">Ecosystem</a></li> <li><a href="/wiki/Ecosystem_ecology" title="Ecosystem ecology">Ecosystem ecology</a></li> <li><a href="/wiki/Ecosystem_model" title="Ecosystem model">Ecosystem model</a></li> <li><a href="/wiki/Green_world_hypothesis" title="Green world hypothesis">Green world hypothesis</a></li> <li><a href="/wiki/Keystone_species" title="Keystone species">Keystone species</a></li> <li><a href="/wiki/List_of_feeding_behaviours" title="List of feeding behaviours">List of feeding behaviours</a></li> <li><a href="/wiki/Metabolic_theory_of_ecology" title="Metabolic theory of ecology">Metabolic theory of ecology</a></li> <li><a href="/wiki/Productivity_(ecology)" title="Productivity (ecology)">Productivity</a></li> <li><a href="/wiki/Resource_(biology)" title="Resource (biology)">Resource</a></li> <li><a href="/wiki/Restoration_ecology" class="mw-redirect" title="Restoration ecology">Restoration</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Autotroph" title="Autotroph">Producers</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Autotroph" title="Autotroph">Autotrophs</a></li> <li><a href="/wiki/Chemosynthesis" title="Chemosynthesis">Chemosynthesis</a></li> <li><a href="/wiki/Chemotroph" title="Chemotroph">Chemotrophs</a></li> <li><a href="/wiki/Foundation_species" title="Foundation species">Foundation species</a></li> <li><a href="/wiki/Kinetotroph" title="Kinetotroph">Kinetotrophs</a></li> <li><a href="/wiki/Mixotroph" title="Mixotroph">Mixotrophs</a></li> <li><a href="/wiki/Myco-heterotrophy" title="Myco-heterotrophy">Myco-heterotrophy</a></li> <li><a href="/wiki/Mycotroph" title="Mycotroph">Mycotroph</a></li> <li><a href="/wiki/Organotroph" title="Organotroph">Organotrophs</a></li> <li><a href="/wiki/Photoheterotroph" title="Photoheterotroph">Photoheterotrophs</a></li> <li><a href="/wiki/Photosynthesis" title="Photosynthesis">Photosynthesis</a></li> <li><a href="/wiki/Photosynthetic_efficiency" title="Photosynthetic efficiency">Photosynthetic efficiency</a></li> <li><a href="/wiki/Phototroph" title="Phototroph">Phototrophs</a></li> <li><a href="/wiki/Primary_nutritional_groups" title="Primary nutritional groups">Primary nutritional groups</a></li> <li><a href="/wiki/Primary_production" title="Primary production">Primary production</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Consumer_(food_chain)" title="Consumer (food chain)">Consumers</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Apex_predator" title="Apex predator">Apex predator</a></li> <li><a href="/wiki/Bacterivore" title="Bacterivore">Bacterivore</a></li> <li><a href="/wiki/Carnivore" title="Carnivore">Carnivores</a></li> <li><a href="/wiki/Chemoorganotroph" class="mw-redirect" title="Chemoorganotroph">Chemoorganotroph</a></li> <li><a href="/wiki/Foraging" title="Foraging">Foraging</a></li> <li><a href="/wiki/Generalist_and_specialist_species" title="Generalist and specialist species">Generalist and specialist species</a></li> <li><a href="/wiki/Intraguild_predation" title="Intraguild predation">Intraguild predation</a></li> <li><a href="/wiki/Herbivore" title="Herbivore">Herbivores</a></li> <li><a href="/wiki/Heterotroph" title="Heterotroph">Heterotroph</a></li> <li><a href="/wiki/Heterotrophic_nutrition" title="Heterotrophic nutrition">Heterotrophic nutrition</a></li> <li><a href="/wiki/Insectivore" title="Insectivore">Insectivore</a></li> <li><a href="/wiki/Mesopredator" title="Mesopredator">Mesopredators</a></li> <li><a href="/wiki/Mesopredator_release_hypothesis" title="Mesopredator release hypothesis">Mesopredator release hypothesis</a></li> <li><a href="/wiki/Omnivore" title="Omnivore">Omnivores</a></li> <li><a href="/wiki/Optimal_foraging_theory" title="Optimal foraging theory">Optimal foraging theory</a></li> <li><a href="/wiki/Planktivore" title="Planktivore">Planktivore</a></li> <li><a href="/wiki/Predation" title="Predation">Predation</a></li> <li><a href="/wiki/Prey_switching" title="Prey switching">Prey switching</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Decomposer" title="Decomposer">Decomposers</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Chemoorganoheterotrophy" class="mw-redirect" title="Chemoorganoheterotrophy">Chemoorganoheterotrophy</a></li> <li><a href="/wiki/Decomposition" title="Decomposition">Decomposition</a></li> <li><a href="/wiki/Detritivore" title="Detritivore">Detritivores</a></li> <li><a href="/wiki/Detritus" title="Detritus">Detritus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Microorganism#Habitats_and_ecology" title="Microorganism">Microorganisms</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archaea" title="Archaea">Archaea</a></li> <li><a href="/wiki/Bacteriophage" title="Bacteriophage">Bacteriophage</a></li> <li><a href="/wiki/Lithoautotroph" title="Lithoautotroph">Lithoautotroph</a></li> <li><a href="/wiki/Lithotroph" title="Lithotroph">Lithotrophy</a></li> <li><a href="/wiki/Marine_microorganisms" title="Marine microorganisms">Marine</a></li> <li><a href="/wiki/Microbial_cooperation" title="Microbial cooperation">Microbial cooperation</a></li> <li><a href="/wiki/Microbial_ecology" title="Microbial ecology">Microbial ecology</a></li> <li><a href="/wiki/Microbial_food_web" title="Microbial food web">Microbial food web</a></li> <li><a href="/wiki/Microbial_intelligence" title="Microbial intelligence">Microbial intelligence</a></li> <li><a href="/wiki/Microbial_loop" title="Microbial loop">Microbial loop</a></li> <li><a href="/wiki/Microbial_mat" title="Microbial mat">Microbial mat</a></li> <li><a href="/wiki/Microbial_metabolism" title="Microbial metabolism">Microbial metabolism</a></li> <li><a href="/wiki/Phage_ecology" title="Phage ecology">Phage ecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Food_web" title="Food web">Food webs</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biomagnification" title="Biomagnification">Biomagnification</a></li> <li><a href="/wiki/Ecological_efficiency" title="Ecological efficiency">Ecological efficiency</a></li> <li><a href="/wiki/Ecological_pyramid" title="Ecological pyramid">Ecological pyramid</a></li> <li><a href="/wiki/Energy_flow_(ecology)" title="Energy flow (ecology)">Energy flow</a></li> <li><a href="/wiki/Food_chain" title="Food chain">Food chain</a></li> <li><a href="/wiki/Trophic_level" title="Trophic level">Trophic level</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Example webs</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lake_ecosystem#Trophic_relationships" title="Lake ecosystem">Lakes</a></li> <li><a href="/wiki/River_ecosystem#Trophic_relationships" title="River ecosystem">Rivers</a></li> <li><a href="/wiki/Soil_food_web" title="Soil food web">Soil</a></li> <li><a href="/wiki/Tritrophic_interactions_in_plant_defense" title="Tritrophic interactions in plant defense">Tritrophic interactions in plant defense</a></li> <li><a href="/wiki/Marine_food_web" title="Marine food web">Marine food webs</a> <ul><li><a href="/wiki/Cold_seep" title="Cold seep">cold seeps</a></li> <li><a href="/wiki/Hydrothermal_vent#Biological_communities" title="Hydrothermal vent">hydrothermal vents</a></li> <li><a href="/wiki/Intertidal_ecology" title="Intertidal ecology">intertidal</a></li> <li><a href="/wiki/Kelp_forest#Trophic_ecology" title="Kelp forest">kelp forests</a></li> <li><a href="/wiki/North_Pacific_Gyre" title="North Pacific Gyre">North Pacific Gyre</a></li> <li><a href="/wiki/Ecology_of_the_San_Francisco_Estuary#Food_web" title="Ecology of the San Francisco Estuary">San Francisco Estuary</a></li> <li><a href="/wiki/Tide_pool" title="Tide pool">tide pool</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Processes</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ascendency" title="Ascendency">Ascendency</a></li> <li><a href="/wiki/Bioaccumulation" title="Bioaccumulation">Bioaccumulation</a></li> <li><a href="/wiki/Cascade_effect_(ecology)" title="Cascade effect (ecology)">Cascade effect</a></li> <li><a href="/wiki/Climax_community" title="Climax community">Climax community</a></li> <li><a href="/wiki/Competitive_exclusion_principle" title="Competitive exclusion principle">Competitive exclusion principle</a></li> <li><a href="/wiki/Consumer%E2%80%93resource_interactions" title="Consumer–resource interactions">Consumer–resource interactions</a></li> <li><a href="/wiki/Copiotroph" title="Copiotroph">Copiotrophs</a></li> <li><a href="/wiki/Dominance_(ecology)" title="Dominance (ecology)">Dominance</a></li> <li><a href="/wiki/Ecological_network" title="Ecological network">Ecological network</a></li> <li><a href="/wiki/Ecological_succession" title="Ecological succession">Ecological succession</a></li> <li><a href="/wiki/Energy_quality" title="Energy quality">Energy quality</a></li> <li><a href="/wiki/Energy_systems_language" title="Energy systems language">Energy systems language</a></li> <li><a href="/wiki/F-ratio_(oceanography)" title="F-ratio (oceanography)">f-ratio</a></li> <li><a href="/wiki/Feed_conversion_ratio" title="Feed conversion ratio">Feed conversion ratio</a></li> <li><a href="/wiki/Feeding_frenzy" title="Feeding frenzy">Feeding frenzy</a></li> <li><a href="/wiki/Mesotrophic_soil" title="Mesotrophic soil">Mesotrophic soil</a></li> <li><a href="/wiki/Nutrient_cycle" title="Nutrient cycle">Nutrient cycle</a></li> <li><a href="/wiki/Oligotroph" title="Oligotroph">Oligotroph</a></li> <li><a href="/wiki/Paradox_of_the_plankton" title="Paradox of the plankton">Paradox of the plankton</a></li> <li><a href="/wiki/Trophic_cascade" title="Trophic cascade">Trophic cascade</a></li> <li><a href="/wiki/Trophic_mutualism" title="Trophic mutualism">Trophic mutualism</a></li> <li><a href="/wiki/Trophic_state_index" title="Trophic state index">Trophic state index</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Defense,<br />counter</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Animal_coloration" title="Animal coloration">Animal coloration</a></li> <li><a href="/wiki/Anti-predator_adaptation" title="Anti-predator adaptation">Anti-predator adaptations</a></li> <li><a href="/wiki/Camouflage" title="Camouflage">Camouflage</a></li> <li><a href="/wiki/Deimatic_behaviour" title="Deimatic behaviour">Deimatic behaviour</a></li> <li><a href="/wiki/Herbivore_adaptations_to_plant_defense" title="Herbivore adaptations to plant defense">Herbivore adaptations to plant defense</a></li> <li><a href="/wiki/Mimicry" title="Mimicry">Mimicry</a></li> <li><a href="/wiki/Plant_defense_against_herbivory" title="Plant defense against herbivory">Plant defense against herbivory</a></li> <li><a href="/wiki/Shoaling_and_schooling#Predator_avoidance" title="Shoaling and schooling">Predator avoidance in schooling fish</a></li></ul> </div></td></tr></tbody></table></div><div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Ecology:_Modelling_ecosystems:_Other_components" style="padding:3px"><table class="nowraplinks hlist mw-collapsible uncollapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Modelling_ecosystems" title="Template:Modelling ecosystems"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Modelling_ecosystems" title="Template talk:Modelling ecosystems"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Modelling_ecosystems" title="Special:EditPage/Template:Modelling ecosystems"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Ecology:_Modelling_ecosystems:_Other_components" style="font-size:114%;margin:0 4em"><a href="/wiki/Ecology" title="Ecology">Ecology</a>: <a href="/wiki/Ecosystem_model" title="Ecosystem model">Modelling ecosystems</a>: Other components</div></th></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Population_ecology" title="Population ecology">Population<br />ecology</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abundance_(ecology)" title="Abundance (ecology)">Abundance</a></li> <li><a href="/wiki/Allee_effect" title="Allee effect">Allee effect</a></li> <li><a href="/wiki/Consumer-resource_model" title="Consumer-resource model">Consumer-resource model</a></li> <li><a href="/wiki/Depensation" title="Depensation">Depensation</a></li> <li><a href="/wiki/Ecological_yield" title="Ecological yield">Ecological yield</a></li> <li><a href="/wiki/Effective_population_size" title="Effective population size">Effective population size</a></li> <li><a href="/wiki/Intraspecific_competition" title="Intraspecific competition">Intraspecific competition</a></li> <li><a href="/wiki/Logistic_function" title="Logistic function">Logistic function</a></li> <li><a href="/wiki/Malthusian_growth_model" title="Malthusian growth model">Malthusian growth model</a></li> <li><a href="/wiki/Maximum_sustainable_yield" title="Maximum sustainable yield">Maximum sustainable yield</a></li> <li><a href="/wiki/Overpopulation" title="Overpopulation">Overpopulation</a></li> <li><a href="/wiki/Overexploitation" title="Overexploitation">Overexploitation</a></li> <li><a href="/wiki/Population_cycle" title="Population cycle">Population cycle</a></li> <li><a href="/wiki/Population_dynamics" title="Population dynamics">Population dynamics</a></li> <li><a href="/wiki/Population_model" title="Population model">Population modeling</a></li> <li><a href="/wiki/Population_size" title="Population size">Population size</a></li> <li><a href="/wiki/Lotka%E2%80%93Volterra_equations" title="Lotka–Volterra equations">Predator–prey (Lotka–Volterra) equations</a></li> <li><a href="/wiki/Recruitment_(biology)" title="Recruitment (biology)">Recruitment</a></li> <li><a href="/wiki/Small_population_size" title="Small population size">Small population size</a></li> <li><a href="/wiki/Ecological_stability" title="Ecological stability">Stability</a> <ul><li><a href="/wiki/Ecological_resilience" title="Ecological resilience">Resilience</a></li> <li><a href="/wiki/Resistance_(ecology)" title="Resistance (ecology)">Resistance</a></li></ul></li> <li><a href="/wiki/Random_generalized_Lotka%E2%80%93Volterra_model" title="Random generalized Lotka–Volterra model">Random generalized Lotka–Volterra model</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Species</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biodiversity" title="Biodiversity">Biodiversity</a></li> <li><a href="/wiki/Density_dependence" title="Density dependence">Density-dependent inhibition</a></li> <li><a href="/wiki/Ecological_effects_of_biodiversity" title="Ecological effects of biodiversity">Ecological effects of biodiversity</a></li> <li><a href="/wiki/Ecological_extinction" title="Ecological extinction">Ecological extinction</a></li> <li><a href="/wiki/Endemism" title="Endemism">Endemic species</a></li> <li><a href="/wiki/Flagship_species" title="Flagship species">Flagship species</a></li> <li><a href="/wiki/Ordination_(statistics)" title="Ordination (statistics)">Gradient analysis</a></li> <li><a href="/wiki/Bioindicator" title="Bioindicator">Indicator species</a></li> <li><a href="/wiki/Introduced_species" title="Introduced species">Introduced species</a></li> <li><a href="/wiki/Invasive_species" title="Invasive species">Invasive species</a> / <a href="/wiki/Native_species" title="Native species">Native species</a></li> <li><a href="/wiki/Latitudinal_gradients_in_species_diversity" title="Latitudinal gradients in species diversity">Latitudinal gradients in species diversity</a></li> <li><a href="/wiki/Minimum_viable_population" title="Minimum viable population">Minimum viable population</a></li> <li><a href="/wiki/Unified_neutral_theory_of_biodiversity" title="Unified neutral theory of biodiversity">Neutral theory</a></li> <li><a href="/wiki/Occupancy%E2%80%93abundance_relationship" title="Occupancy–abundance relationship">Occupancy–abundance relationship</a></li> <li><a href="/wiki/Population_viability_analysis" title="Population viability analysis">Population viability analysis</a></li> <li><a href="/wiki/Priority_effect" title="Priority effect">Priority effect</a></li> <li><a href="/wiki/Rapoport%27s_rule" title="Rapoport's rule">Rapoport's rule</a></li> <li><a href="/wiki/Relative_abundance_distribution" title="Relative abundance distribution">Relative abundance distribution</a></li> <li><a href="/wiki/Relative_species_abundance" title="Relative species abundance">Relative species abundance</a></li> <li><a href="/wiki/Species_diversity" title="Species diversity">Species diversity</a></li> <li><a href="/wiki/Species_homogeneity" title="Species homogeneity">Species homogeneity</a></li> <li><a href="/wiki/Species_richness" title="Species richness">Species richness</a></li> <li><a href="/wiki/Species_distribution" title="Species distribution">Species distribution</a></li> <li><a href="/wiki/Species%E2%80%93area_relationship" title="Species–area relationship">Species–area curve</a></li> <li><a href="/wiki/Umbrella_species" title="Umbrella species">Umbrella species</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Species<br />interaction</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Antibiosis" title="Antibiosis">Antibiosis</a></li> <li><a href="/wiki/Biological_interaction" title="Biological interaction">Biological interaction</a></li> <li><a href="/wiki/Commensalism" title="Commensalism">Commensalism</a></li> <li><a href="/wiki/Community_(ecology)" title="Community (ecology)">Community ecology</a></li> <li><a href="/wiki/Ecological_facilitation" title="Ecological facilitation">Ecological facilitation</a></li> <li><a href="/wiki/Interspecific_competition" title="Interspecific competition">Interspecific competition</a></li> <li><a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">Mutualism</a></li> <li><a href="/wiki/Parasitism" title="Parasitism">Parasitism</a></li> <li><a href="/wiki/Storage_effect" title="Storage effect">Storage effect</a></li> <li><a href="/wiki/Symbiosis" title="Symbiosis">Symbiosis</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Spatial_ecology" title="Spatial ecology">Spatial<br />ecology</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a href="/wiki/Cross-boundary_subsidy" title="Cross-boundary subsidy">Cross-boundary subsidy</a></li> <li><a href="/wiki/Cline_(biology)" title="Cline (biology)">Ecocline</a></li> <li><a href="/wiki/Ecotone" title="Ecotone">Ecotone</a></li> <li><a href="/wiki/Ecotype" title="Ecotype">Ecotype</a></li> <li><a href="/wiki/Disturbance_(ecology)" title="Disturbance (ecology)">Disturbance</a></li> <li><a href="/wiki/Edge_effects" title="Edge effects">Edge effects</a></li> <li><a href="/wiki/Foster%27s_rule" title="Foster's rule">Foster's rule</a></li> <li><a href="/wiki/Habitat_fragmentation" title="Habitat fragmentation">Habitat fragmentation</a></li> <li><a href="/wiki/Ideal_free_distribution" title="Ideal free distribution">Ideal free distribution</a></li> <li><a href="/wiki/Intermediate_disturbance_hypothesis" title="Intermediate disturbance hypothesis">Intermediate disturbance hypothesis</a></li> <li><a href="/wiki/Insular_biogeography" title="Insular biogeography">Insular biogeography</a></li> <li><a href="/wiki/Land_change_modeling" title="Land change modeling">Land change modeling</a></li> <li><a href="/wiki/Landscape_ecology" title="Landscape ecology">Landscape ecology</a></li> <li><a href="/wiki/Landscape_epidemiology" title="Landscape epidemiology">Landscape epidemiology</a></li> <li><a href="/wiki/Landscape_limnology" title="Landscape limnology">Landscape limnology</a></li> <li><a href="/wiki/Metapopulation" title="Metapopulation">Metapopulation</a></li> <li><a href="/wiki/Patch_dynamics" title="Patch dynamics">Patch dynamics</a></li> <li><a href="/wiki/R/K_selection_theory" title="R/K selection theory"><i>r</i>/<i>K</i> selection theory</a></li> <li><a href="/wiki/Resource_selection_function" title="Resource selection function">Resource selection function</a></li> <li><a href="/wiki/Source%E2%80%93sink_dynamics" title="Source–sink dynamics">Source–sink dynamics</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Ecological_niche" title="Ecological niche">Niche</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ecological_niche" title="Ecological niche">Ecological niche</a></li> <li><a href="/wiki/Ecological_trap" title="Ecological trap">Ecological trap</a></li> <li><a href="/wiki/Ecosystem_engineer" title="Ecosystem engineer">Ecosystem engineer</a></li> <li><a href="/wiki/Species_distribution_modelling" title="Species distribution modelling">Environmental niche modelling</a></li> <li><a href="/wiki/Guild_(ecology)" title="Guild (ecology)">Guild</a></li> <li><a href="/wiki/Habitat" title="Habitat">Habitat</a> <ul><li><a href="/wiki/Marine_habitat" title="Marine habitat">marine habitats</a></li></ul></li> <li><a href="/wiki/Limiting_similarity" title="Limiting similarity">Limiting similarity</a></li> <li><a href="/wiki/Niche_apportionment_models" title="Niche apportionment models">Niche apportionment models</a></li> <li><a href="/wiki/Niche_construction" title="Niche construction">Niche construction</a></li> <li><a href="/wiki/Niche_differentiation" class="mw-redirect" title="Niche differentiation">Niche differentiation</a></li> <li><a href="/wiki/Ontogenetic_niche_shift" title="Ontogenetic niche shift">Ontogenetic niche shift</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Non-trophic_networks" title="Non-trophic networks">Other<br />networks</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Assembly_rules" title="Assembly rules">Assembly rules</a></li> <li><a href="/wiki/Bateman%27s_principle" title="Bateman's principle">Bateman's principle</a></li> <li><a href="/wiki/Bioluminescence" title="Bioluminescence">Bioluminescence</a></li> <li><a href="/wiki/Ecological_collapse" class="mw-redirect" title="Ecological collapse">Ecological collapse</a></li> <li><a href="/wiki/Ecological_debt" title="Ecological debt">Ecological debt</a></li> <li><a href="/wiki/Ecological_debt" title="Ecological debt">Ecological deficit</a></li> <li><a href="/wiki/Energy_flow_(ecology)" title="Energy flow (ecology)">Ecological energetics</a></li> <li><a href="/wiki/Ecological_indicator" title="Ecological indicator">Ecological indicator</a></li> <li><a href="/wiki/Ecological_threshold" title="Ecological threshold">Ecological threshold</a></li> <li><a href="/wiki/Ecosystem_diversity" title="Ecosystem diversity">Ecosystem diversity</a></li> <li><a href="/wiki/Emergence" title="Emergence">Emergence</a></li> <li><a href="/wiki/Extinction_debt" title="Extinction debt">Extinction debt</a></li> <li><a href="/wiki/Kleiber%27s_law" title="Kleiber's law">Kleiber's law</a></li> <li><a href="/wiki/Liebig%27s_law_of_the_minimum" title="Liebig's law of the minimum">Liebig's law of the minimum</a></li> <li><a href="/wiki/Marginal_value_theorem" title="Marginal value theorem">Marginal value theorem</a></li> <li><a href="/wiki/Thorson%27s_rule" title="Thorson's rule">Thorson's rule</a></li> <li><a href="/wiki/Xerosere" title="Xerosere">Xerosere</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Allometry" title="Allometry">Allometry</a></li> <li><a href="/wiki/Alternative_stable_state" title="Alternative stable state">Alternative stable state</a></li> <li><a href="/wiki/Balance_of_nature" title="Balance of nature">Balance of nature</a></li> <li><a href="/wiki/Biological_data_visualization" title="Biological data visualization">Biological data visualization</a></li> <li><a href="/wiki/Ecological_economics" title="Ecological economics">Ecological economics</a></li> <li><a href="/wiki/Ecological_footprint" title="Ecological footprint">Ecological footprint</a></li> <li><a href="/wiki/Ecological_forecasting" title="Ecological forecasting">Ecological forecasting</a></li> <li><a href="/wiki/Environmental_humanities" title="Environmental humanities">Ecological humanities</a></li> <li><a href="/wiki/Ecological_stoichiometry" title="Ecological stoichiometry">Ecological stoichiometry</a></li> <li><a href="/wiki/Ecopath" title="Ecopath">Ecopath</a></li> <li><a href="/wiki/Ecosystem_based_fisheries" class="mw-redirect" title="Ecosystem based fisheries">Ecosystem based fisheries</a></li> <li><a href="/wiki/Endolith" title="Endolith">Endolith</a></li> <li><a href="/wiki/Evolutionary_ecology" title="Evolutionary ecology">Evolutionary ecology</a></li> <li><a href="/wiki/Functional_ecology" title="Functional ecology">Functional ecology</a></li> <li><a href="/wiki/Industrial_ecology" title="Industrial ecology">Industrial ecology</a></li> <li><a href="/wiki/Macroecology" title="Macroecology">Macroecology</a></li> <li><a href="/wiki/Microecosystem" title="Microecosystem">Microecosystem</a></li> <li><a href="/wiki/Natural_environment" title="Natural environment">Natural environment</a></li> <li><a href="/wiki/Regime_shift" title="Regime shift">Regime shift</a></li> <li><a href="/wiki/Sexecology" title="Sexecology">Sexecology</a></li> <li><a class="mw-selflink selflink">Systems ecology</a></li> <li><a href="/wiki/Urban_ecology" title="Urban ecology">Urban ecology</a></li> <li><a href="/wiki/Theoretical_ecology" title="Theoretical ecology">Theoretical ecology</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><a href="/wiki/Outline_of_ecology" title="Outline of ecology">Outline of ecology</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Systems_science" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Systems_science" title="Template:Systems science"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Systems_science" title="Template talk:Systems science"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Systems_science" title="Special:EditPage/Template:Systems science"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Systems_science" style="font-size:114%;margin:0 4em"><a href="/wiki/Systems_science" title="Systems science">Systems science</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/System" title="System">System</a><br />types</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Systems_art" title="Systems art">Art</a></li> <li><a href="/wiki/Biological_system" title="Biological system">Biological</a></li> <li><a href="/wiki/Complex_system" title="Complex system">Complex</a></li> <li><a href="/wiki/Coupled_human%E2%80%93environment_system" title="Coupled human–environment system">Coupled human–environment</a></li> <li><a href="/wiki/Ecosystem" title="Ecosystem">Ecological</a></li> <li><a href="/wiki/Economic_system" title="Economic system">Economic</a></li> <li><a href="/wiki/Information_system" title="Information system">Information</a></li> <li><a href="/wiki/Multi-agent_system" title="Multi-agent system">Multi-agent</a></li> <li><a href="/wiki/Nervous_system" title="Nervous system">Nervous</a></li> <li><a href="/wiki/Recommender_system" title="Recommender system">Recommender</a></li> <li><a href="/wiki/Social_system" title="Social system">Social</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Concepts</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Doubling_time" title="Doubling time">Doubling time</a></li> <li><a href="/wiki/Twelve_leverage_points" title="Twelve leverage points">Leverage points</a></li> <li><a href="/wiki/Limiting_factor" title="Limiting factor">Limiting factor</a></li> <li><a href="/wiki/Negative_feedback" title="Negative feedback">Negative feedback</a></li> <li><a href="/wiki/Positive_feedback" title="Positive feedback">Positive feedback</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Theoretical<br />fields</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Control_theory" title="Control theory">Control theory</a></li> <li><a href="/wiki/Cybernetics" title="Cybernetics">Cybernetics</a></li> <li><a href="/wiki/Earth_system_science" title="Earth system science">Earth system science</a></li> <li><a href="/wiki/Living_systems" title="Living systems">Living systems</a></li> <li><a href="/wiki/Sociotechnical_system" title="Sociotechnical system">Sociotechnical system</a></li> <li><a href="/wiki/Systemics" title="Systemics">Systemics</a></li> <li><a href="/wiki/Urban_metabolism" title="Urban metabolism">Urban metabolism</a></li> <li><a href="/wiki/World-systems_theory" title="World-systems theory">World-systems theory</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Systems_analysis" title="Systems analysis">Analysis</a></li> <li><a href="/wiki/Systems_biology" title="Systems biology">Biology</a></li> <li><a href="/wiki/System_dynamics" title="System dynamics">Dynamics</a></li> <li><a class="mw-selflink selflink">Ecology</a></li> <li><a href="/wiki/Systems_engineering" title="Systems engineering">Engineering</a></li> <li><a href="/wiki/Systems_neuroscience" title="Systems neuroscience">Neuroscience</a></li> <li><a href="/wiki/Systems_pharmacology" title="Systems pharmacology">Pharmacology</a></li> <li><a href="/wiki/Systems_philosophy" title="Systems philosophy">Philosophy</a></li> <li><a href="/wiki/Systems_psychology" title="Systems psychology">Psychology</a></li> <li><a href="/wiki/Systems_theory" title="Systems theory">Theory</a> (<a href="/wiki/Systems_thinking" title="Systems thinking">Systems thinking</a>)</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/List_of_systems_scientists" title="List of systems scientists">Scientists</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Russell_L._Ackoff" title="Russell L. Ackoff">Russell L. Ackoff</a></li> <li><a href="/wiki/Victor_Aladjev" title="Victor Aladjev">Victor Aladjev</a></li> <li><a href="/wiki/William_Ross_Ashby" class="mw-redirect" title="William Ross Ashby">William Ross Ashby</a></li> <li><a href="/wiki/Ruzena_Bajcsy" title="Ruzena Bajcsy">Ruzena Bajcsy</a></li> <li><a href="/wiki/B%C3%A9la_H._B%C3%A1n%C3%A1thy" title="Béla H. Bánáthy">Béla H. Bánáthy</a></li> <li><a href="/wiki/Gregory_Bateson" title="Gregory Bateson">Gregory Bateson</a></li> <li><a href="/wiki/Anthony_Stafford_Beer" class="mw-redirect" title="Anthony Stafford Beer">Anthony Stafford Beer</a></li> <li><a href="/wiki/Richard_E._Bellman" title="Richard E. Bellman">Richard E. Bellman</a></li> <li><a href="/wiki/Ludwig_von_Bertalanffy" title="Ludwig von Bertalanffy">Ludwig von Bertalanffy</a></li> <li><a href="/wiki/Margaret_Boden" title="Margaret Boden">Margaret Boden</a></li> <li><a href="/wiki/Alexander_Bogdanov" title="Alexander Bogdanov">Alexander Bogdanov</a></li> <li><a href="/wiki/Kenneth_E._Boulding" title="Kenneth E. Boulding">Kenneth E. Boulding</a></li> <li><a href="/wiki/Murray_Bowen" title="Murray Bowen">Murray Bowen</a></li> <li><a href="/wiki/Kathleen_Carley" title="Kathleen Carley">Kathleen Carley</a></li> <li><a href="/wiki/Mary_Cartwright" title="Mary Cartwright">Mary Cartwright</a></li> <li><a href="/wiki/C._West_Churchman" title="C. West Churchman">C. West Churchman</a></li> <li><a href="/wiki/Manfred_Clynes" title="Manfred Clynes">Manfred Clynes</a></li> <li><a href="/wiki/George_Dantzig" title="George Dantzig">George Dantzig</a></li> <li><a href="/wiki/Edsger_W._Dijkstra" title="Edsger W. Dijkstra">Edsger W. Dijkstra</a></li> <li><a href="/wiki/Fred_Emery" title="Fred Emery">Fred Emery</a></li> <li><a href="/wiki/Heinz_von_Foerster" title="Heinz von Foerster">Heinz von Foerster</a></li> <li><a href="/wiki/Stephanie_Forrest" title="Stephanie Forrest">Stephanie Forrest</a></li> <li><a href="/wiki/Jay_Wright_Forrester" title="Jay Wright Forrester">Jay Wright Forrester</a></li> <li><a href="/wiki/Barbara_J._Grosz" title="Barbara J. Grosz">Barbara Grosz</a></li> <li><a href="/wiki/Charles_A._S._Hall" title="Charles A. S. Hall">Charles A. S. Hall</a></li> <li><a href="/wiki/Mike_Jackson_(systems_scientist)" title="Mike Jackson (systems scientist)">Mike Jackson</a></li> <li><a href="/wiki/Lydia_Kavraki" title="Lydia Kavraki">Lydia Kavraki</a></li> <li><a href="/wiki/James_J._Kay" title="James J. Kay">James J. Kay</a></li> <li><a href="/wiki/Faina_Mihajlovna_Kirillova" class="mw-redirect" title="Faina Mihajlovna Kirillova">Faina M. Kirillova</a></li> <li><a href="/wiki/George_Klir" title="George Klir">George Klir</a></li> <li><a href="/wiki/Allenna_Leonard" title="Allenna Leonard">Allenna Leonard</a></li> <li><a href="/wiki/Edward_Norton_Lorenz" title="Edward Norton Lorenz">Edward Norton Lorenz</a></li> <li><a href="/wiki/Niklas_Luhmann" title="Niklas Luhmann">Niklas Luhmann</a></li> <li><a href="/wiki/Humberto_Maturana" title="Humberto Maturana">Humberto Maturana</a></li> <li><a href="/wiki/Margaret_Mead" title="Margaret Mead">Margaret Mead</a></li> <li><a href="/wiki/Donella_Meadows" title="Donella Meadows">Donella Meadows</a></li> <li><a href="/wiki/Mihajlo_D._Mesarovic" title="Mihajlo D. Mesarovic">Mihajlo D. Mesarovic</a></li> <li><a href="/wiki/James_Grier_Miller" title="James Grier Miller">James Grier Miller</a></li> <li><a href="/wiki/Radhika_Nagpal" title="Radhika Nagpal">Radhika Nagpal</a></li> <li><a href="/wiki/Howard_T._Odum" title="Howard T. Odum">Howard T. Odum</a></li> <li><a href="/wiki/Talcott_Parsons" title="Talcott Parsons">Talcott Parsons</a></li> <li><a href="/wiki/Ilya_Prigogine" title="Ilya Prigogine">Ilya Prigogine</a></li> <li><a href="/wiki/Qian_Xuesen" title="Qian Xuesen">Qian Xuesen</a></li> <li><a href="/wiki/Anatol_Rapoport" title="Anatol Rapoport">Anatol Rapoport</a></li> <li><a href="/wiki/John_Seddon" title="John Seddon">John Seddon</a></li> <li><a href="/wiki/Peter_Senge" title="Peter Senge">Peter Senge</a></li> <li><a href="/wiki/Claude_Shannon" title="Claude Shannon">Claude Shannon</a></li> <li><a href="/wiki/Katia_Sycara" title="Katia Sycara">Katia Sycara</a></li> <li><a href="/wiki/Eric_Trist" title="Eric Trist">Eric Trist</a></li> <li><a href="/wiki/Francisco_Varela" title="Francisco Varela">Francisco Varela</a></li> <li><a href="/wiki/Manuela_M._Veloso" title="Manuela M. Veloso">Manuela M. Veloso</a></li> <li><a href="/wiki/Kevin_Warwick" title="Kevin Warwick">Kevin Warwick</a></li> <li><a href="/wiki/Norbert_Wiener" title="Norbert Wiener">Norbert Wiener</a></li> <li><a href="/wiki/Jennifer_Wilby" title="Jennifer Wilby">Jennifer Wilby</a></li> <li><a href="/wiki/Anthony_Wilden" title="Anthony Wilden">Anthony Wilden</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Applications</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Systems_theory_in_anthropology" title="Systems theory in anthropology">Systems theory in anthropology</a></li> <li><a href="/wiki/Systems_theory_in_archaeology" title="Systems theory in archaeology">Systems theory in archaeology</a></li> <li><a href="/wiki/Systems_theory_in_political_science" title="Systems theory in political science">Systems theory in political science</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Organizations</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/List_of_systems_sciences_organizations" title="List of systems sciences organizations">List</a></li> <li><a href="/wiki/Principia_Cybernetica" title="Principia Cybernetica">Principia Cybernetica</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <b><a href="/wiki/Category:Systems_science" title="Category:Systems science">Category</a></b></li> <li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Symbol_portal_class.svg" class="mw-file-description" title="Portal"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/16px-Symbol_portal_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/23px-Symbol_portal_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/31px-Symbol_portal_class.svg.png 2x" data-file-width="180" data-file-height="185" /></a></span> <b><a href="/wiki/Portal:Systems_science" title="Portal:Systems science">Portal</a></b></li> <li><span class="noviewer" typeof="mw:File"><span title="Commons page"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" decoding="async" width="12" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/18px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/24px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span> <b><a href="https://commons.wikimedia.org/wiki/Category:Systems_theory" class="extiw" title="commons:Category:Systems theory">Commons</a></b></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Environmental_science" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Environmental_science" title="Template:Environmental science"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Environmental_science" title="Template talk:Environmental science"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Environmental_science" title="Special:EditPage/Template:Environmental science"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Environmental_science" style="font-size:114%;margin:0 4em"><a href="/wiki/Environmental_science" title="Environmental science">Environmental science</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Main fields</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Atmospheric_science" title="Atmospheric science">Atmospheric science</a></li> <li><a href="/wiki/Biogeochemistry" title="Biogeochemistry">Biogeochemistry</a></li> <li><a href="/wiki/Ecology" title="Ecology">Ecology</a></li> <li><a href="/wiki/Environmental_chemistry" title="Environmental chemistry">Environmental chemistry</a></li> <li><a href="/wiki/Earth_science" title="Earth science">Geosciences</a></li> <li><a href="/wiki/Hydrology" title="Hydrology">Hydrology</a></li> <li><a href="/wiki/Limnology" title="Limnology">Limnology</a></li> <li><a href="/wiki/Oceanography" title="Oceanography">Oceanography</a></li> <li><a href="/wiki/Environmental_soil_science" title="Environmental soil science">Soil science</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:The_Earth_seen_from_Apollo_17.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/97/The_Earth_seen_from_Apollo_17.jpg/100px-The_Earth_seen_from_Apollo_17.jpg" decoding="async" width="100" height="100" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/97/The_Earth_seen_from_Apollo_17.jpg/150px-The_Earth_seen_from_Apollo_17.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/97/The_Earth_seen_from_Apollo_17.jpg/200px-The_Earth_seen_from_Apollo_17.jpg 2x" data-file-width="3000" data-file-height="3002" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Related fields</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biology" title="Biology">Biology</a></li> <li><a href="/wiki/Chemistry" title="Chemistry">Chemistry</a> <ul><li><a href="/wiki/Green_chemistry" title="Green chemistry">green</a></li></ul></li> <li><a href="/wiki/Ecological_economics" title="Ecological economics">Ecological economics</a></li> <li><a href="/wiki/Environmental_design" title="Environmental design">Environmental design</a></li> <li><a href="/wiki/Environmental_economics" title="Environmental economics">Environmental economics</a></li> <li><a href="/wiki/Environmental_engineering" title="Environmental engineering">Environmental engineering</a></li> <li><a href="/wiki/Environmental_health" title="Environmental health">Environmental health</a> <ul><li><a href="/wiki/Environmental_epidemiology" title="Environmental epidemiology">epidemiology</a></li></ul></li> <li><a href="/wiki/Environmental_studies" title="Environmental studies">Environmental studies</a></li> <li><a href="/wiki/Environmental_humanities" title="Environmental humanities">Environmental humanities</a></li> <li><a href="/wiki/Environmental_statistics" title="Environmental statistics">Environmental statistics</a></li> <li><a href="/wiki/Environmental_toxicology" title="Environmental toxicology">Environmental toxicology</a></li> <li><a href="/wiki/Geodesy" title="Geodesy">Geodesy</a></li> <li><a href="/wiki/Physics" title="Physics">Physics</a></li> <li><a href="/wiki/Radioecology" title="Radioecology">Radioecology</a></li> <li><a href="/wiki/Sustainability_science" title="Sustainability science">Sustainability science</a></li> <li><a class="mw-selflink selflink">Systems ecology</a></li> <li><a href="/wiki/Urban_ecology" title="Urban ecology">Urban ecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Applications</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Energy_conservation" title="Energy conservation">Energy conservation</a></li> <li><a href="/wiki/Environmental_technology" title="Environmental technology">Environmental technology</a></li> <li><a href="/wiki/Natural_resource_management" title="Natural resource management">Natural resource management</a></li> <li><a href="/wiki/Pollution_control" class="mw-redirect" title="Pollution control">Pollution control</a></li> <li><a href="/wiki/Public_transport" title="Public transport">Public transport encouragement</a></li> <li><a href="/wiki/Recycling" title="Recycling">Recycling</a></li> <li><a href="/wiki/Environmental_remediation" title="Environmental remediation">Remediation</a></li> <li><a href="/wiki/Renewable_energy" title="Renewable energy">Renewable energy</a></li> <li><a href="/wiki/Road_ecology" title="Road ecology">Road ecology</a></li> <li><a href="/wiki/Sewage_treatment" title="Sewage treatment">Sewage treatment</a></li> <li><a href="/wiki/Urban_metabolism" title="Urban metabolism">Urban metabolism</a></li> <li><a href="/wiki/Water_purification" title="Water purification">Water purification</a></li> <li><a href="/wiki/Waste_management" title="Waste management">Waste management</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Lists</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/List_of_environmental_degrees" title="List of environmental degrees">Degrees</a></li> <li><a href="/wiki/List_of_environmental_journals#Environmental_sciences" title="List of environmental journals">Journals</a></li> <li><a href="/wiki/List_of_environmental_research_institutes" title="List of environmental research institutes">Research institutes</a></li> <li><a href="/wiki/Glossary_of_environmental_science" title="Glossary of environmental science">Glossary</a></li> <li><a href="/wiki/List_of_years_in_the_environment" title="List of years in the environment">Environment by year</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">See also</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Human_impact_on_the_environment" title="Human impact on the environment">Human impact on the environment</a></li> <li><a href="/wiki/Sustainability" title="Sustainability">Sustainability</a></li> <li><a href="/wiki/Technogaianism" title="Technogaianism">Technogaianism</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <ul><li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <b><a href="/wiki/Category:Environmental_science" title="Category:Environmental science">Category</a></b> <ul><li><a href="/wiki/Category:Environmental_scientists" title="Category:Environmental scientists">scientists</a></li></ul></li> <li><span class="noviewer" typeof="mw:File"><span title="Commons page"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" decoding="async" width="12" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/18px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/24px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span> <b><a href="https://commons.wikimedia.org/wiki/Category:Environmental_sciences" class="extiw" title="commons:Category:Environmental sciences">Commons</a></b></li> <li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Symbol_portal_class.svg" class="mw-file-description" title="Portal"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/16px-Symbol_portal_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/23px-Symbol_portal_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/31px-Symbol_portal_class.svg.png 2x" data-file-width="180" data-file-height="185" /></a></span> <b><a href="/wiki/Portal:Environment" title="Portal:Environment">Environment portal</a></b></li> <li><span class="noviewer" typeof="mw:File"><span title="WikiProject"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/37/People_icon.svg/16px-People_icon.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/37/People_icon.svg/24px-People_icon.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/37/People_icon.svg/32px-People_icon.svg.png 2x" data-file-width="100" data-file-height="100" /></span></span> <b><a href="/wiki/Wikipedia:WikiProject_Environment" title="Wikipedia:WikiProject Environment">WikiProject</a></b></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Environmental_social_science" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Environmental_social_science" title="Template:Environmental social science"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Environmental_social_science" title="Template talk:Environmental social science"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Environmental_social_science" title="Special:EditPage/Template:Environmental social science"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Environmental_social_science" style="font-size:114%;margin:0 4em"><a href="/wiki/Environmental_social_science" title="Environmental social science">Environmental social science</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Fields</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ecological_anthropology" title="Ecological anthropology">Ecological anthropology</a></li> <li><a href="/wiki/Ecological_economics" title="Ecological economics">Ecological economics</a></li> <li><a href="/wiki/Environmental_anthropology" title="Environmental anthropology">Environmental anthropology</a></li> <li><a href="/wiki/Environmental_crime" title="Environmental crime">Environmental crime</a></li> <li><a href="/wiki/Environmental_economics" title="Environmental economics">Environmental economics</a></li> <li><a href="/wiki/Environmental_communication" title="Environmental communication">Environmental communication</a></li> <li><a href="/wiki/Environmental_history" title="Environmental history">Environmental history</a></li> <li><a href="/wiki/Environmental_politics" title="Environmental politics">Environmental politics</a></li> <li><a href="/wiki/Environmental_psychology" title="Environmental psychology">Environmental psychology</a></li> <li><a href="/wiki/Environmental_sociology" title="Environmental sociology">Environmental sociology</a></li> <li><a href="/wiki/Human_ecology" title="Human ecology">Human ecology</a></li> <li><a href="/wiki/Human_geography" title="Human geography">Human geography</a></li> <li><a href="/wiki/Political_ecology" title="Political ecology">Political ecology</a></li> <li><a href="/wiki/Regional_science" title="Regional science">Regional science</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="3" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Sustainable_development.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Sustainable_development.svg/150px-Sustainable_development.svg.png" decoding="async" width="150" height="96" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Sustainable_development.svg/225px-Sustainable_development.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/70/Sustainable_development.svg/300px-Sustainable_development.svg.png 2x" data-file-width="512" data-file-height="326" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Related</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agroecology" title="Agroecology">Agroecology</a></li> <li><a href="/wiki/Anthrozoology" title="Anthrozoology">Anthrozoology</a></li> <li><a href="/wiki/Behavioral_geography" title="Behavioral geography">Behavioral geography</a></li> <li><a href="/wiki/Community_studies" title="Community studies">Community studies</a></li> <li><a href="/wiki/Demography" title="Demography">Demography</a></li> <li>Design <ul><li><a href="/wiki/Ecological_design" title="Ecological design">ecological</a></li> <li><a href="/wiki/Environmental_design" title="Environmental design">environmental</a></li></ul></li> <li><a href="/wiki/Ecological_humanities" class="mw-redirect" title="Ecological humanities">Ecological humanities</a></li> <li>Economics <ul><li><a href="/wiki/Energy_economics" title="Energy economics">energy</a></li> <li><a href="/wiki/Thermoeconomics" title="Thermoeconomics">thermo</a></li></ul></li> <li>Environmental <ul><li><a href="/wiki/Environmental_education" title="Environmental education">education</a></li> <li><a href="/wiki/Environmental_ethics" title="Environmental ethics">ethics</a></li> <li><a href="/wiki/Environmental_law" title="Environmental law">law</a></li> <li><a href="/wiki/Environmental_science" title="Environmental science">science</a></li> <li><a href="/wiki/Environmental_studies" title="Environmental studies">studies</a></li> <li><a href="/wiki/Environmental_justice" title="Environmental justice">justice</a></li> <li><a href="/wiki/Environmental_racism" title="Environmental racism">racism</a></li></ul></li> <li><a href="/wiki/Ethnobiology" title="Ethnobiology">Ethnobiology</a> <ul><li><a href="/wiki/Ethnobotany" title="Ethnobotany">botany</a></li> <li><a href="/wiki/Ethnoecology" title="Ethnoecology">ecology</a></li> <li><a href="/wiki/Ethnozoology" title="Ethnozoology">zoology</a></li></ul></li> <li><a href="/wiki/Forestry" title="Forestry">Forestry</a></li> <li><a href="/wiki/Industrial_ecology" title="Industrial ecology">Industrial ecology</a></li> <li><a href="/wiki/Integrated_geography" title="Integrated geography">Integrated geography</a></li> <li><a href="/wiki/Permaculture" title="Permaculture">Permaculture</a></li> <li><a href="/wiki/Rural_sociology" title="Rural sociology">Rural sociology</a></li> <li><a href="/wiki/Sexecology" title="Sexecology">Sexecology</a></li> <li><a href="/wiki/Science,_technology_and_society" class="mw-redirect" title="Science, technology and society">Science, technology and society</a> <ul><li><a href="/wiki/Science_studies" title="Science studies">science studies</a></li></ul></li> <li>Sustainability <ul><li><a href="/wiki/Sustainability_science" title="Sustainability science">science</a></li> <li><a href="/wiki/Sustainability_studies" title="Sustainability studies">studies</a></li></ul></li> <li><a class="mw-selflink selflink">Systems ecology</a></li> <li>Urban <ul><li><a href="/wiki/Urban_ecology" title="Urban ecology">ecology</a></li> <li><a href="/wiki/Urban_geography" title="Urban geography">geography</a></li> <li><a href="/wiki/Urban_metabolism" title="Urban metabolism">metabolism</a></li> <li><a href="/wiki/Urban_studies" title="Urban studies">studies</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Applied</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Architecture <ul><li><a href="/wiki/Landscape_architecture" title="Landscape architecture">landscape</a></li> <li><a href="/wiki/Sustainable_architecture" title="Sustainable architecture">sustainable</a></li></ul></li> <li><a href="/wiki/Ecopsychology" title="Ecopsychology">Ecopsychology</a></li> <li>Engineering <ul><li><a href="/wiki/Ecological_engineering" title="Ecological engineering">ecological</a></li> <li><a href="/wiki/Environmental_engineering" title="Environmental engineering">environmental</a></li></ul></li> <li><a href="/wiki/Green_criminology" title="Green criminology">Green criminology</a></li> <li>Health <ul><li><a href="/wiki/Environmental_health" title="Environmental health">environmental</a></li> <li><a href="/wiki/Environmental_epidemiology" title="Environmental epidemiology">epidemiology</a></li> <li><a href="/wiki/Occupational_safety_and_health" title="Occupational safety and health">occupational</a></li> <li><a href="/wiki/Public_health" title="Public health">public</a></li></ul></li> <li>Management <ul><li><a href="/wiki/Environmental_resources_management" class="mw-redirect" title="Environmental resources management">environmental</a></li> <li><a href="/wiki/Fisheries_management" title="Fisheries management">fisheries</a></li> <li><a href="/wiki/Sustainable_forest_management" class="mw-redirect" title="Sustainable forest management">forest</a></li> <li><a href="/wiki/Natural_resource_management" title="Natural resource management">natural resource</a></li> <li><a href="/wiki/Waste_management" title="Waste management">waste</a></li></ul></li> <li>Planning <ul><li><a href="/wiki/Environmental_planning" title="Environmental planning">environmental</a></li> <li><a href="/wiki/Land-use_planning" title="Land-use planning">land use</a></li> <li><a href="/wiki/Regional_planning" title="Regional planning">regional</a></li> <li><a href="/wiki/Spatial_planning" title="Spatial planning">spatial</a></li> <li><a href="/wiki/Urban_planning" title="Urban planning">urban</a></li></ul></li> <li>Policy <ul><li><a href="/wiki/Energy_policy" title="Energy policy">energy</a></li> <li><a href="/wiki/Environmental_policy" title="Environmental policy">environmental</a></li></ul></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <ul><li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Symbol_portal_class.svg" class="mw-file-description" title="Portal"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/16px-Symbol_portal_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/23px-Symbol_portal_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/31px-Symbol_portal_class.svg.png 2x" data-file-width="180" data-file-height="185" /></a></span> <a href="/wiki/Portal:Environment" title="Portal:Environment">Environment portal</a></li> <li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Environmental_social_science" title="Category:Environmental social science">Category</a></li> <li><a href="/wiki/Category:Environmental_social_science_concepts" title="Category:Environmental social science concepts">Concepts</a></li> <li><a href="/wiki/List_of_environmental_degrees" title="List of environmental degrees">Degrees</a></li> <li><a href="/wiki/List_of_environmental_social_science_journals" title="List of environmental social science journals">Journals</a></li> <li><a href="/wiki/List_of_environment_research_institutes" class="mw-redirect" title="List of environment research institutes">Research institutes</a></li> <li><a href="/wiki/Category:Environmental_social_scientists" title="Category:Environmental social scientists">Scholars</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Branches_of_ecology" style="border-spacing:0.2em 0;;padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Branches_of_ecology" title="Template:Branches of ecology"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Branches_of_ecology" title="Template talk:Branches of ecology"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Branches_of_ecology" title="Special:EditPage/Template:Branches of ecology"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Branches_of_ecology" style="font-size:114%;margin:0 4em"><a href="/wiki/Outline_of_ecology#Subdisciplines_of_ecology,_and_subdiscipline_classification" title="Outline of ecology">Branches of ecology</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Methodology" title="Methodology">Methodology</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Field_research" title="Field research">Field ecology</a></li> <li><a href="/wiki/Quantitative_ecology" title="Quantitative ecology">Quantitative ecology</a></li> <li><a href="/wiki/Theoretical_ecology" title="Theoretical ecology">Theoretical ecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Spatial_scale" title="Spatial scale">Spatial scale</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Microecology" title="Microecology">Microecology</a></li> <li><a href="/wiki/Macroecology" title="Macroecology">Macroecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Biological_organisation" title="Biological organisation">Organisation</a> or scope</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Autecology" title="Autecology">Autecology</a></li> <li><a href="/wiki/Synecology" class="mw-redirect" title="Synecology">Synecology</a> <ul><li><a href="/wiki/Population_ecology" title="Population ecology">Population ecology</a></li> <li><a href="/wiki/Community_ecology" class="mw-redirect" title="Community ecology">Community ecology</a></li> <li><a href="/wiki/Ecosystem_ecology" title="Ecosystem ecology">Ecosystem ecology</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Taxonomy</a> or <a href="/wiki/Taxon" title="Taxon">taxon</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Microbial_ecology" title="Microbial ecology">Microbial ecology</a></li> <li><a href="/wiki/Plant_ecology" title="Plant ecology">Plant ecology</a></li> <li><a href="/wiki/Insect_ecology" title="Insect ecology">Insect ecology</a></li> <li><a href="/wiki/Human_ecology" title="Human ecology">Human ecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Biome" title="Biome">Biome</a> or <a href="/wiki/Biogeographic" class="mw-redirect" title="Biogeographic">biogeographical</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Polar_ecology" title="Polar ecology">Polar ecology</a> <ul><li><a href="/wiki/Arctic_ecology" title="Arctic ecology">Arctic ecology</a></li></ul></li> <li><a href="/wiki/Desert_ecology" title="Desert ecology">Desert ecology</a></li> <li><a href="/wiki/Forest_ecology" title="Forest ecology">Forest ecology</a></li> <li><a href="/wiki/Soil_ecology" title="Soil ecology">Soil ecology</a></li> <li><a href="/wiki/Tropical_ecology" title="Tropical ecology">Tropical ecology</a></li> <li><a href="/wiki/Urban_ecology" title="Urban ecology">Urban ecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Phenomena#Science" class="mw-redirect" title="Phenomena">Ecological phenomena</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Behavioral_ecology" title="Behavioral ecology">Behavioral ecology</a></li> <li><a href="/wiki/Chemical_ecology" title="Chemical ecology">Chemical ecology</a></li> <li><a href="/wiki/Disease_ecology" title="Disease ecology">Disease ecology</a></li> <li><a href="/wiki/Ecophysiology" title="Ecophysiology">Ecophysiology</a></li> <li><a href="/wiki/Ecotoxicology" title="Ecotoxicology">Ecotoxicology</a></li> <li><a href="/wiki/Evolutionary_ecology" title="Evolutionary ecology">Evolutionary ecology</a></li> <li><a href="/wiki/Fire_ecology" title="Fire ecology">Fire ecology</a></li> <li><a href="/wiki/Functional_ecology" title="Functional ecology">Functional ecology</a></li> <li><a href="/wiki/Genetic_ecology" title="Genetic ecology">Genetic ecology</a></li> <li><a href="/wiki/Landscape_ecology" title="Landscape ecology">Landscape ecology</a></li> <li><a href="/wiki/Molecular_ecology" title="Molecular ecology">Molecular ecology</a></li> <li><a href="/wiki/Paleoecology" title="Paleoecology">Paleoecology</a></li> <li><a href="/wiki/Social_ecology_(academic_field)" title="Social ecology (academic field)">Social ecology</a></li> <li><a href="/wiki/Spatial_ecology" title="Spatial ecology">Spatial ecology</a></li> <li><a href="/wiki/Thermal_ecology" title="Thermal ecology">Thermal ecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Interdisciplinarity" title="Interdisciplinarity">Interdisciplinarity</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agroecology" title="Agroecology">Agroecology</a></li> <li><a href="/wiki/Ecological_anthropology" title="Ecological anthropology">Ecological anthropology</a></li> <li><a href="/wiki/Ecological_economics" title="Ecological economics">Ecological economics</a></li> <li><a href="/wiki/Ecological_engineering" title="Ecological engineering">Ecological engineering</a></li> <li><a href="/wiki/Political_ecology" title="Political ecology">Political ecology</a></li> <li><a class="mw-selflink selflink">Systems ecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/History_of_ecology" title="History of ecology">History of ecology</a></li> <li><a href="/wiki/Glossary_of_ecology" title="Glossary of ecology">Glossary of ecology</a></li> <li><a href="/wiki/Applied_ecology" title="Applied ecology">Applied ecology</a> <ul><li><a href="/wiki/Restoration_ecology" class="mw-redirect" title="Restoration ecology">Restoration ecology</a></li></ul></li> <li><a href="/wiki/Ecosophy" title="Ecosophy">Ecosophy</a></li> <li><a href="/wiki/Natural_history" title="Natural history">Natural history</a></li> <li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li></ul> </div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐f69cdc8f6‐7dxkw Cached time: 20241122211405 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 0.571 seconds Real time usage: 0.779 seconds Preprocessor visited node count: 1504/1000000 Post‐expand include size: 119139/2097152 bytes Template argument size: 1026/2097152 bytes Highest expansion depth: 14/100 Expensive parser function count: 12/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 75950/5000000 bytes Lua time usage: 0.297/10.000 seconds Lua memory usage: 5862403/52428800 bytes Number of Wikibase entities loaded: 1/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 576.344 1 -total 25.16% 145.000 7 Template:Navbox 23.88% 137.631 1 Template:Reflist 20.11% 115.920 5 Template:Cite_book 18.29% 105.411 1 Template:Modelling_ecosystems 16.34% 94.175 1 Template:Commonscat 15.74% 90.745 1 Template:Sister_project 15.39% 88.673 1 Template:Side_box 11.72% 67.530 1 Template:Short_description 6.98% 40.257 2 Template:Pagetype --> <!-- Saved in parser cache with key enwiki:pcache:3253299:|#|:idhash:canonical and timestamp 20241122211405 and revision id 1185361143. Rendering was triggered because: page-view --> </div><!--esi <esi:include src="/esitest-fa8a495983347898/content" /> --><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Systems_ecology&oldid=1185361143">https://en.wikipedia.org/w/index.php?title=Systems_ecology&oldid=1185361143</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Categories</a>: <ul><li><a href="/wiki/Category:Systems_ecology" title="Category:Systems ecology">Systems ecology</a></li><li><a href="/wiki/Category:Environmental_science" title="Category:Environmental science">Environmental science</a></li><li><a href="/wiki/Category:Environmental_social_science" title="Category:Environmental social science">Environmental social science</a></li><li><a href="/wiki/Category:Formal_sciences" title="Category:Formal sciences">Formal sciences</a></li><li><a href="/wiki/Category:Systems_theory" title="Category:Systems theory">Systems theory</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks mw-hidden-cats-hidden">Hidden categories: <ul><li><a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">CS1 maint: multiple names: authors list</a></li><li><a href="/wiki/Category:CS1_maint:_numeric_names:_authors_list" title="Category:CS1 maint: numeric names: authors list">CS1 maint: numeric names: authors list</a></li><li><a href="/wiki/Category:Articles_with_short_description" title="Category:Articles with short description">Articles with short description</a></li><li><a href="/wiki/Category:Short_description_matches_Wikidata" title="Category:Short description matches Wikidata">Short description matches Wikidata</a></li><li><a href="/wiki/Category:Commons_category_link_from_Wikidata" title="Category:Commons category link from Wikidata">Commons category link from Wikidata</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> This page was last edited on 16 November 2023, at 06:30<span class="anonymous-show"> (UTC)</span>.</li> <li id="footer-info-copyright">Text is available under the <a href="/wiki/Wikipedia:Text_of_the_Creative_Commons_Attribution-ShareAlike_4.0_International_License" title="Wikipedia:Text of the Creative Commons Attribution-ShareAlike 4.0 International License">Creative Commons Attribution-ShareAlike 4.0 License</a>; additional terms may apply. By using this site, you agree to the <a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Terms_of_Use" class="extiw" title="foundation:Special:MyLanguage/Policy:Terms of Use">Terms of Use</a> and <a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy" class="extiw" title="foundation:Special:MyLanguage/Policy:Privacy policy">Privacy Policy</a>. Wikipedia® is a registered trademark of the <a rel="nofollow" class="external text" href="https://wikimediafoundation.org/">Wikimedia Foundation, Inc.</a>, a non-profit organization.</li> </ul> <ul id="footer-places"> <li id="footer-places-privacy"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy">Privacy policy</a></li> <li id="footer-places-about"><a href="/wiki/Wikipedia:About">About Wikipedia</a></li> <li id="footer-places-disclaimers"><a href="/wiki/Wikipedia:General_disclaimer">Disclaimers</a></li> <li id="footer-places-contact"><a href="//en.wikipedia.org/wiki/Wikipedia:Contact_us">Contact Wikipedia</a></li> <li id="footer-places-wm-codeofconduct"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Universal_Code_of_Conduct">Code of Conduct</a></li> <li id="footer-places-developers"><a href="https://developer.wikimedia.org">Developers</a></li> <li id="footer-places-statslink"><a href="https://stats.wikimedia.org/#/en.wikipedia.org">Statistics</a></li> <li id="footer-places-cookiestatement"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Cookie_statement">Cookie statement</a></li> <li id="footer-places-mobileview"><a href="//en.m.wikipedia.org/w/index.php?title=Systems_ecology&mobileaction=toggle_view_mobile" class="noprint stopMobileRedirectToggle">Mobile view</a></li> </ul> <ul id="footer-icons" class="noprint"> <li id="footer-copyrightico"><a href="https://wikimediafoundation.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/static/images/footer/wikimedia-button.svg" width="84" height="29" alt="Wikimedia Foundation" loading="lazy"></a></li> <li id="footer-poweredbyico"><a href="https://www.mediawiki.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/w/resources/assets/poweredby_mediawiki.svg" alt="Powered by MediaWiki" width="88" height="31" loading="lazy"></a></li> </ul> </footer> </div> </div> </div> <div class="vector-settings" id="p-dock-bottom"> <ul></ul> </div><script>(RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgHostname":"mw-web.codfw.main-f69cdc8f6-fb7ms","wgBackendResponseTime":160,"wgPageParseReport":{"limitreport":{"cputime":"0.571","walltime":"0.779","ppvisitednodes":{"value":1504,"limit":1000000},"postexpandincludesize":{"value":119139,"limit":2097152},"templateargumentsize":{"value":1026,"limit":2097152},"expansiondepth":{"value":14,"limit":100},"expensivefunctioncount":{"value":12,"limit":500},"unstrip-depth":{"value":1,"limit":20},"unstrip-size":{"value":75950,"limit":5000000},"entityaccesscount":{"value":1,"limit":400},"timingprofile":["100.00% 576.344 1 -total"," 25.16% 145.000 7 Template:Navbox"," 23.88% 137.631 1 Template:Reflist"," 20.11% 115.920 5 Template:Cite_book"," 18.29% 105.411 1 Template:Modelling_ecosystems"," 16.34% 94.175 1 Template:Commonscat"," 15.74% 90.745 1 Template:Sister_project"," 15.39% 88.673 1 Template:Side_box"," 11.72% 67.530 1 Template:Short_description"," 6.98% 40.257 2 Template:Pagetype"]},"scribunto":{"limitreport-timeusage":{"value":"0.297","limit":"10.000"},"limitreport-memusage":{"value":5862403,"limit":52428800}},"cachereport":{"origin":"mw-web.codfw.main-f69cdc8f6-7dxkw","timestamp":"20241122211405","ttl":2592000,"transientcontent":false}}});});</script> <script type="application/ld+json">{"@context":"https:\/\/schema.org","@type":"Article","name":"Systems ecology","url":"https:\/\/en.wikipedia.org\/wiki\/Systems_ecology","sameAs":"http:\/\/www.wikidata.org\/entity\/Q3076438","mainEntity":"http:\/\/www.wikidata.org\/entity\/Q3076438","author":{"@type":"Organization","name":"Contributors to Wikimedia projects"},"publisher":{"@type":"Organization","name":"Wikimedia Foundation, Inc.","logo":{"@type":"ImageObject","url":"https:\/\/www.wikimedia.org\/static\/images\/wmf-hor-googpub.png"}},"datePublished":"2005-11-27T08:10:36Z","dateModified":"2023-11-16T06:30:21Z","image":"https:\/\/upload.wikimedia.org\/wikipedia\/commons\/1\/1c\/Genomics_GTL_Program_Payoffs.jpg","headline":"holistic approach to the study of ecological systems"}</script> </body> </html>