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class="vector-toc-numb">2</span> <span>Professional career</span> </div> </a> <ul id="toc-Professional_career-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Yamanaka's_Nobel_Prize–winning_research_in_iPS_cells" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Yamanaka's_Nobel_Prize–winning_research_in_iPS_cells"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Yamanaka's Nobel Prize–winning research in iPS cells</span> </div> </a> <button aria-controls="toc-Yamanaka's_Nobel_Prize–winning_research_in_iPS_cells-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Yamanaka's Nobel Prize–winning research in iPS cells subsection</span> </button> <ul id="toc-Yamanaka's_Nobel_Prize–winning_research_in_iPS_cells-sublist" class="vector-toc-list"> <li id="toc-Background-different_cell_types" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Background-different_cell_types"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.1</span> <span>Background-different cell types</span> </div> </a> <ul id="toc-Background-different_cell_types-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Background-different_stem_cell_techniques" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Background-different_stem_cell_techniques"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.2</span> <span>Background-different stem cell techniques</span> </div> </a> <ul id="toc-Background-different_stem_cell_techniques-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Historical_background" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Historical_background"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.3</span> <span>Historical background</span> </div> </a> <ul id="toc-Historical_background-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Yamanaka's_research" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Yamanaka's_research"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.4</span> <span>Yamanaka's research</span> </div> </a> <ul id="toc-Yamanaka's_research-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Further_research_and_future_prospects" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Further_research_and_future_prospects"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.5</span> <span>Further research and future prospects</span> </div> </a> <ul id="toc-Further_research_and_future_prospects-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Recognition" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Recognition"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Recognition</span> </div> </a> <ul id="toc-Recognition-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Interest_in_sports" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Interest_in_sports"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Interest in sports</span> </div> </a> <ul id="toc-Interest_in_sports-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" title="Table of Contents" > <input type="checkbox" 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type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 56 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-56" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">56 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B4%D9%8A%D9%86%D9%8A%D8%A7_%D9%8A%D8%A7%D9%85%D8%A7%D9%86%D8%A7%D9%83%D8%A7" title="شينيا ياماناكا – Arabic" lang="ar" hreflang="ar" data-title="شينيا ياماناكا" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/%C5%9Einya_Yamanaka" title="Şinya Yamanaka – Azerbaijani" lang="az" hreflang="az" data-title="Şinya Yamanaka" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%B4%D9%86%D8%A7%DB%8C%D8%A7_%DB%8C%D8%A7%D9%85%D8%A7%D9%86%D8%A7%DA%A9%D8%A7" title="شنایا یاماناکا – South Azerbaijani" lang="azb" hreflang="azb" data-title="شنایا یاماناکا" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B6%E0%A6%BF%E0%A6%A8%E0%A6%BF%E0%A6%AF%E0%A6%BC%E0%A6%BE_%E0%A6%87%E0%A6%AF%E0%A6%BC%E0%A6%BE%E0%A6%AE%E0%A6%BE%E0%A6%A8%E0%A6%BE%E0%A6%95%E0%A6%BE" title="শিনিয়া ইয়ামানাকা – Bangla" lang="bn" hreflang="bn" data-title="শিনিয়া ইয়ামানাকা" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Yamanaka_Sin%27ya" title="Yamanaka Sin'ya – Minnan" lang="nan" hreflang="nan" data-title="Yamanaka Sin'ya" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A1%D1%96%D0%BD%E2%80%99%D1%8F_%D0%AF%D0%BC%D0%B0%D0%BD%D0%B0%D0%BA%D0%B0" title="Сін’я Яманака – Belarusian" lang="be" hreflang="be" data-title="Сін’я Яманака" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A8%D0%B8%D0%BD%D1%8F_%D0%AF%D0%BC%D0%B0%D0%BD%D0%B0%D0%BA%D0%B0" title="Шиня Яманака – Bulgarian" lang="bg" hreflang="bg" data-title="Шиня Яманака" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Catalan" lang="ca" hreflang="ca" data-title="Shinya Yamanaka" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/%C5%A0in%E2%80%99ja_Jamanaka" title="Šin’ja Jamanaka – Czech" lang="cs" hreflang="cs" data-title="Šin’ja Jamanaka" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Shin%E2%80%99ya_Yamanaka" title="Shin’ya Yamanaka – German" lang="de" hreflang="de" data-title="Shin’ya Yamanaka" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Estonian" lang="et" hreflang="et" data-title="Shinya Yamanaka" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%A3%CE%AF%CE%BD%CE%B9%CE%B1_%CE%93%CE%B9%CE%B1%CE%BC%CE%B1%CE%BD%CE%AC%CE%BA%CE%B1" title="Σίνια Γιαμανάκα – Greek" lang="el" hreflang="el" data-title="Σίνια Γιαμανάκα" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Shin%27ya_Yamanaka" title="Shin'ya Yamanaka – Spanish" lang="es" hreflang="es" data-title="Shin'ya Yamanaka" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Basque" lang="eu" hreflang="eu" data-title="Shinya Yamanaka" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B4%D9%86%D8%A7%DB%8C%D8%A7_%DB%8C%D8%A7%D9%85%D8%A7%D9%86%D8%A7%DA%A9%D8%A7" title="شنایا یاماناکا – Persian" lang="fa" hreflang="fa" data-title="شنایا یاماناکا" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – French" lang="fr" hreflang="fr" data-title="Shinya Yamanaka" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Irish" lang="ga" hreflang="ga" data-title="Shinya Yamanaka" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Galician" lang="gl" hreflang="gl" data-title="Shinya Yamanaka" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%95%BC%EB%A7%88%EB%82%98%EC%B9%B4_%EC%8B%A0%EC%95%BC" title="야마나카 신야 – Korean" lang="ko" hreflang="ko" data-title="야마나카 신야" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%87%D5%AB%D5%B6%D5%B5%D5%A1_%D5%85%D5%A1%D5%B4%D5%A1%D5%B6%D5%A1%D5%AF%D5%A1" title="Շինյա Յամանակա – Armenian" lang="hy" hreflang="hy" data-title="Շինյա Յամանակա" data-language-autonym="Հայերեն" data-language-local-name="Armenian" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Ido" lang="io" hreflang="io" data-title="Shinya Yamanaka" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Indonesian" lang="id" hreflang="id" data-title="Shinya Yamanaka" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Shin%27ya_Yamanaka" title="Shin'ya Yamanaka – Italian" lang="it" hreflang="it" data-title="Shin'ya Yamanaka" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A9%D7%99%D7%A0%D7%99%D7%94_%D7%99%D7%90%D7%9E%D7%90%D7%A0%D7%A7%D7%94" title="שיניה יאמאנקה – Hebrew" lang="he" hreflang="he" data-title="שיניה יאמאנקה" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A1%D0%B8%D0%BD%D1%8A%D1%8F_%D0%AF%D0%BC%D0%B0%D0%BD%D0%B0%D0%BA%D0%B0" title="Синъя Яманака – Kazakh" lang="kk" hreflang="kk" data-title="Синъя Яманака" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Swahili" lang="sw" hreflang="sw" data-title="Shinya Yamanaka" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Latin" lang="la" hreflang="la" data-title="Shinya Yamanaka" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Jamanaka_Sinja" title="Jamanaka Sinja – Hungarian" lang="hu" hreflang="hu" data-title="Jamanaka Sinja" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%B4%D9%8A%D9%86%D9%8A%D8%A7_%D9%8A%D8%A7%D9%85%D8%A7%D9%86%D8%A7%D9%83%D8%A7" title="شينيا ياماناكا – Egyptian Arabic" lang="arz" hreflang="arz" data-title="شينيا ياماناكا" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%B4%D9%86%D8%A7%DB%8C%D8%A7_%DB%8C%D8%A7%D9%85%D8%A7%D9%86%D8%A7%DA%A9%D8%A7" title="شنایا یاماناکا – Mazanderani" lang="mzn" hreflang="mzn" data-title="شنایا یاماناکا" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Malay" lang="ms" hreflang="ms" data-title="Shinya Yamanaka" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Dutch" lang="nl" hreflang="nl" data-title="Shinya Yamanaka" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E5%B1%B1%E4%B8%AD%E4%BC%B8%E5%BC%A5" title="山中伸弥 – Japanese" lang="ja" hreflang="ja" data-title="山中伸弥" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Shinya Yamanaka" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Occitan" lang="oc" hreflang="oc" data-title="Shinya Yamanaka" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%B4%D9%86%DB%8C%D8%A7_%DB%8C%D8%A7%D9%85%D8%A7%D9%86%D8%A7%DA%A9%D8%A7" title="شنیا یاماناکا – Western Punjabi" lang="pnb" hreflang="pnb" data-title="شنیا یاماناکا" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Shin%E2%80%99ya_Yamanaka" title="Shin’ya Yamanaka – Low German" lang="nds" hreflang="nds" data-title="Shin’ya Yamanaka" data-language-autonym="Plattdüütsch" data-language-local-name="Low German" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Shin%E2%80%99ya_Yamanaka" title="Shin’ya Yamanaka – Polish" lang="pl" hreflang="pl" data-title="Shin’ya Yamanaka" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Portuguese" lang="pt" hreflang="pt" data-title="Shinya Yamanaka" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Romanian" lang="ro" hreflang="ro" data-title="Shinya Yamanaka" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%AF%D0%BC%D0%B0%D0%BD%D0%B0%D0%BA%D0%B0,_%D0%A1%D0%B8%D0%BD%D1%8A%D1%8F" title="Яманака, Синъя – Russian" lang="ru" hreflang="ru" data-title="Яманака, Синъя" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-scn mw-list-item"><a href="https://scn.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Sicilian" lang="scn" hreflang="scn" data-title="Shinya Yamanaka" data-language-autonym="Sicilianu" data-language-local-name="Sicilian" class="interlanguage-link-target"><span>Sicilianu</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Simple English" lang="en-simple" hreflang="en-simple" data-title="Shinya Yamanaka" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/%C5%A0inja_Jamanaka" title="Šinja Jamanaka – Slovenian" lang="sl" hreflang="sl" data-title="Šinja Jamanaka" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D8%B4%DB%8C%D9%86%DB%8C%D8%A7_%DB%8C%D8%A7%D9%85%D8%A7%D9%86%D8%A7%DA%A9%D8%A7" title="شینیا یاماناکا – Central Kurdish" lang="ckb" hreflang="ckb" data-title="شینیا یاماناکا" data-language-autonym="کوردی" data-language-local-name="Central Kurdish" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Finnish" lang="fi" hreflang="fi" data-title="Shinya Yamanaka" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Swedish" lang="sv" hreflang="sv" data-title="Shinya Yamanaka" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%9A%E0%AE%BF%E0%AE%A9%E0%AF%8D%E0%AE%AF%E0%AE%BE_%E0%AE%AF%E0%AE%BE%E0%AE%AE%E0%AE%BE%E0%AE%A9%E0%AE%BE%E0%AE%95%E0%AF%8D%E0%AE%95%E0%AE%BE" title="சின்யா யாமானாக்கா – Tamil" lang="ta" hreflang="ta" data-title="சின்யா யாமானாக்கா" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%A1%D0%B8%D0%BD%D1%8C%D0%B5_%D0%AF%D0%BC%D0%B0%D0%BD%D0%B0%D0%BA%D0%B5" title="Синье Яманаке – Tatar" lang="tt" hreflang="tt" data-title="Синье Яманаке" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatar" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Turkish" lang="tr" hreflang="tr" data-title="Shinya Yamanaka" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%AF%D0%BC%D0%B0%D0%BD%D0%B0%D0%BA%D0%B0_%D0%A1%D1%96%D0%BD%27%D1%8F" title="Яманака Сін'я – Ukrainian" lang="uk" hreflang="uk" data-title="Яманака Сін'я" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%B4%D9%86%DB%8C%D8%A7_%DB%8C%D8%A7%D9%85%D8%A7%D9%86%D8%A7%DA%A9%D8%A7" title="شنیا یاماناکا – Urdu" lang="ur" hreflang="ur" data-title="شنیا یاماناکا" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Yamanaka_Shin%27ya" title="Yamanaka Shin'ya – Vietnamese" lang="vi" hreflang="vi" data-title="Yamanaka Shin'ya" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Shinya_Yamanaka" title="Shinya Yamanaka – Yoruba" lang="yo" hreflang="yo" data-title="Shinya Yamanaka" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E5%B1%B1%E4%B8%AD%E4%BC%B8%E5%BD%8C" title="山中伸彌 – Cantonese" lang="yue" hreflang="yue" data-title="山中伸彌" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%B1%B1%E4%B8%AD%E4%BC%B8%E5%BD%8C" title="山中伸彌 – Chinese" lang="zh" hreflang="zh" data-title="山中伸彌" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span 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<div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Japanese stem cell researcher (born 1962)</div> <style data-mw-deduplicate="TemplateStyles:r1251242444">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output .ambox-content{border-left:10px solid #f28500}.mw-parser-output .ambox-style{border-left:10px solid #fc3}.mw-parser-output .ambox-move{border-left:10px solid #9932cc}.mw-parser-output .ambox-protection{border-left:10px solid #a2a9b1}.mw-parser-output .ambox .mbox-text{border:none;padding:0.25em 0.5em;width:100%}.mw-parser-output .ambox .mbox-image{border:none;padding:2px 0 2px 0.5em;text-align:center}.mw-parser-output .ambox .mbox-imageright{border:none;padding:2px 0.5em 2px 0;text-align:center}.mw-parser-output .ambox .mbox-empty-cell{border:none;padding:0;width:1px}.mw-parser-output .ambox .mbox-image-div{width:52px}@media(min-width:720px){.mw-parser-output .ambox{margin:0 10%}}@media print{body.ns-0 .mw-parser-output .ambox{display:none!important}}</style><table class="box-More_footnotes_needed plainlinks metadata ambox ambox-style ambox-More_footnotes_needed" role="presentation"><tbody><tr><td class="mbox-image"><div class="mbox-image-div"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Text_document_with_red_question_mark.svg/40px-Text_document_with_red_question_mark.svg.png" decoding="async" width="40" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Text_document_with_red_question_mark.svg/60px-Text_document_with_red_question_mark.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Text_document_with_red_question_mark.svg/80px-Text_document_with_red_question_mark.svg.png 2x" data-file-width="48" data-file-height="48" /></span></span></div></td><td class="mbox-text"><div class="mbox-text-span">This article includes a list of <a href="/wiki/Wikipedia:Citing_sources#General_references" title="Wikipedia:Citing sources">general references</a>, but <b>it lacks sufficient corresponding <a href="/wiki/Wikipedia:Citing_sources#Inline_citations" title="Wikipedia:Citing sources">inline citations</a></b>.<span class="hide-when-compact"> Please help to <a href="/wiki/Wikipedia:WikiProject_Reliability" title="Wikipedia:WikiProject Reliability">improve</a> this article by <a href="/wiki/Wikipedia:When_to_cite" title="Wikipedia:When to cite">introducing</a> more precise citations.</span> <span class="date-container"><i>(<span class="date">January 2015</span>)</i></span><span class="hide-when-compact"><i> (<small><a href="/wiki/Help:Maintenance_template_removal" title="Help:Maintenance template removal">Learn how and when to remove this message</a></small>)</i></span></div></td></tr></tbody></table> <p class="mw-empty-elt"> </p> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">The native form of this <a href="/wiki/Personal_name" title="Personal name">personal name</a> is <i>Yamanaka Shin'ya</i>. This article uses <a href="/wiki/Personal_name#Western_name_order" title="Personal name">Western name order</a> when mentioning individuals.</div> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">Shinya Yamanaka</div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Shinya_Yamanaka_20141226.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fe/Shinya_Yamanaka_20141226.jpg/220px-Shinya_Yamanaka_20141226.jpg" decoding="async" width="220" height="330" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fe/Shinya_Yamanaka_20141226.jpg/330px-Shinya_Yamanaka_20141226.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fe/Shinya_Yamanaka_20141226.jpg/440px-Shinya_Yamanaka_20141226.jpg 2x" data-file-width="2176" data-file-height="3265" /></a></span><div class="infobox-caption">Yamanaka in 2014</div></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><span style="display:none"> (<span class="bday">1962-09-04</span>) </span>September 4, 1962<span class="noprint ForceAgeToShow"> (age 62)</span><br /><div style="display:inline" class="birthplace"><a href="/wiki/Higashi%C5%8Dsaka,_Osaka" class="mw-redirect" title="Higashiōsaka, Osaka">Higashiōsaka, Osaka</a>, Japan</div></td></tr><tr><th scope="row" class="infobox-label">Nationality</th><td class="infobox-data category">Japanese</td></tr><tr><th scope="row" class="infobox-label">Alma mater</th><td class="infobox-data"><a href="/wiki/Kobe_University" title="Kobe University">Kobe University</a> (<a href="/wiki/Doctor_of_Medicine" title="Doctor of Medicine">MD</a>)<br /><a href="/wiki/Osaka_City_University" title="Osaka City University">Osaka City University</a> (<a href="/wiki/PhD" class="mw-redirect" title="PhD">PhD</a>)</td></tr><tr><th scope="row" class="infobox-label">Known for</th><td class="infobox-data"><a href="/wiki/Induced_pluripotent_stem_cell" title="Induced pluripotent stem cell">Induced pluripotent stem cell</a></td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><a href="/wiki/Meyenburg_Prize" title="Meyenburg Prize">Meyenburg Prize</a> (2007)<br /><a href="/wiki/Massry_Prize" title="Massry Prize">Massry Prize</a> (2008)<br /><a href="/wiki/Robert_Koch_Prize" class="mw-redirect" title="Robert Koch Prize">Robert Koch Prize</a> (2008)<br /><a href="/wiki/Shaw_Prize" title="Shaw Prize">Shaw Prize</a> (2008)<br /><a href="/wiki/Gairdner_Foundation_International_Award" class="mw-redirect" title="Gairdner Foundation International Award">Gairdner Foundation International Award</a> (2009)<br /><a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Basic Medical Research Award</a> (2009) <br /> <a href="/wiki/Balzan_Prize" title="Balzan Prize">Balzan Prize</a> (2010) <br /> <a href="/wiki/Kyoto_Prize" title="Kyoto Prize">Kyoto Prize</a> (2010) <br /> <a href="/wiki/BBVA_Foundation_Frontiers_of_Knowledge_Award" title="BBVA Foundation Frontiers of Knowledge Award">BBVA Foundation Frontiers of Knowledge Award</a> (2010)<br /><a href="/wiki/Wolf_Prize" title="Wolf Prize">Wolf Prize</a> (2011)<br /><a rel="nofollow" class="external text" href="http://www.isscr.org/McEwen_Award_for_Innovation/5323.htm">McEwen Award for Innovation</a> (2011)<br />Fellow of the <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">National Academy of Sciences</a><sup id="cite_ref-nas_1-0" class="reference"><a href="#cite_note-nas-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> (2012)<br /><a href="/wiki/Millennium_Technology_Prize" title="Millennium Technology Prize">Millennium Technology Prize</a> (2012)<br /><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> (2012)</td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Stem_cell" title="Stem cell">Stem cell</a> research<sup id="cite_ref-stem1_2-0" class="reference"><a href="#cite_note-stem1-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-stem2_3-0" class="reference"><a href="#cite_note-stem2-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-stem3_4-0" class="reference"><a href="#cite_note-stem3-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><a href="/wiki/Kyoto_University" title="Kyoto University">Kyoto University</a><br /><a href="/wiki/Nara_Institute_of_Science_and_Technology" title="Nara Institute of Science and Technology">Nara Institute of Science and Technology</a><br /><a href="/wiki/J._David_Gladstone_Institutes" class="mw-redirect" title="J. David Gladstone Institutes">Gladstone Institute of Cardiovascular disease</a><br /><a href="/wiki/University_of_California,_San_Francisco" title="University of California, San Francisco">University of California, San Francisco</a></td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <figure class="mw-default-size" typeof="mw:File/Thumb"><span><video id="mwe_player_0" poster="//upload.wikimedia.org/wikipedia/commons/thumb/b/b9/A_single_cardiomyocyte_beating%2C_five_days_after_purification_from_cell_culture.ogv/220px--A_single_cardiomyocyte_beating%2C_five_days_after_purification_from_cell_culture.ogv.jpg" controls="" preload="none" data-mw-tmh="" class="mw-file-element" width="220" height="165" data-durationhint="14" data-mwtitle="A_single_cardiomyocyte_beating,_five_days_after_purification_from_cell_culture.ogv" data-mwprovider="wikimediacommons" resource="/wiki/File:A_single_cardiomyocyte_beating,_five_days_after_purification_from_cell_culture.ogv"><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/b/b9/A_single_cardiomyocyte_beating%2C_five_days_after_purification_from_cell_culture.ogv/A_single_cardiomyocyte_beating%2C_five_days_after_purification_from_cell_culture.ogv.360p.webm" type="video/webm; codecs="vp8, vorbis"" data-transcodekey="360p.webm" data-width="274" data-height="206" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/b/b9/A_single_cardiomyocyte_beating%2C_five_days_after_purification_from_cell_culture.ogv/A_single_cardiomyocyte_beating%2C_five_days_after_purification_from_cell_culture.ogv.240p.vp9.webm" type="video/webm; codecs="vp9, opus"" data-transcodekey="240p.vp9.webm" data-width="274" data-height="206" /><source src="//upload.wikimedia.org/wikipedia/commons/b/b9/A_single_cardiomyocyte_beating%2C_five_days_after_purification_from_cell_culture.ogv" type="video/ogg; codecs="theora"" data-width="274" data-height="206" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/b/b9/A_single_cardiomyocyte_beating%2C_five_days_after_purification_from_cell_culture.ogv/A_single_cardiomyocyte_beating%2C_five_days_after_purification_from_cell_culture.ogv.144p.mjpeg.mov" type="video/quicktime" data-transcodekey="144p.mjpeg.mov" data-width="192" data-height="144" /></video></span><figcaption>Video of a single beating <a href="/wiki/Cardiomyocyte" class="mw-redirect" title="Cardiomyocyte">cardiomyocyte</a>, taken from an <a href="/wiki/Open-access" class="mw-redirect" title="Open-access">open-access</a> <a href="//doi.org/10.1371/journal.pone.0012743" class="extiw" title="doi:10.1371/journal.pone.0012743">article</a> co-authored by Yamanaka.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> Isolating cells by cell type is an important step in <a href="/wiki/Stem_cell_therapy" class="mw-redirect" title="Stem cell therapy">stem cell therapy</a>.</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Shinya_Yamanaka_giving_a_lecture_at_NIH.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7a/Shinya_Yamanaka_giving_a_lecture_at_NIH.jpg/220px-Shinya_Yamanaka_giving_a_lecture_at_NIH.jpg" decoding="async" width="220" height="130" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/7/7a/Shinya_Yamanaka_giving_a_lecture_at_NIH.jpg 1.5x" data-file-width="250" data-file-height="148" /></a><figcaption>Shinya Yamanaka speaking at a lecture on 2010 January 14</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Pre-Ceremony,_50th_All-Japan_Rugby_Football_Championship_%E2%80%9003.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/36/Pre-Ceremony%2C_50th_All-Japan_Rugby_Football_Championship_%E2%80%9003.JPG/220px-Pre-Ceremony%2C_50th_All-Japan_Rugby_Football_Championship_%E2%80%9003.JPG" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/36/Pre-Ceremony%2C_50th_All-Japan_Rugby_Football_Championship_%E2%80%9003.JPG/330px-Pre-Ceremony%2C_50th_All-Japan_Rugby_Football_Championship_%E2%80%9003.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/36/Pre-Ceremony%2C_50th_All-Japan_Rugby_Football_Championship_%E2%80%9003.JPG/440px-Pre-Ceremony%2C_50th_All-Japan_Rugby_Football_Championship_%E2%80%9003.JPG 2x" data-file-width="3648" data-file-height="2736" /></a><figcaption>Yamanaka and <a href="/wiki/Ry%C5%8Dji_Noyori" title="Ryōji Noyori">Ryōji Noyori</a> participating in the ceremony of the <a href="/wiki/50th_All_Japan_Rugby_Football_Championship" title="50th All Japan Rugby Football Championship">50th All Japan Rugby Football Championship</a></figcaption></figure> <p><b>Shinya Yamanaka</b><span style="font-weight: normal"> (<span title="Japanese-language text"><span lang="ja">山中伸弥</span></span>, <span title="Hepburn transliteration"><i lang="ja-Latn">Yamanaka Shin'ya</i></span>, born September 4, 1962)</span> is a Japanese <a href="/wiki/Stem_cell" title="Stem cell">stem cell</a> researcher and a <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> laureate.<sup id="cite_ref-stem1_2-1" class="reference"><a href="#cite_note-stem1-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-stem2_3-1" class="reference"><a href="#cite_note-stem2-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-stem3_4-1" class="reference"><a href="#cite_note-stem3-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> He is a professor and the director emeritus of Center for iPS Cell (<a href="/wiki/Induced_Pluripotent_Stem_Cell" class="mw-redirect" title="Induced Pluripotent Stem Cell">induced Pluripotent Stem Cell</a>) Research and Application, <a href="/wiki/Kyoto_University" title="Kyoto University">Kyoto University</a>;<sup id="cite_ref-:2_6-0" class="reference"><a href="#cite_note-:2-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> as a senior investigator at the <a href="/wiki/UCSF" class="mw-redirect" title="UCSF">UCSF</a>-affiliated <a href="/wiki/Gladstone_Institutes" title="Gladstone Institutes">Gladstone Institutes</a> in San Francisco, California; and as a professor of anatomy at <a href="/wiki/University_of_California,_San_Francisco" title="University of California, San Francisco">University of California, San Francisco</a> (UCSF). Yamanaka is also a past president of the <a href="/wiki/International_Society_for_Stem_Cell_Research" title="International Society for Stem Cell Research">International Society for Stem Cell Research</a> (ISSCR). </p><p>He received the 2010 <a href="/wiki/BBVA_Foundation_Frontiers_of_Knowledge_Award" title="BBVA Foundation Frontiers of Knowledge Award">BBVA Foundation Frontiers of Knowledge Award</a> in the biomedicine category, the 2011 <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a> with <a href="/wiki/Rudolf_Jaenisch" title="Rudolf Jaenisch">Rudolf Jaenisch</a>,<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> and the 2012 <a href="/wiki/Millennium_Technology_Prize" title="Millennium Technology Prize">Millennium Technology Prize</a> together with <a href="/wiki/Linus_Torvalds" title="Linus Torvalds">Linus Torvalds</a>. In 2012, he and <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> were awarded the <a href="/wiki/Nobel_Prize_for_Physiology_or_Medicine" class="mw-redirect" title="Nobel Prize for Physiology or Medicine">Nobel Prize for Physiology or Medicine</a> for the discovery that mature cells can be converted to <a href="/wiki/Stem_cells" class="mw-redirect" title="Stem cells">stem cells</a>.<sup id="cite_ref-:0_8-0" class="reference"><a href="#cite_note-:0-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> In 2013, he was awarded the $3 million <a href="/wiki/Breakthrough_Prize_in_Life_Sciences" title="Breakthrough Prize in Life Sciences">Breakthrough Prize in Life Sciences</a> for his work. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Education">Education</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=1" title="Edit section: Education"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Yamanaka was born in <a href="/wiki/Higashi%C5%8Dsaka,_Osaka" class="mw-redirect" title="Higashiōsaka, Osaka">Higashiōsaka</a>, Japan, in 1962. After graduating from Tennōji High School attached to <a href="/wiki/Osaka_Kyoiku_University" title="Osaka Kyoiku University">Osaka Kyoiku University</a>, he received his <a href="/wiki/Medical_Doctor" class="mw-redirect" title="Medical Doctor">M.D.</a> degree at <a href="/wiki/Kobe_University" title="Kobe University">Kobe University</a> in 1987 and his <a href="/wiki/Doctor_of_Philosophy" title="Doctor of Philosophy">Ph.D.</a> degree at <a href="/wiki/Osaka_City_University#Graduate_schools" title="Osaka City University">Osaka City University, Graduate School</a> of Medicine in 1993. After this, he went through a residency in orthopedic surgery at National Osaka Hospital and a <a href="/wiki/Postdoctoral" class="mw-redirect" title="Postdoctoral">postdoctoral</a> fellowship at the Gladstone Institute of <a href="/wiki/Cardiovascular_disease" title="Cardiovascular disease">Cardiovascular disease</a>, San Francisco. </p><p>Afterwards, he worked at the Gladstone Institutes in San Francisco, US, and <a href="/wiki/Nara_Institute_of_Science_and_Technology" title="Nara Institute of Science and Technology">Nara Institute of Science and Technology</a> in Japan. Yamanaka is currently a professor and the director emeritus of Center for iPS Research and Application (CiRA), Kyoto University.<sup id="cite_ref-:2_6-1" class="reference"><a href="#cite_note-:2-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> He is also a senior investigator at the Gladstone Institutes.<sup id="cite_ref-:0_8-1" class="reference"><a href="#cite_note-:0-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Professional_career">Professional career</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=2" title="Edit section: Professional career"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Between 1987 and 1989, Yamanaka was a <a href="/wiki/Residency_(medicine)" title="Residency (medicine)">resident</a> in orthopedic surgery at the National Osaka Hospital. His first operation was to remove a benign tumor from his friend Shuichi Hirata, a task he could not complete after one hour when a skilled surgeon would have taken ten minutes or so. Some <a href="/wiki/Senpai_and_k%C5%8Dhai" title="Senpai and kōhai">seniors</a> referred to him as "Jamanaka", a pun on the Japanese word for obstacle.<sup id="cite_ref-ajw.asahi.com_10-0" class="reference"><a href="#cite_note-ajw.asahi.com-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p><p>From 1993 to 1996, he was at the <a href="/wiki/Gladstone_Institutes" title="Gladstone Institutes">Gladstone Institute</a> of Cardiovascular disease. Between 1996 and 1999, he was an assistant professor at Osaka City University Medical School, but found himself mostly looking after mice in the laboratory, not doing actual research.<sup id="cite_ref-ajw.asahi.com_10-1" class="reference"><a href="#cite_note-ajw.asahi.com-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p><p>His wife advised him to become a practicing doctor, but instead he applied for a position at the <a href="/wiki/Nara_Institute_of_Science_and_Technology" title="Nara Institute of Science and Technology">Nara Institute of Science and Technology</a>. He stated that he could and would clarify the characteristics of embryonic stem cells, and this can-do attitude won him the job. From 1999 to 2003, he was an <a href="/wiki/Associate_professor" title="Associate professor">associate professor</a> there, and started the research that would later win him the 2012 Nobel Prize. He became a full professor and remained at the institute in that position from 2003 to 2005. Between 2004 and 2010, Yamanaka was a professor at the Institute for Frontier Medical Sciences, Kyoto University.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Between 2010 and 2022, Yamanaka was the director and a professor at the center for iPS Cell Research and Application (CiRA), Kyoto University.<sup id="cite_ref-:2_6-2" class="reference"><a href="#cite_note-:2-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> In April 2022, he stepped down and took place of the director emeritus of CiRA keeping with professor position.<sup id="cite_ref-:2_6-3" class="reference"><a href="#cite_note-:2-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p><p>In 2006, he and his team generated <a href="/wiki/Induced_pluripotent_stem_cell" title="Induced pluripotent stem cell">induced pluripotent stem cells</a> (iPS cells) from adult mouse <a href="/wiki/Fibroblast" title="Fibroblast">fibroblasts</a>.<sup id="cite_ref-stem1_2-2" class="reference"><a href="#cite_note-stem1-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> iPS cells closely resemble <a href="/wiki/Embryonic_stem_cell" title="Embryonic stem cell">embryonic stem cells</a>, the <i>in vitro</i> equivalent of the part of the <a href="/wiki/Blastocyst" title="Blastocyst">blastocyst</a> (the embryo a few days after fertilization) which grows to become the embryo proper. They could show that his iPS cells were <a href="/wiki/Pluripotent" class="mw-redirect" title="Pluripotent">pluripotent</a>, i.e. capable of generating all cell lineages of the body. Later he and his team generated iPS cells from human adult fibroblasts,<sup id="cite_ref-stem2_3-2" class="reference"><a href="#cite_note-stem2-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> again as the first group to do so. A key difference from previous attempts by the field was his team's use of multiple <a href="/wiki/Transcription_factor" title="Transcription factor">transcription factors</a>, instead of <a href="/wiki/Transfection" title="Transfection">transfecting</a> one transcription factor per experiment. They started with 24 transcription factors known to be important in the early embryo, but could in the end reduce it to four transcription factors – <a href="/wiki/Sox2" class="mw-redirect" title="Sox2">Sox2</a>, <a href="/wiki/Oct4" class="mw-redirect" title="Oct4">Oct4</a>, <a href="/wiki/Klf4" class="mw-redirect" title="Klf4">Klf4</a> and <a href="/wiki/C-Myc" class="mw-redirect" title="C-Myc">c-Myc</a>.<sup id="cite_ref-stem1_2-3" class="reference"><a href="#cite_note-stem1-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Yamanaka's_Nobel_Prize–winning_research_in_iPS_cells"><span id="Yamanaka.27s_Nobel_Prize.E2.80.93winning_research_in_iPS_cells"></span>Yamanaka's Nobel Prize–winning research in iPS cells</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=3" title="Edit section: Yamanaka's Nobel Prize–winning research in iPS cells"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The <b>2012 Nobel Prize in Physiology or Medicine</b> was awarded jointly to Sir John B. Gurdon and Shinya Yamanaka "for the discovery that mature cells can be reprogrammed to become <a href="/wiki/Pluripotent" class="mw-redirect" title="Pluripotent">pluripotent</a>."<sup id="cite_ref-:0_8-2" class="reference"><a href="#cite_note-:0-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Background-different_cell_types">Background-different cell types</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=4" title="Edit section: Background-different cell types"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div><p> There are different types of stem cells. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1251242444"></p><table class="box-Example_farm plainlinks metadata ambox ambox-content" role="presentation"><tbody><tr><td class="mbox-image"><div class="mbox-image-div"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/b/b4/Ambox_important.svg/40px-Ambox_important.svg.png" decoding="async" width="40" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/b/b4/Ambox_important.svg/60px-Ambox_important.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/b/b4/Ambox_important.svg/80px-Ambox_important.svg.png 2x" data-file-width="40" data-file-height="40" /></span></span></div></td><td class="mbox-text"><div class="mbox-text-span">This article <b>may contain <a href="/wiki/Wikipedia:Manual_of_Style/Lists#List_size" title="Wikipedia:Manual of Style/Lists">excessive</a> or <a href="/wiki/Wikipedia:Neutral_point_of_view#Due_and_undue_weight" title="Wikipedia:Neutral point of view">irrelevant</a> examples</b>.<span class="hide-when-compact"> Please help <a class="external text" href="https://en.wikipedia.org/w/index.php?title=Shinya_Yamanaka&action=edit">improve the article</a> by adding descriptive text and removing <a href="/wiki/Wikipedia:Example_cruft" title="Wikipedia:Example cruft">less pertinent examples</a>.</span> <span class="date-container"><i>(<span class="date">May 2016</span>)</i></span></div></td></tr></tbody></table><p> These are some types of cells that will help in understanding the material. </p><table class="wikitable"> <tbody><tr> <th>Cell types</th> <th>Characteristics </th></tr> <tr> <td>Totipotent cells</td> <td>Can give rise to all other cell types. Totipotency remains through the first few cell divisions, e.g. the fertilised egg. </td></tr> <tr> <td>Pluripotent cells</td> <td>Can develop into all cell types (except those that form the amniotic sac and the placenta). For example, the early embryo consists mainly of pluripotent stem cells. </td></tr> <tr> <td>Multipotent cells</td> <td>Can develop into any of a family of closely related cell types. For example, blood multipotent cells can develop into various blood cells. </td></tr></tbody></table> <div class="mw-heading mw-heading3"><h3 id="Background-different_stem_cell_techniques">Background-different stem cell techniques</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=5" title="Edit section: Background-different stem cell techniques"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <table class="wikitable"> <tbody><tr> <th>Type</th> <th>Pros</th> <th>Cons </th></tr> <tr> <td>Somatic cell replication Embryonic Stem (ES) cell</td> <td>No immune rejection <p>Theoretically patient-specific transplantations possible </p> </td> <td>No case of success<br />Many human egg cells needed<br />Ethical issue: Can clone humans </td></tr> <tr> <td>Fertilized egg ES cell</td> <td>Pluripotent <p>Much research done </p><p>Immune rejection reducible via stem cell bank </p> </td> <td>Fertilized egg usage<br />Immune rejection<br />Oncogenic potential<br />(can't use for clinical trial) </td></tr> <tr> <td>Induced pluripotent stem (iPS) Cell </td> <td>No ethical issue <p>Pluripotent </p> </td> <td>Oncogenic potential <p>Abnormal aging </p> </td></tr> <tr> <td>Adult stem cell</td> <td>Much research <p>No immune rejection Safe (clinical trials) </p> </td> <td>Not as much differentiation potential as ES cell </td></tr></tbody></table> <div class="mw-heading mw-heading3"><h3 id="Historical_background">Historical background</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=6" title="Edit section: Historical background"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The prevalent view during the early 20th century was that mature cells were permanently locked into the differentiated state and cannot return to a fully immature, pluripotent stem cell state. It was thought that cellular differentiation can only be a unidirectional process. Therefore, non-differentiated egg/early embryo cells can only develop into specialized cells. However, stem cells with limited potency (adult stem cells) remain in bone marrow, intestine, skin etc. to act as a source of cell replacement.<sup id="cite_ref-:1_13-0" class="reference"><a href="#cite_note-:1-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>The fact that differentiated cell types had specific patterns of proteins suggested irreversible epigenetic modifications or genetic alterations to be the cause of unidirectional cell differentiation. So, cells progressively become more restricted in the differentiation potential and eventually lose pluripotency.<sup id="cite_ref-:1_13-1" class="reference"><a href="#cite_note-:1-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>In 1962, <a href="/wiki/John_Gurdon" title="John Gurdon">John B. Gurdon</a> demonstrated that the nucleus from a differentiated frog intestinal epithelial cell can generate a fully functional tadpole via transplantation to an enucleated egg. Gurdon used <a href="/wiki/Somatic_cell_nuclear_transfer" title="Somatic cell nuclear transfer">somatic cell nuclear transfer</a> (SCNT) as a method to understand reprogramming and how cells change in specialization. He concluded that differentiated somatic cell nuclei had the potential to revert to pluripotency. This was a paradigm shift at the time. It showed that a differentiated cell nucleus has retained the capacity to successfully revert to an undifferentiated state, with the potential to restart development (pluripotent capacity). </p><p>However, the question still remained whether an intact differentiated cell could be fully reprogrammed to become pluripotent. </p> <div class="mw-heading mw-heading3"><h3 id="Yamanaka's_research"><span id="Yamanaka.27s_research"></span>Yamanaka's research</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=7" title="Edit section: Yamanaka's research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Shinya Yamanaka proved that introduction of a small set of transcription factors into a differentiated cell was sufficient to revert the cell to a pluripotent state. Yamanaka focused on factors that are important for maintaining pluripotency in embryonic stem (ES) cells. This was the first time an intact differentiated somatic cell could be reprogrammed to become pluripotent. </p><p>Knowing that transcription factors were involved in the maintenance of the pluripotent state, he selected a set of 24 ES cell transcriptional factors as candidates to reinstate pluripotency in somatic cells. First, he collected the 24 candidate factors. When all 24 genes encoding these transcription factors were introduced into skin fibroblasts, few actually generated colonies that were remarkably similar to ES cells. Secondly, further experiments were conducted with smaller numbers of transcription factors added to identify the key factors, through a very simple and yet sensitive assay system. Lastly, he identified the four key genes. They found that 4 transcriptional factors (Myc, Oct3/4, Sox2 and Klf4) were sufficient to convert mouse embryonic or adult fibroblasts to pluripotent stem cells (capable of producing teratomas <i>in vivo</i> and contributing to chimeric mice). </p><p>These pluripotent cells are called iPS (induced pluripotent stem) cells; they appeared with very low frequency. iPS cells can be selected by inserting the b-geo gene into the Fbx15 locus. The Fbx15 promoter is active in pluripotent stem cells which induce b-geo expression, which in turn gives rise to G418 resistance; this resistance helps us identify the iPS cells in culture. </p><p>Moreover, in 2007, Yamanaka and his colleagues found iPS cells with germline transmission (via selecting for Oct4 or Nanog gene). Also in 2007, they were the first to produce human iPS cells. </p><p>Some issues that current methods of induced pluripotency face are the very low production rate of iPS cells and the fact that the 4 transcriptional factors are shown to be oncogenic. </p><p>In July 2014, during a scandal involving Japanese stem cell researcher <a href="/wiki/Haruko_Obokata" title="Haruko Obokata">Haruko Obokata</a> fabricating data, doctoring images, and plagiarizing the work of others, Yamanaka faced public scrutiny for his associated work lacking full documentation. Yamanaka denied manipulating images in his papers on embryonic mouse stem cells, but he could not find lab notes to confirm that the raw data was consistent with the published results.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Further_research_and_future_prospects">Further research and future prospects</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=8" title="Edit section: Further research and future prospects"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Since the original discovery by Yamanaka, much further research has been done in this field, and many improvements have been made to the technology. Improvements made to Yamanaka's research as well as future prospects of his findings are as follows: </p> <ol><li>The delivery mechanism of pluripotency factors has been improved. At first retroviral vectors, that integrate randomly in the genome and cause deregulation of genes that contribute to tumor formation, were used. However, now, non-integrating viruses, stabilised RNAs or proteins, or episomal plasmids (integration-free delivery mechanism) are used.</li> <li>Transcription factors required for inducing pluripotency in different cell types have been identified (e.g. neural stem cells).</li> <li>Small substitutive molecules were identified, that can substitute for the function of the transcription factors.</li> <li><a href="/wiki/Transdifferentiation" title="Transdifferentiation">Transdifferentiation</a> experiments were carried out. They tried to change the cell fate without proceeding through a pluripotent state. They were able to systematically identify genes that carry out transdifferentiation using combinations of transcription factors that induce cell fate switches. They found trandifferentiation within germ layer and between germ layers, e.g., exocrine cells to endocrine cells, fibroblast cells to myoblast cells, fibroblast cells to cardiomyocyte cells, fibroblast cells to neurons</li> <li><a href="/wiki/Cell_replacement_therapy" class="mw-redirect" title="Cell replacement therapy">Cell replacement therapy</a> with iPS cells is a possibility. Stem cells can replace diseased or lost cells in degenerative disorders and they are less prone to immune rejection. However, there is a danger that it may introduce mutations or other genomic abnormalities that render it unsuitable for cell therapy. So, there are still many challenges, but it is a very exciting and promising research area. Further work is required to guarantee safety for patients.</li> <li>Can medically use iPS cells from patients with genetic and other disorders to gain insights into the disease process. - Amyotrophic lateral sclerosis (ALS), Rett syndrome, spinal muscular atrophy (SMA), α1-antitrypsin deficiency, familial hypercholesterolemia and glycogen storage disease type 1A. - For cardiovascular disease, Timothy syndrome, LEOPARD syndrome, type 1 and 2 long QT syndrome - Alzheimer's, Spinocerebellar ataxia, Huntington's etc.</li> <li>iPS cells provide screening platforms for development and validation of therapeutic compounds. For example, kinetin was a novel compound found in iPS cells from familial dysautonomia and beta blockers & ion channel blockers for long QT syndrome were identified with iPS cells.</li></ol> <p>Yamanaka's research has "opened a new door and the world's scientists have set forth on a long journey of exploration, hoping to find our cells’ true potential."<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p><p>In 2013, iPS cells were used to generate a human vascularized and functional liver in mice in Japan. Multiple stem cells were used to differentiate the component parts of the liver, which then self-organized into the complex structure. When placed into a mouse host, the liver vessels connected to the hosts vessels and performed normal liver functions, including breaking down of drugs and liver secretions.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p><p>In 2022, Yamanaka factors were shown to effect age related measures in aged mice.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Recognition">Recognition</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=9" title="Edit section: Recognition"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In 2007, Yamanaka was recognized as a "Person Who Mattered" in the <a href="/wiki/Time_Person_of_the_Year" title="Time Person of the Year">Time Person of the Year</a> edition of <i><a href="/wiki/Time_(magazine)" title="Time (magazine)">Time</a></i> magazine.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> Yamanaka was also nominated as a 2008 <a href="/wiki/Time_100" title="Time 100">Time 100</a> Finalist.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> In June 2010, Yamanaka was awarded the <a href="/wiki/Kyoto_Prize" title="Kyoto Prize">Kyoto Prize</a> for reprogramming adult skin cells to pluripotential precursors. Yamanaka developed the method as an alternative to embryonic stem cells, thus circumventing an approach in which embryos would be destroyed. </p><p>In May 2010, Yamanaka was given "Doctor of Science honorary degree" by <a href="/wiki/Mount_Sinai_Hospital,_New_York" class="mw-redirect" title="Mount Sinai Hospital, New York">Mount Sinai School of Medicine</a>.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> </p><p>In September 2010, he was awarded the <a href="/wiki/Balzan_Prize" title="Balzan Prize">Balzan Prize</a> for his work on biology and stem cells.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p><p>Yamanaka has been listed as one of the 15 Asian Scientists To Watch by <i><a href="/wiki/Asian_Scientist" title="Asian Scientist">Asian Scientist</a></i> magazine on May 15, 2011.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> In June 2011, he was awarded the inaugural McEwen Award for Innovation; he shared the $100,000 prize with Kazutoshi Takahashi, who was the lead author on the paper describing the generation of induced pluripotent stem cells.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup> </p><p>In June 2012, he was awarded the <a href="/wiki/Millennium_Technology_Prize" title="Millennium Technology Prize">Millennium Technology Prize</a> for his work in stem cells.<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup> He shared the 1.2 million euro prize with <a href="/wiki/Linus_Torvalds" title="Linus Torvalds">Linus Torvalds</a>, the creator of the Linux kernel. In October 2012, he and fellow stem cell researcher <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> were awarded the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> "for the discovery that mature cells can be reprogrammed to become pluripotent."<sup id="cite_ref-NobelPrize_28-0" class="reference"><a href="#cite_note-NobelPrize-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col"> <ul><li>2007 – Osaka Science Prize<sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup></li> <li>2007 – Inoue Prize for Science<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (October 2012)">citation needed</span></a></i>]</sup></li> <li>2007 – <a href="/wiki/Asahi_Prize" title="Asahi Prize">Asahi Prize</a></li> <li>2007 – <a href="/wiki/Meyenburg_Cancer_Research_Award" class="mw-redirect" title="Meyenburg Cancer Research Award">Meyenburg Cancer Research Award</a><sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-31" class="reference"><a href="#cite_note-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup></li> <li>2008 – <a href="/wiki/Yamazaki-Teiichi_Prize" title="Yamazaki-Teiichi Prize">Yamazaki-Teiichi Prize</a> in <a rel="nofollow" class="external text" href="http://www.mst.or.jp/prize/2008/bio_en.html">Biological Science & Technology</a></li> <li>2008 – <a href="/wiki/Robert_Koch_Prize" class="mw-redirect" title="Robert Koch Prize">Robert Koch Prize</a></li> <li>2008 – <a href="/wiki/Medals_of_Honor_(Japan)" title="Medals of Honor (Japan)">Medals of Honor (Japan)</a> (with purple ribbon)</li> <li>2008 – <a href="/wiki/Shaw_Prize" title="Shaw Prize">Shaw Prize</a> in Life Science & Medicine<sup id="cite_ref-32" class="reference"><a href="#cite_note-32"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-33" class="reference"><a href="#cite_note-33"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-34" class="reference"><a href="#cite_note-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup></li> <li>2008 – Sankyo Takamine Memorial Award<sup id="cite_ref-35" class="reference"><a href="#cite_note-35"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup></li> <li>2008 – <a href="/wiki/Massry_Prize" title="Massry Prize">Massry Prize</a> from the <a href="/wiki/Keck_School_of_Medicine" class="mw-redirect" title="Keck School of Medicine">Keck School of Medicine</a>, <a href="/wiki/University_of_Southern_California" title="University of Southern California">University of Southern California</a></li> <li>2008 - Golden Plate Award of the <a href="/wiki/Academy_of_Achievement" title="Academy of Achievement">American Academy of Achievement</a><sup id="cite_ref-36" class="reference"><a href="#cite_note-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup></li> <li>2009 – Lewis S. <a href="/wiki/Rosenstiel_Award" title="Rosenstiel Award">Rosenstiel Award</a> for Distinguished Work in Basic Medical Research<sup id="cite_ref-37" class="reference"><a href="#cite_note-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup></li> <li>2009 – <a href="/wiki/Gairdner_Foundation_International_Award" class="mw-redirect" title="Gairdner Foundation International Award">Gairdner Foundation International Award</a><sup id="cite_ref-38" class="reference"><a href="#cite_note-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup></li> <li>2009 – <a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award for Basic Medical Research</a></li> <li>2010 – <a href="/wiki/Balzan_Prize" title="Balzan Prize">Balzan Prize</a> for Stem Cells: Biology and potential applications</li> <li>2010 – <a href="/wiki/March_of_Dimes_Prize_in_Developmental_Biology" title="March of Dimes Prize in Developmental Biology">March of Dimes Prize in Developmental Biology</a></li> <li>2010 – <a href="/wiki/Kyoto_Prize" title="Kyoto Prize">Kyoto Prize</a> in Biotechnology and medical technology</li> <li>2010 – <a href="/wiki/Person_of_Cultural_Merit" title="Person of Cultural Merit">Person of Cultural Merit</a></li> <li>2010 – BBVA Foundation Frontiers of Knowledge Award in the Biomedicine Category<sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">[</span>39<span class="cite-bracket">]</span></a></sup></li> <li>2011 – <a href="/wiki/Albany_Medical_Center_Prize" title="Albany Medical Center Prize">Albany Medical Center Prize</a> in biomedicine</li> <li>2011 – <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a></li> <li>2011 – <a href="/wiki/King_Faisal_International_Prize#Medicine" class="mw-redirect" title="King Faisal International Prize">King Faisal International Prize for Medicine</a></li> <li>2011 – <a rel="nofollow" class="external text" href="https://web.archive.org/web/20121008001055/http://www.isscr.org/McEwen_Award_for_Innovation/5323.htm">McEwen Award for Innovation</a></li> <li>2012 – <a href="/wiki/Millennium_Technology_Prize" title="Millennium Technology Prize">Millennium Technology Prize</a></li> <li>2012 – Fellow of the <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">National Academy of Sciences</a><sup id="cite_ref-nas_1-1" class="reference"><a href="#cite_note-nas-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></li> <li>2012 – <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a><sup id="cite_ref-NobelPrize_28-1" class="reference"><a href="#cite_note-NobelPrize-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup></li> <li>2012 – <a href="/wiki/Order_of_Culture" title="Order of Culture">Order of Culture</a></li> <li>2013 – <a href="/wiki/Breakthrough_Prize_in_Life_Sciences" title="Breakthrough Prize in Life Sciences">Breakthrough Prize in Life Sciences</a></li> <li>2013 – Member of the <a href="/wiki/Pontifical_Academy_of_Sciences" title="Pontifical Academy of Sciences">Pontifical Academy of Sciences</a><sup id="cite_ref-40" class="reference"><a href="#cite_note-40"><span class="cite-bracket">[</span>40<span class="cite-bracket">]</span></a></sup></li> <li>2014 – <a href="/wiki/UCSF" class="mw-redirect" title="UCSF">UCSF</a> 150th Anniversary Alumni Excellence Awards<sup id="cite_ref-41" class="reference"><a href="#cite_note-41"><span class="cite-bracket">[</span>41<span class="cite-bracket">]</span></a></sup></li> <li>2016 – Honorable Emeritus Professor, <a href="/wiki/Hiroshima_University" title="Hiroshima University">Hiroshima University</a></li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="Interest_in_sports">Interest in sports</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=10" title="Edit section: Interest in sports"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Yamanaka practiced <a href="/wiki/Judo" title="Judo">judo</a> (<a href="/wiki/Dan_(rank)#Ranks_in_Japanese" title="Dan (rank)">2nd Dan</a> black belt) and played rugby as a university student. He also has a history of running marathons. After a 20-year gap, he competed in the inaugural <a href="/wiki/Osaka_Marathon" title="Osaka Marathon">Osaka Marathon</a> in 2011 as a charity runner with a time of 4:29:53. He took part in <a href="/wiki/Kyoto_Marathon" title="Kyoto Marathon">Kyoto Marathon</a> to raise money for iPS research since 2012. His personal best is 3:25:20 at 2018 <a href="/wiki/Beppu-%C5%8Cita_Marathon" title="Beppu-Ōita Marathon">Beppu-Ōita Marathon</a>. </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=11" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1266661725">.mw-parser-output .portalbox{padding:0;margin:0.5em 0;display:table;box-sizing:border-box;max-width:175px;list-style:none}.mw-parser-output .portalborder{border:1px solid var(--border-color-base,#a2a9b1);padding:0.1em;background:var(--background-color-neutral-subtle,#f8f9fa)}.mw-parser-output .portalbox-entry{display:table-row;font-size:85%;line-height:110%;height:1.9em;font-style:italic;font-weight:bold}.mw-parser-output .portalbox-image{display:table-cell;padding:0.2em;vertical-align:middle;text-align:center}.mw-parser-output .portalbox-link{display:table-cell;padding:0.2em 0.2em 0.2em 0.3em;vertical-align:middle}@media(min-width:720px){.mw-parser-output .portalleft{margin:0.5em 1em 0.5em 0}.mw-parser-output .portalright{clear:right;float:right;margin:0.5em 0 0.5em 1em}}</style><ul role="navigation" aria-label="Portals" class="noprint portalbox portalborder portalright"> <li class="portalbox-entry"><span class="portalbox-image"><span class="mw-image-border noviewer" typeof="mw:File"><span><img alt="flag" src="//upload.wikimedia.org/wikipedia/en/thumb/9/9e/Flag_of_Japan.svg/32px-Flag_of_Japan.svg.png" decoding="async" width="32" height="21" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/9e/Flag_of_Japan.svg/48px-Flag_of_Japan.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/9e/Flag_of_Japan.svg/64px-Flag_of_Japan.svg.png 2x" data-file-width="900" data-file-height="600" /></span></span></span><span class="portalbox-link"><a href="/wiki/Portal:Japan" title="Portal:Japan">Japan portal</a></span></li><li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Stylised_atom_with_three_Bohr_model_orbits_and_stylised_nucleus.svg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6f/Stylised_atom_with_three_Bohr_model_orbits_and_stylised_nucleus.svg/25px-Stylised_atom_with_three_Bohr_model_orbits_and_stylised_nucleus.svg.png" decoding="async" width="25" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6f/Stylised_atom_with_three_Bohr_model_orbits_and_stylised_nucleus.svg/37px-Stylised_atom_with_three_Bohr_model_orbits_and_stylised_nucleus.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6f/Stylised_atom_with_three_Bohr_model_orbits_and_stylised_nucleus.svg/49px-Stylised_atom_with_three_Bohr_model_orbits_and_stylised_nucleus.svg.png 2x" data-file-width="530" data-file-height="600" /></a></span></span><span class="portalbox-link"><a href="/wiki/Portal:Physics" title="Portal:Physics">Physics portal</a></span></li><li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/6/69/P_vip.svg/28px-P_vip.svg.png" decoding="async" width="28" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/6/69/P_vip.svg/41px-P_vip.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/6/69/P_vip.svg/55px-P_vip.svg.png 2x" data-file-width="1911" data-file-height="1944" /></span></span></span><span class="portalbox-link"><a href="/wiki/Portal:Biography" title="Portal:Biography">Biography portal</a></span></li></ul> <ul><li><a href="/wiki/Catherine_Verfaillie" title="Catherine Verfaillie">Catherine Verfaillie</a></li> <li><a href="/wiki/List_of_Japanese_Nobel_laureates" class="mw-redirect" title="List of Japanese Nobel laureates">List of Japanese Nobel laureates</a></li> <li><a href="/wiki/List_of_Nobel_laureates_affiliated_with_Kyoto_University" class="mw-redirect" title="List of Nobel laureates affiliated with Kyoto University">List of Nobel laureates affiliated with Kyoto University</a></li> <li><a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=12" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>General references: </p> <style data-mw-deduplicate="TemplateStyles:r1239549316">.mw-parser-output .refbegin{margin-bottom:0.5em}.mw-parser-output .refbegin-hanging-indents>ul{margin-left:0}.mw-parser-output .refbegin-hanging-indents>ul>li{margin-left:0;padding-left:3.2em;text-indent:-3.2em}.mw-parser-output .refbegin-hanging-indents ul,.mw-parser-output .refbegin-hanging-indents ul li{list-style:none}@media(max-width:720px){.mw-parser-output .refbegin-hanging-indents>ul>li{padding-left:1.6em;text-indent:-1.6em}}.mw-parser-output .refbegin-columns{margin-top:0.3em}.mw-parser-output .refbegin-columns ul{margin-top:0}.mw-parser-output .refbegin-columns li{page-break-inside:avoid;break-inside:avoid-column}@media screen{.mw-parser-output .refbegin{font-size:90%}}</style><div class="refbegin" style=""> <ul><li><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFFackler2007" class="citation news cs1">Fackler, Martin (December 11, 2007). <a rel="nofollow" class="external text" href="https://www.nytimes.com/2007/12/11/science/11prof.html?pagewanted=all">"Risk Taking Is in His Genes"</a>. <i>The New York Times</i><span class="reference-accessdate">. Retrieved <span class="nowrap">December 11,</span> 2007</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+New+York+Times&rft.atitle=Risk+Taking+Is+in+His+Genes&rft.date=2007-12-11&rft.aulast=Fackler&rft.aufirst=Martin&rft_id=https%3A%2F%2Fwww.nytimes.com%2F2007%2F12%2F11%2Fscience%2F11prof.html%3Fpagewanted%3Dall&rfr_id=info%3Asid%2Fen.wikipedia.org%3AShinya+Yamanaka" class="Z3988"></span></li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20090208224931/http://www.tv.janjan.jp/movie/edit/fccj/080109fccj_shinya_yamanakai_v_01.php">The Discovery and Future of Induced Pluripotent Stem (iPS)</a></li> <li><a rel="nofollow" class="external text" href="https://www.nytimes.com/2012/10/09/health/research/cloning-and-stem-cell-discoveries-earn-nobel-prize-in-medicine.html">Cloning and Stem Cell Discoveries Earn Nobel in Medicine (New York Times, October 8, 2012)</a></li></ul> </div> <p>Specific citations: </p> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-nas-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-nas_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-nas_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFNair2012" class="citation journal cs1">Nair, P. 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Retrieved on 2012-10-08.</span> </li> <li id="cite_note-NobelPrize-28"><span class="mw-cite-backlink">^ <a href="#cite_ref-NobelPrize_28-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-NobelPrize_28-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.nobelprize.org/nobel_prizes/medicine/laureates/2012/">"The Nobel Prize in Physiology or Medicine 2012"</a>. NobelPrize.org. October 8, 2012<span class="reference-accessdate">. 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Archived from <a rel="nofollow" class="external text" href="https://gladstone.org/our-science/cardiovascular-disease/honors-and-awards?page=1">the original</a> on October 29, 2016<span class="reference-accessdate">. Retrieved <span class="nowrap">October 27,</span> 2016</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Honors+and+Awards&rft_id=https%3A%2F%2Fgladstone.org%2Four-science%2Fcardiovascular-disease%2Fhonors-and-awards%3Fpage%3D1&rfr_id=info%3Asid%2Fen.wikipedia.org%3AShinya+Yamanaka" class="Z3988"></span></span> </li> <li id="cite_note-30"><span class="mw-cite-backlink"><b><a href="#cite_ref-30">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.dkfz.de/en/presse/pressemitteilungen/2007/dkfz_pm_07_72_e.php">"Meyenburg Cancer Research Award 2007 given to Dr. Shinya Yamanaka for the artificial generation of stem cells"</a>. <i>German Cancer Research Center</i>. November 23, 2007.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=German+Cancer+Research+Center&rft.atitle=Meyenburg+Cancer+Research+Award+2007+given+to+Dr.+Shinya+Yamanaka+for+the+artificial+generation+of+stem+cells&rft.date=2007-11-23&rft_id=http%3A%2F%2Fwww.dkfz.de%2Fen%2Fpresse%2Fpressemitteilungen%2F2007%2Fdkfz_pm_07_72_e.php&rfr_id=info%3Asid%2Fen.wikipedia.org%3AShinya+Yamanaka" class="Z3988"></span></span> </li> <li id="cite_note-31"><span class="mw-cite-backlink"><b><a href="#cite_ref-31">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20130327090130/http://bio-pro.de/magazin/wissenschaft/archiv_2007/index.html?lang=en">"Meyenburg Cancer Research Award 2007 for artificial stem cell production"</a>. <i>German Cancer Research Center</i>. November 23, 2007. Archived from <a rel="nofollow" class="external text" href="http://www.bio-pro.de/magazin/wissenschaft/archiv_2007/index.html?lang=en&artikelid=/artikel/01983/index.html">the original</a> on March 27, 2013.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=German+Cancer+Research+Center&rft.atitle=Meyenburg+Cancer+Research+Award+2007+for+artificial+stem+cell+production&rft.date=2007-11-23&rft_id=http%3A%2F%2Fwww.bio-pro.de%2Fmagazin%2Fwissenschaft%2Farchiv_2007%2Findex.html%3Flang%3Den%26artikelid%3D%2Fartikel%2F01983%2Findex.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AShinya+Yamanaka" class="Z3988"></span></span> </li> <li id="cite_note-32"><span class="mw-cite-backlink"><b><a href="#cite_ref-32">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFTucker,_Valerie2008" class="citation web cs1">Tucker, Valerie (June 11, 2008). <a rel="nofollow" class="external text" href="http://pub.ucsf.edu/today/cache/feature/200806103.html">"Gladstone's Shinya Yamanaka Wins Prestigious Shaw Prize for Stem Cell Discoveries"</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Gladstone%27s+Shinya+Yamanaka+Wins+Prestigious+Shaw+Prize+for+Stem+Cell+Discoveries&rft.date=2008-06-11&rft.au=Tucker%2C+Valerie&rft_id=http%3A%2F%2Fpub.ucsf.edu%2Ftoday%2Fcache%2Ffeature%2F200806103.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AShinya+Yamanaka" class="Z3988"></span></span> </li> <li id="cite_note-33"><span class="mw-cite-backlink"><b><a href="#cite_ref-33">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20200930143027/http://www.bio-medicine.org/biology-news-1/Gladstones-Shinya-Yamanaka-wins-prestigious-Shaw-Prize-3608-1/">"Gladstone's Shinya Yamanaka wins prestigious Shaw Prize"</a>. <i>Bio-Medicine.org</i>. 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Retrieved <span class="nowrap">July 31,</span> 2008</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Kyoto+University&rft.atitle=Professor+Shinya+Yamanaka+Awarded+the+Shaw+Prize&rft.date=2008-06-17&rft_id=http%3A%2F%2Fwww.kyoto-u.ac.jp%2Fen%2Fnews_data%2Fh%2Fh1%2F2008%2Fnews7%2F080617_2.htm&rfr_id=info%3Asid%2Fen.wikipedia.org%3AShinya+Yamanaka" class="Z3988"></span></span> </li> <li id="cite_note-35"><span class="mw-cite-backlink"><b><a href="#cite_ref-35">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20081028200838/http://www.icems.kyoto-u.ac.jp/cira/doc/080715_takamine_e.pdf">"Professor Shinya Granted Sixth Sankyo Takamine Memorial Award"</a> <span class="cs1-format">(PDF)</span>. <i>CiRA News Release</i>. July 15, 2008. 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Rosenstiel Award for Distinguished Work in Basic Medical Science"</a>. Archived from <a rel="nofollow" class="external text" href="http://www.rose.brandeis.edu/Center/rose_past.html">the original</a> on May 15, 2014.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Lewis+S.+Rosenstiel+Award+for+Distinguished+Work+in+Basic+Medical+Science&rft_id=http%3A%2F%2Fwww.rose.brandeis.edu%2FCenter%2Frose_past.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AShinya+Yamanaka" class="Z3988"></span></span> </li> <li id="cite_note-38"><span class="mw-cite-backlink"><b><a href="#cite_ref-38">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.gairdner.org/content/shinya-yamanaka">Shinya Yamanaka, Recipient of the Canada Gairdner International Award, 2009</a>. gairdner.org</span> </li> <li id="cite_note-39"><span class="mw-cite-backlink"><b><a href="#cite_ref-39">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.fbbva.es/awards">BBVA Foundation Frontiers of Knowledge Award</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20110703112316/http://www.fbbva.es/awards/">Archived</a> July 3, 2011, at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>. 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Archived from <a rel="nofollow" class="external text" href="http://www.ucsfalumni.org/s/1420/index_1col.aspx?sid=1420&gid=1&pgid=1610">the original</a> on June 12, 2021<span class="reference-accessdate">. Retrieved <span class="nowrap">January 18,</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=University+of+California%2C+San+Francisco&rft.atitle=UCSF+150th+Anniversary+Alumni+Excellence+Awards&rft.date=2014&rft.aulast=Minihane&rft.aufirst=Aine&rft_id=http%3A%2F%2Fwww.ucsfalumni.org%2Fs%2F1420%2Findex_1col.aspx%3Fsid%3D1420%26gid%3D1%26pgid%3D1610&rfr_id=info%3Asid%2Fen.wikipedia.org%3AShinya+Yamanaka" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Shinya_Yamanaka&action=edit&section=13" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Commons-logo.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png" decoding="async" width="30" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/45px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/59px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></a></span></div> <div class="side-box-text plainlist">Wikimedia Commons has media related to <span style="font-weight: bold; font-style: italic;"><a href="https://commons.wikimedia.org/wiki/Category:Shinya_Yamanaka" class="extiw" title="commons:Category:Shinya Yamanaka">Shinya Yamanaka</a></span>.</div></div> </div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1235681985"><div class="side-box metadata side-box-right"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Scholia_logo.svg/40px-Scholia_logo.svg.png" decoding="async" width="40" height="39" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Scholia_logo.svg/60px-Scholia_logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Scholia_logo.svg/80px-Scholia_logo.svg.png 2x" data-file-width="107" data-file-height="104" /></span></span></div> <div class="side-box-text plainlist"><a href="https://www.wikidata.org/wiki/Wikidata:Scholia" class="extiw" title="d:Wikidata:Scholia">Scholia</a> has an <i>author</i> profile for <i><b><a href="https://iw.toolforge.org/scholia/author/Q80917" class="extiw" title="toolforge:scholia/author/Q80917">Shinya Yamanaka</a></b></i>.</div></div> </div> <ul><li><a rel="nofollow" class="external text" href="https://www.cira.kyoto-u.ac.jp/e/research/yamanaka_summary.html?lang=en">Shinya Yamanaka</a>, Center for iPS Cell Research and Application (CiRA), Kyoto University</li> <li><a rel="nofollow" class="external text" href="https://www.isscr.org">International Society for Stem Cell Research</a> (ISSCR)</li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.balzan.org/en/prizewinners/shinya-yamanaka">"Shinya Yamanaka 2010 Balzan Prize for Stem Cells: Biology and Potential Applications"</a>. International Balzan Prize Foundation. 2010.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Shinya+Yamanaka+2010+Balzan+Prize+for+Stem+Cells%3A+Biology+and+Potential+Applications&rft.pub=International+Balzan+Prize+Foundation&rft.date=2010&rft_id=https%3A%2F%2Fwww.balzan.org%2Fen%2Fprizewinners%2Fshinya-yamanaka&rfr_id=info%3Asid%2Fen.wikipedia.org%3AShinya+Yamanaka" class="Z3988"></span></li> <li><a rel="nofollow" class="external text" href="https://charlierose.com/videos/13943">Shinya Yamanaka</a> on <a href="/wiki/Charlie_Rose_(talk_show)" title="Charlie Rose (talk show)"><i>Charlie Rose</i></a></li> <li><a rel="nofollow" class="external text" href="https://www.nobelprize.org/laureate/875">Shinya Yamanaka</a> on Nobelprize.org <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q80917#P8024" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></li></ul> <table class="wikitable succession-box noprint" style="margin:0.5em auto; font-size:small;clear:both;"> <tbody><tr> <th colspan="3" style="border-top: 5px solid #FFF179;">Awards </th></tr> <tr style="text-align:center;"> <td style="width:30%;" rowspan="1">Preceded by<div style="font-weight: bold"><a href="/wiki/Michael_Gr%C3%A4tzel" title="Michael Grätzel">Michael Grätzel</a></div> </td> <td style="width: 40%; text-align: center;" rowspan="1"><b> <a href="/wiki/Millennium_Technology_Prize" title="Millennium Technology Prize">Millennium Technology Prize</a> winner </b><br />2012 </td> <td style="width: 30%; text-align: center;" rowspan="1">Succeeded by<div style="font-weight: bold"><a href="/wiki/Stuart_Parkin" title="Stuart Parkin">Stuart Parkin</a></div> </td></tr> </tbody></table> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist 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Schmidt</a> (2006)</li> <li><a href="/wiki/Peter_Goldreich" title="Peter Goldreich">Peter Goldreich</a> (2007)</li> <li><a href="/wiki/Reinhard_Genzel" title="Reinhard Genzel">Reinhard Genzel</a> (2008)</li> <li><a href="/wiki/Frank_Shu" title="Frank Shu">Frank Shu</a> (2009)</li> <li><a href="/wiki/Charles_L._Bennett" title="Charles L. Bennett">Charles Bennett</a>, <a href="/wiki/Lyman_Page" title="Lyman Page">Lyman Page</a> and <a href="/wiki/David_Spergel" title="David Spergel">David Spergel</a> (2010)</li> <li><a href="/wiki/Enrico_Costa_(physicist)" title="Enrico Costa (physicist)">Enrico Costa</a> and <a href="/wiki/Gerald_J._Fishman" title="Gerald J. Fishman">Gerald Fishman</a> (2011)</li> <li><a href="/wiki/David_C._Jewitt" title="David C. Jewitt">David C. Jewitt</a> and <a href="/wiki/Jane_Luu" title="Jane Luu">Jane Luu</a> (2012)</li> <li><a href="/wiki/Steven_Balbus" title="Steven Balbus">Steven Balbus</a> and <a href="/wiki/John_F._Hawley" title="John F. Hawley">John F. Hawley</a> (2013)</li> <li><a href="/wiki/Daniel_Eisenstein" title="Daniel Eisenstein">Daniel Eisenstein</a>, <a href="/wiki/Shaun_Cole" title="Shaun Cole">Shaun Cole</a> and <a href="/wiki/John_A._Peacock" title="John A. Peacock">John A. Peacock</a> (2014)</li> <li><a href="/wiki/William_J._Borucki" title="William J. Borucki">William J. Borucki</a> (2015)</li> <li><a href="/wiki/Ronald_Drever" title="Ronald Drever">Ronald Drever</a>, <a href="/wiki/Kip_Thorne" title="Kip Thorne">Kip Thorne</a> and <a href="/wiki/Rainer_Weiss" title="Rainer Weiss">Rainer Weiss</a> (2016)</li> <li><a href="/wiki/Simon_White" title="Simon White">Simon White</a> (2017)</li> <li><a href="/wiki/Jean-Loup_Puget" title="Jean-Loup Puget">Jean-Loup Puget</a> (2018)</li> <li><a href="/wiki/Edward_C._Stone" title="Edward C. Stone">Edward C. Stone</a> (2019)</li> <li><a href="/wiki/Roger_Blandford" title="Roger Blandford">Roger Blandford</a> (2020)</li> <li><a href="/wiki/Victoria_Kaspi" title="Victoria Kaspi">Victoria Kaspi</a> and <a href="/wiki/Chryssa_Kouveliotou" title="Chryssa Kouveliotou">Chryssa Kouveliotou</a> (2021)</li> <li><a href="/wiki/Lennart_Lindegren" title="Lennart Lindegren">Lennart Lindegren</a> and <a href="/wiki/Michael_Perryman" title="Michael Perryman">Michael Perryman</a> (2022)</li> <li><a href="/wiki/Matthew_Bailes" title="Matthew Bailes">Matthew Bailes</a>, <a href="/wiki/Duncan_Lorimer" title="Duncan Lorimer">Duncan Lorimer</a> and <a href="/wiki/Maura_McLaughlin" title="Maura McLaughlin">Maura McLaughlin</a> (2023)</li> <li><a href="/wiki/Shrinivas_R._Kulkarni" class="mw-redirect" title="Shrinivas R. Kulkarni">Shrinivas R. Kulkarni</a> (2024)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">Life science<br />and medicine</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a>, <a href="/wiki/Herbert_Boyer" title="Herbert Boyer">Herbert Boyer</a>, <a href="/wiki/Yuet_Wai_Kan" title="Yuet Wai Kan">Yuet-Wai Kan</a> and <a href="/wiki/Richard_Doll" title="Richard Doll">Richard Doll</a> (2004)</li> <li><a href="/wiki/Michael_Berridge" title="Michael Berridge">Michael Berridge</a> (2005)</li> <li><a href="/wiki/Xiaodong_Wang_(biochemist)" title="Xiaodong Wang (biochemist)">Xiaodong Wang</a> (2006)</li> <li><a href="/wiki/Robert_Lefkowitz" title="Robert Lefkowitz">Robert Lefkowitz</a> (2007)</li> <li><a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Ian Wilmut</a>, <a href="/wiki/Keith_Campbell_(biologist)" title="Keith Campbell (biologist)">Keith H. S. Campbell</a> and <a class="mw-selflink selflink">Shinya Yamanaka</a> (2008)</li> <li><a href="/wiki/Douglas_L._Coleman" title="Douglas L. Coleman">Douglas Coleman</a> and <a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey Friedman</a> (2009)</li> <li><a href="/wiki/David_Julius" title="David Julius">David Julius</a> (2010)</li> <li><a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules Hoffmann</a>, <a href="/wiki/Ruslan_Medzhitov" title="Ruslan Medzhitov">Ruslan Medzhitov</a> and <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> (2011)</li> <li><a href="/wiki/Franz-Ulrich_Hartl" title="Franz-Ulrich Hartl">Franz-Ulrich Hartl</a> and <a href="/wiki/Arthur_L._Horwich" title="Arthur L. Horwich">Arthur L. Horwich</a> (2012)</li> <li><a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> and <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a> (2013)</li> <li><a href="/wiki/Kazutoshi_Mori" title="Kazutoshi Mori">Kazutoshi Mori</a> and <a href="/wiki/Peter_Walter" title="Peter Walter">Peter Walter</a> (2014)</li> <li><a href="/wiki/Bonnie_Bassler" title="Bonnie Bassler">Bonnie Bassler</a> and <a href="/wiki/Everett_Peter_Greenberg" title="Everett Peter Greenberg">Everett Peter Greenberg</a> (2015)</li> <li><a href="/wiki/Adrian_Bird" title="Adrian Bird">Adrian Bird</a> and <a href="/wiki/Huda_Zoghbi" title="Huda Zoghbi">Huda Zoghbi</a> (2016)</li> <li><a href="/wiki/Ian_R._Gibbons" title="Ian R. Gibbons">Ian R. Gibbons</a> and <a href="/wiki/Ronald_Vale" title="Ronald Vale">Ronald Vale</a> (2017)</li> <li><a href="/wiki/Mary-Claire_King" title="Mary-Claire King">Mary-Claire King</a> (2018)</li> <li><a href="/wiki/Maria_Jasin" title="Maria Jasin">Maria Jasin</a> (2019)</li> <li><a href="/wiki/Gero_Miesenb%C3%B6ck" title="Gero Miesenböck">Gero Miesenböck</a>, <a href="/wiki/Peter_Hegemann" title="Peter Hegemann">Peter Hegemann</a> and <a href="/wiki/Georg_Nagel" title="Georg Nagel">Georg Nagel</a> (2020)</li> <li><a href="/wiki/Scott_D._Emr" title="Scott D. Emr">Scott D. Emr</a> (2021)</li> <li><a href="/wiki/Paul_Negulescu" title="Paul Negulescu">Paul A. Negulescu</a> and <a href="/wiki/Michael_J._Welsh_(biologist)" title="Michael J. Welsh (biologist)">Michael J. Welsh</a> (2022)</li> <li><a href="/wiki/Patrick_Cramer" title="Patrick Cramer">Patrick Cramer</a> and <a href="/wiki/Eva_Nogales" title="Eva Nogales">Eva Nogales</a> (2023)</li> <li><a href="/wiki/Stuart_H._Orkin" class="mw-redirect" title="Stuart H. Orkin">Stuart H. Orkin</a> and <a href="/wiki/Swee_Lay_Thein" title="Swee Lay Thein">Swee Lay Thein</a> (2024)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">Mathematical<br />science</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Shiing-Shen_Chern" title="Shiing-Shen Chern">Shiing-Shen Chern</a> (2004)</li> <li><a href="/wiki/Andrew_Wiles" title="Andrew Wiles">Andrew Wiles</a> (2005)</li> <li><a href="/wiki/David_Mumford" title="David Mumford">David Mumford</a> and <a href="/wiki/Wu_Wenjun" title="Wu Wenjun">Wentsun Wu</a> (2006)</li> <li><a href="/wiki/Robert_Langlands" title="Robert Langlands">Robert Langlands</a> and <a href="/wiki/Richard_Taylor_(mathematician)" title="Richard Taylor (mathematician)">Richard Taylor</a> (2007)</li> <li><a href="/wiki/Vladimir_Arnold" title="Vladimir Arnold">Vladimir Arnold</a> and <a href="/wiki/Ludvig_Faddeev" title="Ludvig Faddeev">Ludwig Faddeev</a> (2008)</li> <li><a href="/wiki/Simon_Donaldson" title="Simon Donaldson">Simon Donaldson</a> and <a href="/wiki/Clifford_Taubes" title="Clifford Taubes">Clifford Taubes</a> (2009)</li> <li><a href="/wiki/Jean_Bourgain" title="Jean Bourgain">Jean Bourgain</a> (2010)</li> <li><a href="/wiki/Demetrios_Christodoulou" title="Demetrios Christodoulou">Demetrios Christodoulou</a> and <a href="/wiki/Richard_S._Hamilton" title="Richard S. Hamilton">Richard S. Hamilton</a> (2011)</li> <li><a href="/wiki/Maxim_Kontsevich" title="Maxim Kontsevich">Maxim Kontsevich</a> (2012)</li> <li><a href="/wiki/David_Donoho" title="David Donoho">David Donoho</a> (2013)</li> <li><a href="/wiki/George_Lusztig" title="George Lusztig">George Lusztig</a> (2014)</li> <li><a href="/wiki/Gerd_Faltings" title="Gerd Faltings">Gerd Faltings</a> and <a href="/wiki/Henryk_Iwaniec" title="Henryk Iwaniec">Henryk Iwaniec</a> (2015)</li> <li><a href="/wiki/Nigel_Hitchin" title="Nigel Hitchin">Nigel Hitchin</a> (2016)</li> <li><a href="/wiki/J%C3%A1nos_Koll%C3%A1r" title="János Kollár">János Kollár</a> and <a href="/wiki/Claire_Voisin" title="Claire Voisin">Claire Voisin</a> (2017)</li> <li><a href="/wiki/Luis_Caffarelli" title="Luis Caffarelli">Luis Caffarelli</a> (2018)</li> <li><a href="/wiki/Michel_Talagrand" title="Michel Talagrand">Michel Talagrand</a> (2019)</li> <li><a href="/wiki/Alexander_Beilinson" title="Alexander Beilinson">Alexander Beilinson</a> and <a href="/wiki/David_Kazhdan" title="David Kazhdan">David Kazhdan</a> (2020)</li> <li><a href="/wiki/Jean-Michel_Bismut" title="Jean-Michel Bismut">Jean-Michel Bismut</a> and <a href="/wiki/Jeff_Cheeger" title="Jeff Cheeger">Jeff Cheeger</a> (2021)</li> <li><a href="/wiki/Noga_Alon" title="Noga Alon">Noga Alon</a> and <a href="/wiki/Ehud_Hrushovski" title="Ehud Hrushovski">Ehud Hrushovski</a> (2022)</li> <li><a href="/wiki/Vladimir_Drinfeld" title="Vladimir Drinfeld">Vladimir Drinfeld</a> and <a href="/wiki/Shing-Tung_Yau" title="Shing-Tung Yau">Shing-Tung Yau</a> (2023)</li> <li><a href="/wiki/Peter_Sarnak" title="Peter Sarnak">Peter Sarnak</a> (2024)</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Laureates_of_the_Wolf_Prize_in_Medicine73" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background:#d3d3d3;"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Wolf_Prize_in_Medicine" title="Template:Wolf Prize in Medicine"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Wolf_Prize_in_Medicine" title="Template talk:Wolf Prize in Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Wolf_Prize_in_Medicine" title="Special:EditPage/Template:Wolf Prize in Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Wolf_Prize_in_Medicine73" style="font-size:114%;margin:0 4em">Laureates of the <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1970s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/Jon_van_Rood" title="Jon van Rood">Jon J. van Rood</a> (1978)</li> <li><a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Oleh_Hornykiewicz" title="Oleh Hornykiewicz">Oleh Hornykiewicz</a> (1979)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1980s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> / <a href="/wiki/Leo_Sachs" title="Leo Sachs">Leo Sachs</a> / <a href="/wiki/James_Learmonth_Gowans" title="James Learmonth Gowans">James L. Gowans</a> (1980)</li> <li><a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a> / <a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a> (1981)</li> <li><a href="/wiki/Jean-Pierre_Changeux" title="Jean-Pierre Changeux">Jean-Pierre Changeux</a> / <a href="/wiki/Solomon_H._Snyder" title="Solomon H. Snyder">Solomon H. Snyder</a> / <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> (1982)</li> <li><a href="/wiki/Donald_F._Steiner" title="Donald F. Steiner">Donald F. Steiner</a> (1984/5)</li> <li><a href="/wiki/Osamu_Hayaishi" title="Osamu Hayaishi">Osamu Hayaishi</a> (1986)</li> <li><a href="/wiki/Pedro_Cuatrecasas" title="Pedro Cuatrecasas">Pedro Cuatrecasas</a> / <a href="/wiki/Meir_Wilchek" title="Meir Wilchek">Meir Wilchek</a> (1987)</li> <li><a href="/wiki/Henri_G._Hers" title="Henri G. Hers">Henri G. Hers</a> / <a href="/wiki/Elizabeth_F._Neufeld" title="Elizabeth F. Neufeld">Elizabeth F. Neufeld</a> (1988)</li> <li><a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> (1989)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1990s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Maclyn_McCarty" title="Maclyn McCarty">Maclyn McCarty</a> (1990)</li> <li><a href="/wiki/Seymour_Benzer" title="Seymour Benzer">Seymour Benzer</a> (1991)</li> <li><a href="/wiki/Judah_Folkman" title="Judah Folkman">Judah Folkman</a> (1992)</li> <li><a href="/wiki/Michael_Berridge" title="Michael Berridge">Michael Berridge</a> / <a href="/wiki/Yasutomi_Nishizuka" title="Yasutomi Nishizuka">Yasutomi Nishizuka</a> (1994/5)</li> <li><a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a> (1995/6)</li> <li><a href="/wiki/Mary_F._Lyon" title="Mary F. Lyon">Mary F. Lyon</a> (1996/7)</li> <li><a href="/wiki/Michael_Sela" title="Michael Sela">Michael Sela</a> / <a href="/wiki/Ruth_Arnon" title="Ruth Arnon">Ruth Arnon</a> (1998)</li> <li><a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a> (1999)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2000s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Avram_Hershko" title="Avram Hershko">Avram Hershko</a> / <a href="/wiki/Alexander_Varshavsky" title="Alexander Varshavsky">Alexander Varshavsky</a> (2001)</li> <li><a href="/wiki/Ralph_L._Brinster" title="Ralph L. Brinster">Ralph L. Brinster</a> / <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a> (2002/3)</li> <li><a href="/wiki/Robert_Weinberg_(biologist)" title="Robert Weinberg (biologist)">Robert Weinberg</a> / <a href="/wiki/Roger_Y._Tsien" title="Roger Y. Tsien">Roger Y. Tsien</a> (2004)</li> <li><a href="/wiki/Alexander_Levitzki" title="Alexander Levitzki">Alexander Levitzki</a> / <a href="/wiki/Anthony_R._Hunter" title="Anthony R. Hunter">Anthony R. Hunter</a> / <a href="/wiki/Tony_Pawson_(biochemist)" title="Tony Pawson (biochemist)">Tony Pawson</a> (2005)</li> <li><a href="/wiki/Howard_Cedar" title="Howard Cedar">Howard Cedar</a> / <a href="/wiki/Aharon_Razin" title="Aharon Razin">Aharon Razin</a> (2008)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2010s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Axel_Ullrich" title="Axel Ullrich">Axel Ullrich</a> (2010)</li> <li><a class="mw-selflink selflink">Shinya Yamanaka</a> / <a href="/wiki/Rudolf_Jaenisch" title="Rudolf Jaenisch">Rudolf Jaenisch</a> (2011)</li> <li><a href="/wiki/Ronald_M._Evans" title="Ronald M. Evans">Ronald M. Evans</a> (2012)</li> <li><a href="/wiki/Nahum_Sonenberg" title="Nahum Sonenberg">Nahum Sonenberg</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a> / <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> (2014)</li> <li><a href="/wiki/John_Kappler" title="John Kappler">John Kappler</a> / <a href="/wiki/Philippa_Marrack" title="Philippa Marrack">Philippa Marrack</a> / <a href="/wiki/Jeffrey_V._Ravetch" title="Jeffrey V. Ravetch">Jeffrey V. Ravetch</a> (2015)</li> <li><a href="/wiki/Lewis_C._Cantley" title="Lewis C. Cantley">Lewis C. Cantley</a> / <a href="/wiki/C._Ronald_Kahn" title="C. Ronald Kahn">C. Ronald Kahn</a> (2016)</li> <li><a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> (2017)</li> <li><a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey M. Friedman</a> (2019)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2020s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a> / <a href="/wiki/Jennifer_Doudna" title="Jennifer Doudna">Jennifer Doudna</a> (2020)</li> <li><a href="/wiki/Joan_Steitz" class="mw-redirect" title="Joan Steitz">Joan Steitz</a> / <a href="/wiki/Lynne_Maquat" class="mw-redirect" title="Lynne Maquat">Lynne Elizabeth Maquat</a> / <a href="/wiki/Adrian_R._Krainer" class="mw-redirect" title="Adrian R. Krainer">Adrian Krainer</a> (2021)</li> <li><a href="/wiki/Daniel_J._Drucker" title="Daniel J. Drucker">Daniel J. Drucker</a> (2023)</li> <li><a href="/wiki/Botond_Roska" title="Botond Roska">Botond Roska</a> / <a href="/wiki/Jos%C3%A9-Alain_Sahel" title="José-Alain Sahel">José-Alain Sahel</a> (2024)</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r1239334494">@media screen{html.skin-theme-clientpref-night .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"></div><div role="navigation" class="navbox" aria-labelledby="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine170" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Template:Nobel Prize in Physiology or Medicine"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Nobel_Prize_in_Physiology_or_Medicine" title="Template talk:Nobel Prize in Physiology or Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Special:EditPage/Template:Nobel Prize in Physiology or Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine170" style="font-size:114%;margin:0 4em"><a href="/wiki/List_of_Nobel_laureates_in_Physiology_or_Medicine" title="List of Nobel laureates in Physiology or Medicine">Laureates</a> of the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Emil_von_Behring" title="Emil von Behring">Emil Behring</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ronald Ross</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Niels Finsen</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Ivan Pavlov</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Camillo Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Santiago Ramón y Cajal</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Alphonse Laveran</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/%C3%89lie_Metchnikoff" title="Élie Metchnikoff">Élie Metchnikoff</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Paul Ehrlich</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Emil Kocher</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Albrecht Kossel</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Allvar Gullstrand</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Alexis Carrel</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Charles Richet</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" class="mw-redirect" title="Róbert Bárány">Róbert Bárány</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1915</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1917</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1918</span></span></li> <li><span class="nowrap">1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Jules Bordet</a></span></li> <li><span class="nowrap">1920: <a href="/wiki/August_Krogh" title="August Krogh">August Krogh</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1921</span></span></li> <li><span class="nowrap">1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Archibald Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Otto Meyerhof</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Frederick Banting</a> / <a href="/wiki/John_Macleod_(physiologist)" title="John Macleod (physiologist)">John Macleod</a></span></li> <li><span class="nowrap">1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Willem Einthoven</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1925</span></span></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/80px-Nobel_Prize.png" decoding="async" width="80" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/120px-Nobel_Prize.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/160px-Nobel_Prize.png 2x" data-file-width="500" data-file-height="492" /></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Johannes Fibiger</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Charles Nicolle</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Christiaan Eijkman</a> / <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Frederick Gowland Hopkins</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Karl Landsteiner</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Otto Warburg</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Charles Scott Sherrington</a> / <a href="/wiki/Edgar_Adrian" title="Edgar Adrian">Edgar Adrian</a></span></li> <li><span class="nowrap">1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Morgan</a></span></li> <li><span class="nowrap">1934: <a href="/wiki/George_Whipple" title="George Whipple">George Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">George Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">William Murphy</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Hans Spemann</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Otto Loewi</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Albert Szent-Györgyi</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Corneille Heymans</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Gerhard Domagk</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Henrik Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Edward Doisy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Joseph Erlanger</a> / <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Herbert Gasser</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Alexander Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Ernst Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Howard Florey</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Hermann Muller</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">Carl Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">Gerty Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Bernardo Houssay</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Paul Müller</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Walter Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">António Egas Moniz</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Edward Kendall</a> / <a href="/wiki/Tadeusz_Reichstein" title="Tadeusz Reichstein">Tadeusz Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Philip Hench</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Max Theiler</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Selman Waksman</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hans_Krebs_(biochemist)" title="Hans Krebs (biochemist)">Hans Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Lipmann</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Thomas Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Frederick Robbins</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Hugo Theorell</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">André Cournand</a> / <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Werner Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Dickinson W. Richards</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Daniel Bovet</a></span></li> <li><span class="nowrap">1958: <a href="/wiki/George_Beadle" title="George Beadle">George Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Macfarlane_Burnet" title="Macfarlane Burnet">Frank Burnet</a> / <a href="/wiki/Peter_Medawar" title="Peter Medawar">Peter Medawar</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Georg von Békésy</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> / <a href="/wiki/James_Watson" title="James Watson">James Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Maurice Wilkins</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">John Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Alan Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Andrew Huxley</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Konrad Bloch</a> / <a href="/wiki/Feodor_Lynen" title="Feodor Lynen">Feodor Lynen</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">François Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" title="André Michel Lwoff">André Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Jacques Monod</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Charles B. Huggins</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Ragnar Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Haldan Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">George Wald</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Robert W. Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall Nirenberg</a></span></li> <li><span class="nowrap">1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Max Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Bernard Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf von Euler</a> / <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Julius Axelrod</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Earl Sutherland Jr.</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Gerald Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" class="mw-redirect" title="Rodney Robert Porter">Rodney Porter</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> / <a href="/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen">Nikolaas Tinbergen</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Albert Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Palade</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Renato Dulbecco</a> / <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Baruch Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Andrew Schally</a> / <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Yalow</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> / <a href="/wiki/Hamilton_O._Smith" title="Hamilton O. Smith">Hamilton O. Smith</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Allan_MacLeod_Cormack" title="Allan MacLeod Cormack">Allan Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Godfrey Hounsfield</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/David_H._Hubel" title="David H. Hubel">David H. Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Torsten Wiesel</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Sune Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" title="Bengt I. Samuelsson">Bengt I. Samuelsson</a> / <a href="/wiki/John_Vane" title="John Vane">John Vane</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Georges Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Michael Brown</a> / <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Joseph L. Goldstein</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Stanley_Cohen_(biochemist)" title="Stanley Cohen (biochemist)">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> / <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Gertrude B. Elion</a> / <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">George H. Hitchings</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">J. Michael Bishop</a> / <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Harold E. Varmus</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Joseph_Murray" class="mw-redirect" title="Joseph Murray">Joseph Murray</a> / <a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">E. Donnall Thomas</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Edmond Fischer</a> / <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">Edwin G. Krebs</a></span></li> <li><span class="nowrap">1993: <a href="/wiki/Richard_J._Roberts" title="Richard J. Roberts">Richard J. Roberts</a> / <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Sharp</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Alfred G. Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Martin Rodbell</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Christiane Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Eric F. Wieschaus</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Peter_C._Doherty" class="mw-redirect" title="Peter C. Doherty">Peter C. Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Rolf M. Zinkernagel</a></span></li> <li><span class="nowrap">1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Robert F. Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Louis Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Paul Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Leland H. Hartwell</a> / <a href="/wiki/Tim_Hunt" title="Tim Hunt">Tim Hunt</a> / <a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> / <a href="/wiki/John_Sulston" title="John Sulston">John E. Sulston</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Barry Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Robin Warren</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Andrew Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Craig Mello</a></span></li> <li><span class="nowrap">2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> / <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> / <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Carol W. Greider</a> / <a href="/wiki/Jack_W._Szostak" title="Jack W. Szostak">Jack W. Szostak</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Robert_Edwards_(physiologist)" title="Robert Edwards (physiologist)">Robert G. Edwards</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> / <a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules A. Hoffmann</a> / <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Ralph M. Steinman</a> (<i>posthumously</i>)</span></li> <li><span class="nowrap">2012: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a class="mw-selflink selflink">Shinya Yamanaka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/James_Rothman" title="James Rothman">James Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Randy Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Thomas C. Südhof</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/John_O%27Keefe_(neuroscientist)" title="John O'Keefe (neuroscientist)">John O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">May-Britt Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">Edvard Moser</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/William_C._Campbell_(scientist)" title="William C. Campbell (scientist)">William C. Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Tu Youyou</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Gregg L. Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">William Kaelin Jr.</a></span></li> <li><span class="nowrap"><a href="/wiki/2020_Nobel_Prize_in_Physiology_or_Medicine" title="2020 Nobel Prize in Physiology or Medicine">2020</a>: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Harvey J. Alter</a> / <a href="/wiki/Michael_Houghton_(virologist)" class="mw-redirect" title="Michael Houghton (virologist)">Michael Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Charles M. Rice</a></span></li> <li><span class="nowrap"><a href="/wiki/2021_Nobel_Prize_in_Physiology_or_Medicine" title="2021 Nobel Prize in Physiology or Medicine">2021</a>: <a href="/wiki/David_Julius" title="David Julius">David Julius</a> / <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Ardem Patapoutian</a></span></li> <li><span class="nowrap"><a href="/wiki/2022_Nobel_Prize_in_Physiology_or_Medicine" title="2022 Nobel Prize in Physiology or Medicine">2022</a>: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a></span></li> <li><span class="nowrap"><a href="/wiki/2023_Nobel_Prize_in_Physiology_or_Medicine" title="2023 Nobel Prize in Physiology or Medicine">2023</a>: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style></div><div role="navigation" class="navbox" aria-labelledby="2012_Nobel_Prize_laureates787" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:2012_Nobel_Prize_winners" title="Template:2012 Nobel Prize winners"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:2012_Nobel_Prize_winners" title="Template talk:2012 Nobel Prize winners"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:2012_Nobel_Prize_winners" title="Special:EditPage/Template:2012 Nobel Prize winners"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="2012_Nobel_Prize_laureates787" style="font-size:114%;margin:0 4em">2012 <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Robert_Lefkowitz" title="Robert Lefkowitz">Robert Lefkowitz</a> (United States)</li> <li><a href="/wiki/Brian_Kobilka" title="Brian Kobilka">Brian Kobilka</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a> (<a href="/wiki/2012_Nobel_Prize_in_Literature" title="2012 Nobel Prize in Literature">2012</a>)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mo_Yan" title="Mo Yan">Mo Yan</a> (China)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Peace_Prize" title="Nobel Peace Prize">Peace</a> (<a href="/wiki/2012_Nobel_Peace_Prize" title="2012 Nobel Peace Prize">2012</a>)</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/European_Union" title="European Union">European Union</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Serge_Haroche" title="Serge Haroche">Serge Haroche</a> (France)</li> <li><a href="/wiki/David_J._Wineland" title="David J. Wineland">David J. Wineland</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> (United Kingdom)</li> <li><a class="mw-selflink selflink">Shinya Yamanaka</a> (Japan)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economic Sciences</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Alvin_E._Roth" title="Alvin E. Roth">Alvin E. Roth</a> (United States)</li> <li><a href="/wiki/Lloyd_Shapley" title="Lloyd Shapley">Lloyd Shapley</a> (United States)</li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <dl><dt><span class="nobold"><a href="/wiki/List_of_Nobel_laureates" title="List of Nobel laureates">Nobel Prize recipients</a></span></dt> <dd><a href="/wiki/2007_Nobel_Prizes" title="2007 Nobel Prizes">2007</a></dd> <dd><a href="/wiki/2008_Nobel_Prizes" title="2008 Nobel Prizes">2008</a></dd> <dd><a href="/wiki/2009_Nobel_Prizes" title="2009 Nobel Prizes">2009</a></dd> <dd><a href="/wiki/2010_Nobel_Prizes" title="2010 Nobel Prizes">2010</a></dd> <dd><a href="/wiki/2011_Nobel_Prizes" title="2011 Nobel Prizes">2011</a></dd> <dd><b>2012 </b></dd> <dd><a href="/wiki/2013_Nobel_Prizes" title="2013 Nobel Prizes">2013</a></dd> <dd><a href="/wiki/2014_Nobel_Prizes" title="2014 Nobel Prizes">2014</a></dd> <dd><a href="/wiki/2015_Nobel_Prizes" title="2015 Nobel Prizes">2015</a></dd> <dd><a href="/wiki/2016_Nobel_Prizes" title="2016 Nobel Prizes">2016</a></dd> <dd><a href="/wiki/2017_Nobel_Prizes" title="2017 Nobel Prizes">2017</a></dd></dl> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="23x15px&#124;border_&#124;alt=&#124;link=_Japanese_Nobel_laureates361" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Japanese_Nobel_laureates" title="Template:Japanese Nobel laureates"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Japanese_Nobel_laureates" title="Template talk:Japanese Nobel laureates"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Japanese_Nobel_laureates" title="Special:EditPage/Template:Japanese Nobel laureates"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="23x15px&#124;border_&#124;alt=&#124;link=_Japanese_Nobel_laureates361" style="font-size:114%;margin:0 4em"><span class="flagicon"><span class="mw-image-border" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/9e/Flag_of_Japan.svg/23px-Flag_of_Japan.svg.png" decoding="async" width="23" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/9e/Flag_of_Japan.svg/35px-Flag_of_Japan.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/9e/Flag_of_Japan.svg/45px-Flag_of_Japan.svg.png 2x" data-file-width="900" data-file-height="600" /></span></span></span> <a href="/wiki/List_of_Japanese_Nobel_laureates_and_nominees" title="List of Japanese Nobel laureates and nominees">Japanese Nobel laureates</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Kenichi_Fukui" title="Kenichi Fukui">Kenichi Fukui</a> (1981)</li> <li><a href="/wiki/Hideki_Shirakawa" title="Hideki Shirakawa">Hideki Shirakawa</a> (2000)</li> <li><a href="/wiki/Ry%C5%8Dji_Noyori" title="Ryōji Noyori">Ryōji Noyori</a> (2001)</li> <li><a href="/wiki/Koichi_Tanaka" title="Koichi Tanaka">Koichi Tanaka</a> (2002)</li> <li><a href="/wiki/Osamu_Shimomura" title="Osamu Shimomura">Osamu Shimomura</a> (2008)</li> <li><a href="/wiki/Ei-ichi_Negishi" title="Ei-ichi Negishi">Ei-ichi Negishi</a> / <a href="/wiki/Akira_Suzuki" title="Akira Suzuki">Akira Suzuki</a> (2010)</li> <li><a href="/wiki/Akira_Yoshino" title="Akira Yoshino">Akira Yoshino</a> (2019)</li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/80px-Nobel_Prize.png" decoding="async" width="80" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/120px-Nobel_Prize.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/160px-Nobel_Prize.png 2x" data-file-width="500" data-file-height="492" /></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Yasunari_Kawabata" title="Yasunari Kawabata">Yasunari Kawabata</a> (1968)</li> <li><a href="/wiki/Kenzabur%C5%8D_%C5%8Ce" title="Kenzaburō Ōe">Kenzaburō Ōe</a> (1994)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Peace_Prize" title="Nobel Peace Prize">Peace</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Eisaku_Sat%C5%8D" title="Eisaku Satō">Eisaku Satō</a> (1974)</li> <li><a href="/wiki/Nihon_Hidankyo" title="Nihon Hidankyo">Nihon Hidankyo</a> (2024)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hideki_Yukawa" title="Hideki Yukawa">Hideki Yukawa</a> (1949)</li> <li><a href="/wiki/Shin%27ichir%C5%8D_Tomonaga" title="Shin'ichirō Tomonaga">Shin'ichirō Tomonaga</a> (1965)</li> <li><a href="/wiki/Leo_Esaki" title="Leo Esaki">Leo Esaki</a> (1973)</li> <li><a href="/wiki/Masatoshi_Koshiba" title="Masatoshi Koshiba">Masatoshi Koshiba</a> (2002)</li> <li><a href="/wiki/Makoto_Kobayashi_(physicist)" class="mw-redirect" title="Makoto Kobayashi (physicist)">Makoto Kobayashi</a> / <a href="/wiki/Toshihide_Maskawa" title="Toshihide Maskawa">Toshihide Maskawa</a> (2008)</li> <li><a href="/wiki/Isamu_Akasaki" title="Isamu Akasaki">Isamu Akasaki</a> / <a href="/wiki/Hiroshi_Amano" title="Hiroshi Amano">Hiroshi Amano</a> (2014)</li> <li><a href="/wiki/Takaaki_Kajita" title="Takaaki Kajita">Takaaki Kajita</a> (2015)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a> (1987)</li> <li><a class="mw-selflink selflink">Shinya Yamanaka</a> (2012)</li> <li><a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> (2015)</li> <li><a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a> (2016)</li> <li><a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a> (2018)</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Breakthrough_Prize_laureates203" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Breakthrough_Prize_laureates" title="Template:Breakthrough Prize laureates"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Breakthrough_Prize_laureates" title="Template talk:Breakthrough Prize laureates"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Breakthrough_Prize_laureates" title="Special:EditPage/Template:Breakthrough Prize laureates"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Breakthrough_Prize_laureates203" style="font-size:114%;margin:0 4em"><a href="/wiki/Breakthrough_Prize" title="Breakthrough Prize">Breakthrough Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Breakthrough_Prize_in_Mathematics" title="Breakthrough Prize in Mathematics">Mathematics</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Simon_Donaldson" title="Simon Donaldson">Simon Donaldson</a>, <a href="/wiki/Maxim_Kontsevich" title="Maxim Kontsevich">Maxim Kontsevich</a>, <a href="/wiki/Jacob_Lurie" title="Jacob Lurie">Jacob Lurie</a>, <a href="/wiki/Terence_Tao" title="Terence Tao">Terence Tao</a> and <a href="/wiki/Richard_Taylor_(mathematician)" title="Richard Taylor (mathematician)">Richard Taylor</a> (2015)</li> <li><a href="/wiki/Ian_Agol" title="Ian Agol">Ian Agol</a> (2016)</li> <li><a href="/wiki/Jean_Bourgain" title="Jean Bourgain">Jean Bourgain</a> (2017)</li> <li><a href="/wiki/Christopher_Hacon" title="Christopher Hacon">Christopher Hacon</a>, <a href="/wiki/James_McKernan" title="James McKernan">James McKernan</a> (2018)</li> <li><a href="/wiki/Vincent_Lafforgue" title="Vincent Lafforgue">Vincent Lafforgue</a> (2019)</li> <li><a href="/wiki/Alex_Eskin" title="Alex Eskin">Alex Eskin</a> (2020)</li> <li><a href="/wiki/Martin_Hairer" title="Martin Hairer">Martin Hairer</a> (2021)</li> <li><a href="/wiki/Takuro_Mochizuki" class="mw-redirect" title="Takuro Mochizuki">Takuro Mochizuki</a> (2022)</li> <li><a href="/wiki/Daniel_A._Spielman" class="mw-redirect" title="Daniel A. Spielman">Daniel A. Spielman</a> (2023)</li> <li><a href="/wiki/Simon_Brendle" title="Simon Brendle">Simon Brendle</a> (2024)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Breakthrough_Prize_in_Fundamental_Physics" title="Breakthrough Prize in Fundamental Physics">Fundamental<br />physics</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nima_Arkani-Hamed" title="Nima Arkani-Hamed">Nima Arkani-Hamed</a>, <a href="/wiki/Alan_Guth" title="Alan Guth">Alan Guth</a>, <a href="/wiki/Alexei_Kitaev" title="Alexei Kitaev">Alexei Kitaev</a>, <a href="/wiki/Maxim_Kontsevich" title="Maxim Kontsevich">Maxim Kontsevich</a>, <a href="/wiki/Andrei_Linde" title="Andrei Linde">Andrei Linde</a>, <a href="/wiki/Juan_Mart%C3%ADn_Maldacena" class="mw-redirect" title="Juan Martín Maldacena">Juan Maldacena</a>, <a href="/wiki/Nathan_Seiberg" title="Nathan Seiberg">Nathan Seiberg</a>, <a href="/wiki/Ashoke_Sen" title="Ashoke Sen">Ashoke Sen</a>, <a href="/wiki/Edward_Witten" title="Edward Witten">Edward Witten</a> (2012)</li> <li><i>Special</i>: <a href="/wiki/Stephen_Hawking" title="Stephen Hawking">Stephen Hawking</a>, <a href="/wiki/Peter_Jenni" title="Peter Jenni">Peter Jenni</a>, <a href="/wiki/Fabiola_Gianotti" title="Fabiola Gianotti">Fabiola Gianotti</a> (ATLAS), <a href="/wiki/Michel_Della_Negra" title="Michel Della Negra">Michel Della Negra</a>, <a href="/wiki/Tejinder_Virdee" title="Tejinder Virdee">Tejinder Virdee</a>, <a href="/wiki/Guido_Tonelli" title="Guido Tonelli">Guido Tonelli</a>, <a href="/wiki/Joseph_Incandela" title="Joseph Incandela">Joseph Incandela</a> (CMS) and <a href="/wiki/Lyn_Evans" title="Lyn Evans">Lyn Evans</a> (LHC) (2013)</li> <li><a href="/wiki/Alexander_Markovich_Polyakov" title="Alexander Markovich Polyakov">Alexander Polyakov</a> (2013)</li> <li><a href="/wiki/Michael_Green_(physicist)" title="Michael Green (physicist)">Michael Green</a> and <a href="/wiki/John_Henry_Schwarz" title="John Henry Schwarz">John Henry Schwarz</a> (2014)</li> <li><a href="/wiki/Saul_Perlmutter" title="Saul Perlmutter">Saul Perlmutter</a> and members of the <a href="/wiki/Supernova_Cosmology_Project" title="Supernova Cosmology Project">Supernova Cosmology Project</a>; <a href="/wiki/Brian_Schmidt" title="Brian Schmidt">Brian Schmidt</a>, <a href="/wiki/Adam_Riess" title="Adam Riess">Adam Riess</a> and members of the <a href="/wiki/High-Z_Supernova_Search_Team" title="High-Z Supernova Search Team">High-Z Supernova Team</a> (2015)</li> <li><i>Special</i>: <a href="/wiki/Ronald_Drever" title="Ronald Drever">Ronald Drever</a>, <a href="/wiki/Kip_Thorne" title="Kip Thorne">Kip Thorne</a>, <a href="/wiki/Rainer_Weiss" title="Rainer Weiss">Rainer Weiss</a> and contributors to <a href="/wiki/LIGO" title="LIGO">LIGO</a> project (2016)</li> <li><a href="/wiki/Wang_Yifang" title="Wang Yifang">Yifang Wang</a>, <a href="/wiki/Kam-Biu_Luk" title="Kam-Biu Luk">Kam-Biu Luk</a> and the <a href="/wiki/Daya_Bay_Reactor_Neutrino_Experiment" title="Daya Bay Reactor Neutrino Experiment">Daya Bay team</a>, <a href="/wiki/Atsuto_Suzuki" title="Atsuto Suzuki">Atsuto Suzuki</a> and the <a href="/wiki/Kamioka_Liquid_Scintillator_Antineutrino_Detector" title="Kamioka Liquid Scintillator Antineutrino Detector">KamLAND</a> team, <a href="/wiki/K%C5%8Dichir%C5%8D_Nishikawa" title="Kōichirō Nishikawa">Kōichirō Nishikawa</a> and the <a href="/wiki/K2K_experiment" title="K2K experiment">K2K</a> / <a href="/wiki/T2K_experiment" title="T2K experiment">T2K</a> team, <a href="/wiki/Arthur_B._McDonald" title="Arthur B. McDonald">Arthur B. McDonald</a> and the <a href="/wiki/Sudbury_Neutrino_Observatory" title="Sudbury Neutrino Observatory">Sudbury Neutrino Observatory</a> team, <a href="/wiki/Takaaki_Kajita" title="Takaaki Kajita">Takaaki Kajita</a> and <a href="/wiki/Y%C5%8Dichir%C5%8D_Suzuki" title="Yōichirō Suzuki">Yōichirō Suzuki</a> and the <a href="/wiki/Super-Kamiokande" title="Super-Kamiokande">Super-Kamiokande</a> team (2016)</li> <li><a href="/wiki/Joseph_Polchinski" title="Joseph Polchinski">Joseph Polchinski</a>, <a href="/wiki/Andrew_Strominger" title="Andrew Strominger">Andrew Strominger</a>, <a href="/wiki/Cumrun_Vafa" title="Cumrun Vafa">Cumrun Vafa</a> (2017)</li> <li><a href="/wiki/Charles_L._Bennett" title="Charles L. Bennett">Charles L. Bennett</a>, <a href="/wiki/Gary_Hinshaw" title="Gary Hinshaw">Gary Hinshaw</a>, <a href="/wiki/Norman_Jarosik" title="Norman Jarosik">Norman Jarosik</a>, <a href="/wiki/Lyman_Page" title="Lyman Page">Lyman Page Jr.</a>, <a href="/wiki/David_Spergel" title="David Spergel">David Spergel</a> (2018)</li> <li><i>Special</i>: <a href="/wiki/Jocelyn_Bell_Burnell" title="Jocelyn Bell Burnell">Jocelyn Bell Burnell</a> (2018)</li> <li><a href="/wiki/Charles_L._Kane" title="Charles L. Kane">Charles Kane</a> and <a href="/wiki/Eugene_J._Mele" title="Eugene J. Mele">Eugene Mele</a> (2019)</li> <li><i>Special</i>: <a href="/wiki/Sergio_Ferrara" title="Sergio Ferrara">Sergio Ferrara</a>, <a href="/wiki/Daniel_Z._Freedman" title="Daniel Z. Freedman">Daniel Z. Freedman</a>, <a href="/wiki/Peter_van_Nieuwenhuizen" title="Peter van Nieuwenhuizen">Peter van Nieuwenhuizen</a> (2019)</li> <li><a href="/wiki/The_Event_Horizon_Telescope_Collaboration" class="mw-redirect" title="The Event Horizon Telescope Collaboration">The Event Horizon Telescope Collaboration</a> (2020)</li> <li><a href="/wiki/Eric_Adelberger" class="mw-redirect" title="Eric Adelberger">Eric Adelberger</a>, <a href="/wiki/Jens_H._Gundlach" title="Jens H. Gundlach">Jens H. Gundlach</a> and <a href="/wiki/Blayne_Heckel" title="Blayne Heckel">Blayne Heckel</a> (2021)</li> <li><i>Special</i>: <a href="/wiki/Steven_Weinberg" title="Steven Weinberg">Steven Weinberg</a> (2021)</li> <li><a href="/wiki/Hidetoshi_Katori" title="Hidetoshi Katori">Hidetoshi Katori</a> and <a href="/wiki/Jun_Ye" title="Jun Ye">Jun Ye</a> (2022)</li> <li><a href="/wiki/Charles_H._Bennett_(physicist)" title="Charles H. Bennett (physicist)">Charles H. Bennett</a>, <a href="/wiki/Gilles_Brassard" title="Gilles Brassard">Gilles Brassard</a>, <a href="/wiki/David_Deutsch" title="David Deutsch">David Deutsch</a>, <a href="/wiki/Peter_W._Shor" class="mw-redirect" title="Peter W. Shor">Peter W. Shor</a> (2023)</li> <li><a href="/wiki/John_Cardy" title="John Cardy">John Cardy</a> and <a href="/wiki/Alexander_Zamolodchikov" title="Alexander Zamolodchikov">Alexander Zamolodchikov</a> (2024)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Breakthrough_Prize_in_Life_Sciences" title="Breakthrough Prize in Life Sciences">Life sciences</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cornelia_Bargmann" title="Cornelia Bargmann">Cornelia Bargmann</a>, <a href="/wiki/David_Botstein" title="David Botstein">David Botstein</a>, <a href="/wiki/Lewis_C._Cantley" title="Lewis C. Cantley">Lewis C. Cantley</a>, <a href="/wiki/Hans_Clevers" title="Hans Clevers">Hans Clevers</a>, <a href="/wiki/Titia_de_Lange" title="Titia de Lange">Titia de Lange</a>, <a href="/wiki/Napoleone_Ferrara" title="Napoleone Ferrara">Napoleone Ferrara</a>, <a href="/wiki/Eric_Lander" title="Eric Lander">Eric Lander</a>, <a href="/wiki/Charles_Sawyers" title="Charles Sawyers">Charles Sawyers</a>, <a href="/wiki/Robert_Weinberg_(biologist)" title="Robert Weinberg (biologist)">Robert Weinberg</a>, <a class="mw-selflink selflink">Shinya Yamanaka</a> and <a href="/wiki/Bert_Vogelstein" title="Bert Vogelstein">Bert Vogelstein</a> (2013)</li> <li><a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a>, <a href="/wiki/Mahlon_DeLong" title="Mahlon DeLong">Mahlon DeLong</a>, <a href="/wiki/Michael_N._Hall" title="Michael N. Hall">Michael N. Hall</a>, <a href="/wiki/Robert_S._Langer" title="Robert S. Langer">Robert S. Langer</a>, <a href="/wiki/Richard_P._Lifton" title="Richard P. Lifton">Richard P. Lifton</a> and <a href="/wiki/Alexander_Varshavsky" title="Alexander Varshavsky">Alexander Varshavsky</a> (2014)</li> <li><a href="/wiki/Alim_Louis_Benabid" title="Alim Louis Benabid">Alim Louis Benabid</a>, <a href="/wiki/Charles_David_Allis" title="Charles David Allis">Charles David Allis</a>, <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a>, <a href="/wiki/Jennifer_Doudna" title="Jennifer Doudna">Jennifer Doudna</a> and <a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a> (2015)</li> <li><a href="/wiki/Edward_Boyden" title="Edward Boyden">Edward Boyden</a>, <a href="/wiki/Karl_Deisseroth" title="Karl Deisseroth">Karl Deisseroth</a>, <a href="/wiki/John_Hardy_(geneticist)" title="John Hardy (geneticist)">John Hardy</a>, <a href="/wiki/Helen_Hobbs" title="Helen Hobbs">Helen Hobbs</a> and <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a> (2016)</li> <li><a href="/wiki/Stephen_Elledge" title="Stephen Elledge">Stephen J. Elledge</a>, <a href="/wiki/Harry_F._Noller" title="Harry F. Noller">Harry F. Noller</a>, <a href="/wiki/Roel_Nusse" title="Roel Nusse">Roeland Nusse</a>, <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a>, <a href="/wiki/Huda_Zoghbi" title="Huda Zoghbi">Huda Zoghbi</a> (2017)</li> <li><a href="/wiki/Joanne_Chory" title="Joanne Chory">Joanne Chory</a>, <a href="/wiki/Peter_Walter" title="Peter Walter">Peter Walter</a>, <a href="/wiki/Kazutoshi_Mori" title="Kazutoshi Mori">Kazutoshi Mori</a>, <a href="/wiki/Kim_Nasmyth" title="Kim Nasmyth">Kim Nasmyth</a>, <a href="/wiki/Don_W._Cleveland" title="Don W. Cleveland">Don W. Cleveland</a> (2018)</li> <li><a href="/wiki/C._Frank_Bennett" title="C. Frank Bennett">C. Frank Bennett</a> and <a href="/wiki/Adrian_R._Krainer" class="mw-redirect" title="Adrian R. Krainer">Adrian R. Krainer</a>, <a href="/wiki/Angelika_Amon" title="Angelika Amon">Angelika Amon</a>, <a href="/wiki/Xiaowei_Zhuang" title="Xiaowei Zhuang">Xiaowei Zhuang</a>, <a href="/wiki/Zhijian_Chen" title="Zhijian Chen">Zhijian Chen</a> (2019)</li> <li><a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey M. Friedman</a>, <a href="/wiki/Franz-Ulrich_Hartl" title="Franz-Ulrich Hartl">Franz-Ulrich Hartl</a>, <a href="/wiki/Arthur_L._Horwich" title="Arthur L. Horwich">Arthur L. Horwich</a>, <a href="/wiki/David_Julius" title="David Julius">David Julius</a>, <a href="/wiki/Virginia_Man-Yee_Lee" title="Virginia Man-Yee Lee">Virginia Man-Yee Lee</a> (2020)</li> <li><a href="/wiki/David_Baker_(biochemist)" title="David Baker (biochemist)">David Baker</a>, <a href="/wiki/Catherine_Dulac" title="Catherine Dulac">Catherine Dulac</a>, <a href="/wiki/Dennis_Lo" title="Dennis Lo">Dennis Lo</a>, <a href="/w/index.php?title=Richard_J._Youle&action=edit&redlink=1" class="new" title="Richard J. Youle (page does not exist)">Richard J. Youle</a><span class="noprint" style="font-size:85%; font-style: normal;"> [<a href="https://de.wikipedia.org/wiki/Richard_J._Youle" class="extiw" title="de:Richard J. Youle">de</a>]</span> (2021)</li> <li><a href="/wiki/Jeffery_W._Kelly" title="Jeffery W. Kelly">Jeffery W. Kelly</a>, <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a>, <a href="/wiki/Shankar_Balasubramanian" title="Shankar Balasubramanian">Shankar Balasubramanian</a>, <a href="/wiki/David_Klenerman" title="David Klenerman">David Klenerman</a> and <a href="/wiki/Pascal_Mayer" title="Pascal Mayer">Pascal Mayer</a> (2022)</li> <li><a href="/wiki/Clifford_P._Brangwynne" title="Clifford P. Brangwynne">Clifford P. Brangwynne</a>, <a href="/wiki/Anthony_A._Hyman" title="Anthony A. Hyman">Anthony A. Hyman</a>, <a href="/wiki/Demis_Hassabis" title="Demis Hassabis">Demis Hassabis</a>, <a href="/wiki/John_M._Jumper" title="John M. Jumper">John Jumper</a>, <a href="/wiki/Emmanuel_Mignot" title="Emmanuel Mignot">Emmanuel Mignot</a>, <a href="/wiki/Masashi_Yanagisawa" title="Masashi Yanagisawa">Masashi Yanagisawa</a> (2023)</li> <li><a href="/wiki/Carl_June" class="mw-redirect" title="Carl June">Carl June</a>, <a href="/wiki/Michel_Sadelain" title="Michel Sadelain">Michel Sadelain</a>, <a href="/wiki/Sabine_Hadida" title="Sabine Hadida">Sabine Hadida</a>, <a href="/wiki/Paul_Negulescu" title="Paul Negulescu">Paul Negulescu</a>, <a href="/w/index.php?title=Fredrick_Van_Goor&action=edit&redlink=1" class="new" title="Fredrick Van Goor (page does not exist)">Fredrick Van Goor</a>, <a href="/w/index.php?title=Thomas_Gasser&action=edit&redlink=1" class="new" title="Thomas Gasser (page does not exist)">Thomas Gasser</a>, <a href="/wiki/Ellen_Sidransky" title="Ellen Sidransky">Ellen Sidransky</a> and <a href="/wiki/Andrew_Singleton" title="Andrew Singleton">Andrew Singleton</a> (2024)</li></ul> 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