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class="firstHeading mw-first-heading"><span class="mw-page-title-main">Vascular bundle</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 28 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-28" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">28 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Vaatbundel" title="Vaatbundel – Afrikaans" lang="af" hreflang="af" data-title="Vaatbundel" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AD%D8%B2%D9%85%D8%A9_%D9%88%D8%B9%D8%A7%D8%A6%D9%8A%D8%A9" title="حزمة وعائية – Arabic" lang="ar" hreflang="ar" data-title="حزمة وعائية" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AD%E0%A6%BE%E0%A6%B8%E0%A7%8D%E0%A6%95%E0%A7%81%E0%A6%B2%E0%A6%BE%E0%A6%B0_%E0%A6%AC%E0%A6%BE%E0%A6%A8%E0%A7%8D%E0%A6%A1%E0%A6%B2" title="ভাস্কুলার বান্ডল – Bangla" lang="bn" hreflang="bn" data-title="ভাস্কুলার বান্ডল" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%9F%D1%80%D0%B0%D0%B2%D0%BE%D0%B4%D0%B7%D1%8F%D1%87%D1%8B_%D0%BF%D1%83%D1%87%D0%BE%D0%BA" title="Праводзячы пучок – Belarusian" lang="be" hreflang="be" data-title="Праводзячы пучок" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Provodno_tkivo" title="Provodno tkivo – Bosnian" lang="bs" hreflang="bs" data-title="Provodno tkivo" data-language-autonym="Bosanski" data-language-local-name="Bosnian" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/C%C3%A9vn%C3%AD_svazek" title="Cévní svazek – Czech" lang="cs" hreflang="cs" data-title="Cévní svazek" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Leitb%C3%BCndel" title="Leitbündel – German" lang="de" hreflang="de" data-title="Leitbündel" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Haz_vascular" title="Haz vascular – Spanish" lang="es" hreflang="es" data-title="Haz vascular" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Vaskulaj_faskoj" title="Vaskulaj faskoj – Esperanto" lang="eo" hreflang="eo" data-title="Vaskulaj faskoj" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%AF%D8%B3%D8%AA%D9%87%E2%80%8C%D9%87%D8%A7%DB%8C_%D8%A2%D9%88%D9%86%D8%AF%DB%8C" title="دسته‌های آوندی – Persian" lang="fa" hreflang="fa" data-title="دسته‌های آوندی" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Faisceau_cribro-vasculaire" title="Faisceau cribro-vasculaire – French" lang="fr" hreflang="fr" data-title="Faisceau cribro-vasculaire" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EA%B4%80%EB%8B%A4%EB%B0%9C" title="관다발 – Korean" lang="ko" hreflang="ko" data-title="관다발" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D4%B1%D5%B6%D6%81%D5%AF%D5%A1%D6%81%D5%B6%D5%B8%D5%B2_%D5%AD%D6%80%D5%B1%D5%A5%D6%80" title="Անցկացնող խրձեր – Armenian" lang="hy" hreflang="hy" data-title="Անցկացնող խրձեր" data-language-autonym="Հայերեն" data-language-local-name="Armenian" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Jaringan_pengangkut" title="Jaringan pengangkut – Indonesian" lang="id" hreflang="id" data-title="Jaringan pengangkut" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Fasci_cribro-vascolari" title="Fasci cribro-vascolari – Italian" lang="it" hreflang="it" data-title="Fasci cribro-vascolari" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Ed%C3%A9nynyal%C3%A1b" title="Edénynyaláb – Hungarian" lang="hu" hreflang="hu" data-title="Edénynyaláb" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%B8%E0%B4%82%E0%B4%B5%E0%B4%B9%E0%B4%A8%E0%B4%B5%E0%B5%8D%E0%B4%AF%E0%B5%82%E0%B4%B9%E0%B4%82" title="സംവഹനവ്യൂഹം – Malayalam" lang="ml" hreflang="ml" data-title="സംവഹനവ്യൂഹം" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Vaatbundel" title="Vaatbundel – Dutch" lang="nl" hreflang="nl" data-title="Vaatbundel" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E7%B6%AD%E7%AE%A1%E6%9D%9F" title="維管束 – Japanese" lang="ja" hreflang="ja" data-title="維管束" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Yuqorio%27tkazuvchi_nur" title="Yuqorio'tkazuvchi nur – Uzbek" lang="uz" hreflang="uz" data-title="Yuqorio'tkazuvchi nur" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Wi%C4%85zki_przewodz%C4%85ce" title="Wiązki przewodzące – Polish" lang="pl" hreflang="pl" data-title="Wiązki przewodzące" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Feixe_vascular" title="Feixe vascular – Portuguese" lang="pt" hreflang="pt" data-title="Feixe vascular" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Fascicul_libero-lemnos" title="Fascicul libero-lemnos – Romanian" lang="ro" hreflang="ro" data-title="Fascicul libero-lemnos" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru badge-Q17559452 badge-recommendedarticle mw-list-item" title="recommended article"><a href="https://ru.wikipedia.org/wiki/%D0%9F%D1%80%D0%BE%D0%B2%D0%BE%D0%B4%D1%8F%D1%89%D0%B8%D0%B9_%D0%BF%D1%83%D1%87%D0%BE%D0%BA" title="Проводящий пучок – Russian" lang="ru" hreflang="ru" data-title="Проводящий пучок" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Rastlinska_%C5%BEila" title="Rastlinska žila – Slovenian" lang="sl" hreflang="sl" data-title="Rastlinska žila" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%A2%D1%80%D0%B0%D1%98%D0%BD%D0%B0_%D1%82%D0%BA%D0%B8%D0%B2%D0%B0" title="Трајна ткива – Serbian" lang="sr" hreflang="sr" data-title="Трајна ткива" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E7%B6%AD%E7%AE%A1%E6%9D%9F" title="維管束 – Cantonese" lang="yue" hreflang="yue" data-title="維管束" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E7%BB%B4%E7%AE%A1%E6%9D%9F" title="维管束 – Chinese" lang="zh" hreflang="zh" data-title="维管束" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a 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dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Longitudinal strand of vascular tissue in the roots, stems and leaves of higher plants</div> <style data-mw-deduplicate="TemplateStyles:r1251242444">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output .ambox-content{border-left:10px solid #f28500}.mw-parser-output .ambox-style{border-left:10px solid #fc3}.mw-parser-output .ambox-move{border-left:10px solid #9932cc}.mw-parser-output .ambox-protection{border-left:10px solid #a2a9b1}.mw-parser-output .ambox .mbox-text{border:none;padding:0.25em 0.5em;width:100%}.mw-parser-output .ambox .mbox-image{border:none;padding:2px 0 2px 0.5em;text-align:center}.mw-parser-output .ambox .mbox-imageright{border:none;padding:2px 0.5em 2px 0;text-align:center}.mw-parser-output .ambox .mbox-empty-cell{border:none;padding:0;width:1px}.mw-parser-output .ambox .mbox-image-div{width:52px}@media(min-width:720px){.mw-parser-output .ambox{margin:0 10%}}@media print{body.ns-0 .mw-parser-output .ambox{display:none!important}}</style><table class="box-Expand_language plainlinks metadata ambox ambox-notice skin-invert-image" role="presentation"><tbody><tr><td class="mbox-image"><div class="mbox-image-div"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/Translation_to_english_arrow.svg/50px-Translation_to_english_arrow.svg.png" decoding="async" width="50" height="17" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/Translation_to_english_arrow.svg/75px-Translation_to_english_arrow.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/Translation_to_english_arrow.svg/100px-Translation_to_english_arrow.svg.png 2x" data-file-width="60" data-file-height="20" /></span></span></div></td><td class="mbox-text"><div class="mbox-text-span"><style data-mw-deduplicate="TemplateStyles:r1214851843">.mw-parser-output .hidden-begin{box-sizing:border-box;width:100%;padding:5px;border:none;font-size:95%}.mw-parser-output .hidden-title{font-weight:bold;line-height:1.6;text-align:left}.mw-parser-output .hidden-content{text-align:left}@media all and (max-width:500px){.mw-parser-output .hidden-begin{width:auto!important;clear:none!important;float:none!important}}</style><div class="hidden-begin mw-collapsible mw-collapsed" style="font-size:100%; padding: 0px;"><div class="hidden-title skin-nightmode-reset-color" style="text-align:center; padding-right: 3em; font-weight: normal; text-align: left">You can help <b>expand this article with text translated from <a href="https://fr.wikipedia.org/wiki/Faisceau_cribro-vasculaire" class="extiw" title="fr:Faisceau cribro-vasculaire">the corresponding article</a> in French</b>.  <i><small>(February 2019)</small></i> <small>Click [show] for important translation instructions.</small></div><div class="hidden-content mw-collapsible-content" style=""> <ul><li><a rel="nofollow" class="external text" href="https://translate.google.com/translate?&u=https%3A%2F%2Ffr.wikipedia.org%2Fwiki%2FFaisceau_cribro-vasculaire&sl=fr&tl=en&prev=_t&hl=en">View</a> a machine-translated version of the French article.</li> <li>Machine translation, like <a rel="nofollow" class="external text" href="https://deepl.com">DeepL</a> or <a rel="nofollow" class="external text" href="https://translate.google.com/">Google Translate</a>, is a useful starting point for translations, but translators must revise errors as necessary and confirm that the translation is accurate, rather than simply copy-pasting machine-translated text into the English Wikipedia.</li> <li>Consider <b><a href="/wiki/Template:Expand_French#Topics_and_categorization" title="Template:Expand French">adding a topic</a></b> to this template: there are already 1,627 articles in the <a href="/wiki/Category:Articles_needing_translation_from_French_Wikipedia" title="Category:Articles needing translation from French Wikipedia">main category</a>, and specifying<code class="tpl-para" style="word-break:break-word;">|topic=</code> will aid in categorization.</li> <li>Do not translate text that appears unreliable or low-quality. If possible, verify the text with references provided in the foreign-language article.</li> <li>You <b>must</b> provide <a href="/wiki/Wikipedia:Copying_within_Wikipedia" title="Wikipedia:Copying within Wikipedia">copyright attribution</a> in the <a href="/wiki/Help:Edit_summary" title="Help:Edit summary">edit summary</a> accompanying your translation by providing an <a href="/wiki/Help:Interlanguage_links" title="Help:Interlanguage links">interlanguage link</a> to the source of your translation. A model attribution edit summary is <code>Content in this edit is translated from the existing French Wikipedia article at [[:fr:Faisceau cribro-vasculaire]]; see its history for attribution.</code></li> <li>You may also add the template <code>{{Translated|fr|Faisceau cribro-vasculaire}}</code> to the <a href="/wiki/Talk:Vascular_bundle" title="Talk:Vascular bundle">talk page</a>.</li> <li>For more guidance, see <a href="/wiki/Wikipedia:Translation" title="Wikipedia:Translation">Wikipedia:Translation</a>.</li></ul></div></div></div></td></tr></tbody></table> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Leitb%C3%BCndel.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Leitb%C3%BCndel.svg/220px-Leitb%C3%BCndel.svg.png" decoding="async" width="220" height="154" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Leitb%C3%BCndel.svg/330px-Leitb%C3%BCndel.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Leitb%C3%BCndel.svg/440px-Leitb%C3%BCndel.svg.png 2x" data-file-width="363" data-file-height="254" /></a><figcaption><b>Types of Vascular bundles</b><br /> (black: Xylem, green: Phloem, white: Cambium)<br /> <b>A</b>  concentric, periphloematic(Hadrocentric)<br /> <b>B</b>  concentric, perixylematic(Leptocentric)<br /> <b>C</b>  radial with inner xylem, here with four xylem-poles, left closed, right open<br /> <b>D</b>  collateral closed<br /> <b>E</b>  collateral open<br /> <b>F</b>  bicollateral open</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Celery_cross_section.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/db/Celery_cross_section.jpg/220px-Celery_cross_section.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/db/Celery_cross_section.jpg/330px-Celery_cross_section.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/db/Celery_cross_section.jpg/440px-Celery_cross_section.jpg 2x" data-file-width="1600" data-file-height="1200" /></a><figcaption>Cross section of <a href="/wiki/Celery" title="Celery">celery</a> stalk, showing vascular bundles, which include both phloem and xylem</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Leitbuendel_zea_mays.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/1a/Leitbuendel_zea_mays.jpg/220px-Leitbuendel_zea_mays.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/1a/Leitbuendel_zea_mays.jpg/330px-Leitbuendel_zea_mays.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/1a/Leitbuendel_zea_mays.jpg/440px-Leitbuendel_zea_mays.jpg 2x" data-file-width="640" data-file-height="480" /></a><figcaption>Detail of vascular bundle: closed, collateral vascular bundles of the stem axis of <i><a href="/wiki/Zea_mays" class="mw-redirect" title="Zea mays">Zea mays</a></i></figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Metasequoia_glyptostroboides_leaf_vascular_bundle.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d9/Metasequoia_glyptostroboides_leaf_vascular_bundle.jpg/220px-Metasequoia_glyptostroboides_leaf_vascular_bundle.jpg" decoding="async" width="220" height="225" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d9/Metasequoia_glyptostroboides_leaf_vascular_bundle.jpg/330px-Metasequoia_glyptostroboides_leaf_vascular_bundle.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d9/Metasequoia_glyptostroboides_leaf_vascular_bundle.jpg/440px-Metasequoia_glyptostroboides_leaf_vascular_bundle.jpg 2x" data-file-width="800" data-file-height="817" /></a><figcaption>Vascular bundle in the <a href="/wiki/Leaf" title="Leaf">leaf</a> of <i><a href="/wiki/Metasequoia_glyptostroboides" title="Metasequoia glyptostroboides">Metasequoia glyptostroboides</a></i></figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Pine_Leaf_Vascular_Bundle.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/dc/Pine_Leaf_Vascular_Bundle.jpg/220px-Pine_Leaf_Vascular_Bundle.jpg" decoding="async" width="220" height="293" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/dc/Pine_Leaf_Vascular_Bundle.jpg/330px-Pine_Leaf_Vascular_Bundle.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/dc/Pine_Leaf_Vascular_Bundle.jpg/440px-Pine_Leaf_Vascular_Bundle.jpg 2x" data-file-width="2448" data-file-height="3264" /></a><figcaption>The vascular bundle of pine leaf showing xylem and phloem</figcaption></figure> <p>A <b>vascular bundle</b> is a part of the transport system in <a href="/wiki/Vascular_plant" title="Vascular plant">vascular plants</a>. The transport itself happens in the <a href="/wiki/Plant_stem" title="Plant stem">stem</a>, which exists in two forms: <a href="/wiki/Xylem" title="Xylem">xylem</a> and <a href="/wiki/Phloem" title="Phloem">phloem</a>. Both these tissues are present in a vascular bundle, which in addition will include supporting and protective tissues. There is also a tissue between xylem and phloem, which is the <a href="/wiki/Cambium" title="Cambium">cambium</a>. </p><p>The xylem typically lies towards the axis (<a href="https://en.wiktionary.org/wiki/adaxial" class="extiw" title="wiktionary:adaxial">adaxial</a>) with phloem positioned away from the axis (<a href="https://en.wiktionary.org/wiki/abaxial" class="extiw" title="wiktionary:abaxial">abaxial</a>). In a stem or root this means that the xylem is closer to the centre of the stem or root while the phloem is closer to the exterior. In a leaf, the adaxial surface of the leaf will usually be the upper side, with the abaxial surface the lower side. </p><p>The sugars synthesized by the plant with sun light are transported by the phloem, which is closer to the lower surface. <a href="/wiki/Aphids" class="mw-redirect" title="Aphids">Aphids</a> and <a href="/wiki/Leaf_hopper" class="mw-redirect" title="Leaf hopper">leaf hoppers</a> feed off of these sugars by tapping into the phloem. This is why aphids and leaf hoppers are typically found on the underside of a leaf rather than on the top. The position of vascular bundles relative to each other may vary considerably: see <a href="/wiki/Stele_(biology)" title="Stele (biology)">stele</a>. The vascular bundle are depend on size of veins </p><p><br /> </p><p><br /> </p> <style data-mw-deduplicate="TemplateStyles:r1062633282">@media all and (max-width:720px){.mw-parser-output .content .thumb>div:not(.thumbinner){display:flex;justify-content:center;flex-wrap:wrap;align-content:flex-start;flex-direction:column}}body.skin-vector .mw-parser-output div.thumb>div:not(.thumbinner){font-size:94%;text-align:center;overflow:hidden;min-width:100px}body.skin-minerva .mw-parser-output div.thumb>div:not(.thumbinner){margin:0 auto;max-width:100%!important}</style> <div class="center"> <div class="thumb tnone"> <div style="display:table;width:600px;"> <span typeof="mw:File/Frameless"><a href="/wiki/File:Plant_cell_types.svg" class="mw-file-description" title="Cross section of a leaf showing parts of a vascular bundle"><img alt="Cross section of a leaf showing parts of a vascular bundle" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e7/Plant_cell_types.svg/600px-Plant_cell_types.svg.png" decoding="async" width="600" height="423" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e7/Plant_cell_types.svg/900px-Plant_cell_types.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e7/Plant_cell_types.svg/1200px-Plant_cell_types.svg.png 2x" data-file-width="1900" data-file-height="1340" /></a></span><div class="thumbcaption" style="display:table-caption;caption-side:bottom;box-sizing:border-box;">Cross section of a leaf showing parts of a vascular bundle </div> </div> </div></div> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Bundle-sheath_cells">Bundle-sheath cells</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Vascular_bundle&action=edit&section=1" title="Edit section: Bundle-sheath cells"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The bundle-sheath cells are the photosynthetic cells arranged into a tightly packed sheath around the vein of a leaf. It forms a protective covering on the leaf vein and consists of one or more cell layers, usually <a href="/wiki/Ground_tissue#Parenchyma" title="Ground tissue">parenchyma</a>. Loosely-arranged <a href="/wiki/Leaf#Mesophyll" title="Leaf">mesophyll</a> cells lie between the bundle sheath and the leaf surface. The <a href="/wiki/Calvin_cycle" title="Calvin cycle">Calvin cycle</a> is confined to the <a href="/wiki/Chloroplasts" class="mw-redirect" title="Chloroplasts">chloroplasts</a> of these bundle sheath cells in <a href="/wiki/C4_carbon_fixation" title="C4 carbon fixation">C<sub>4</sub> plants</a>. <a href="/wiki/C2_photosynthesis" class="mw-redirect" title="C2 photosynthesis">C<sub>2</sub> plants</a> also use a variation of this structure.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Vascular_bundle&action=edit&section=2" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFSageKhoshraveshSage2014" class="citation journal cs1">Sage, Rowan F.; Khoshravesh, Roxana; Sage, Tammy L. (1 July 2014). <a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fjxb%2Feru180">"From proto-Kranz to C4 Kranz: building the bridge to C4 photosynthesis"</a>. <i>Journal of Experimental Botany</i>. <b>65</b> (13): 3341–3356. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fjxb%2Feru180">10.1093/jxb/eru180</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/24803502">24803502</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Experimental+Botany&rft.atitle=From+proto-Kranz+to+C4+Kranz%3A+building+the+bridge+to+C4+photosynthesis&rft.volume=65&rft.issue=13&rft.pages=3341-3356&rft.date=2014-07-01&rft_id=info%3Adoi%2F10.1093%2Fjxb%2Feru180&rft_id=info%3Apmid%2F24803502&rft.aulast=Sage&rft.aufirst=Rowan+F.&rft.au=Khoshravesh%2C+Roxana&rft.au=Sage%2C+Tammy+L.&rft_id=https%3A%2F%2Fdoi.org%2F10.1093%252Fjxb%252Feru180&rfr_id=info%3Asid%2Fen.wikipedia.org%3AVascular+bundle" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="Further_reading">Further reading</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Vascular_bundle&action=edit&section=3" title="Edit section: Further reading"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>Campbell, N. A. & Reece, J. B. (2005). Photosynthesis. <i>Biology</i> (7th ed.). San Francisco: Benjamin Cummings.</li></ul> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Vascular_bundle&action=edit&section=4" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="http://botweb.uwsp.edu/anatomy/primaryxylem.htm">Curtis, Lersten, and Nowak</a> cross section of a vascular bundle</li> <li><a rel="nofollow" class="external text" href="http://www.sbs.utexas.edu/mauseth/weblab/webchap8phloem/chapter_8.htm">Mauseth</a> another cross section of a vascular bundle</li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output 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td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"></div><div role="navigation" class="navbox" aria-labelledby="Botany" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2" 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<ul><li><a href="/wiki/History_of_botany" title="History of botany">History</a></li> <li><a href="/wiki/Outline_of_botany" title="Outline of botany">Outline</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Branches_of_botany" title="Branches of botany">Subdisciplines</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archaeobotany" class="mw-redirect" title="Archaeobotany">Archaeobotany</a></li> <li><a href="/wiki/Astrobotany" title="Astrobotany">Astrobotany</a></li> <li><a href="/wiki/Bryology" title="Bryology">Bryology</a></li> <li><a href="/wiki/Dendrology" title="Dendrology">Dendrology</a></li> <li><a href="/wiki/Ethnobotany" title="Ethnobotany">Ethnobotany</a></li> <li><a href="/wiki/Paleobotany" title="Paleobotany">Paleobotany</a></li> <li><a href="/wiki/Phycology" title="Phycology">Phycology</a></li> <li><a href="/wiki/Phytochemistry" title="Phytochemistry">Phytochemistry</a></li> <li><a href="/wiki/Phytogeography" title="Phytogeography">Phytogeography</a> <ul><li><a href="/wiki/Geobotanical_prospecting" title="Geobotanical prospecting">Geobotany</a></li></ul></li> <li><a href="/wiki/Plant_anatomy" title="Plant anatomy">Plant anatomy</a></li> <li><a href="/wiki/Plant_ecology" title="Plant ecology">Plant ecology</a></li> <li><a href="/wiki/Plant_pathology" title="Plant pathology">Plant pathology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant" title="Plant">Plant</a> groups</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Algae" title="Algae">Algae</a></li> <li><a href="/wiki/Archaeplastida" title="Archaeplastida">Archaeplastida</a></li> <li><a href="/wiki/Bryophyte" title="Bryophyte">Bryophyte</a></li> <li><a href="/wiki/Non-vascular_plant" title="Non-vascular plant">Non-vascular plants</a></li> <li><a href="/wiki/Vascular_plant" title="Vascular plant">Vascular plants</a></li> <li><a href="/wiki/Fern" title="Fern">Fern</a></li> <li><a href="/wiki/Lycophyte" title="Lycophyte">Lycophyte</a></li> <li><a href="/wiki/Spermatophyte" class="mw-redirect" title="Spermatophyte">Spermatophytes</a></li> <li><a href="/wiki/Gymnosperm" title="Gymnosperm">Gymnosperm</a></li> <li><a href="/wiki/Flowering_plant" title="Flowering plant">Angiosperm</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant_anatomy" title="Plant anatomy">Plant anatomy</a> <div class="hlist"><ul><li><a href="/wiki/Plant_morphology" title="Plant morphology">Plant morphology</a><br /><span class="nobold">(<a href="/wiki/Glossary_of_plant_morphology" title="Glossary of plant morphology">glossary</a>)</span></li></ul></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%"><a href="/wiki/Plant_cell" title="Plant cell">Plant cells</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cell_wall" title="Cell wall">Cell wall</a></li> <li><a href="/wiki/Phragmoplast" title="Phragmoplast">Phragmoplast</a></li> <li><a href="/wiki/Plastid" title="Plastid">Plastid</a></li> <li><a href="/wiki/Plasmodesma" title="Plasmodesma">Plasmodesma</a></li> <li><a href="/wiki/Vacuole" title="Vacuole">Vacuole</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%"><a href="/wiki/Tissue_(biology)" title="Tissue (biology)">Tissues</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cork_cambium" title="Cork cambium">Cork</a></li> <li><a href="/wiki/Ground_tissue" title="Ground tissue">Ground tissue</a> <ul><li><a href="/wiki/Leaf#Mesophyll" title="Leaf">Mesophyll</a></li></ul></li> <li><a href="/wiki/Meristem" title="Meristem">Meristem</a></li> <li><a href="/wiki/Storage_organ" title="Storage organ">Storage organs</a></li> <li><a href="/wiki/Vascular_tissue" title="Vascular tissue">Vascular tissue</a> <ul><li><a class="mw-selflink selflink">Vascular bundle</a></li></ul></li> <li><a href="/wiki/Wood" title="Wood">Wood</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%">Vegetative</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bulb" title="Bulb">Bulb</a></li> <li><a href="/wiki/Root" title="Root">Root</a></li> <li><a href="/wiki/Rhizoid" title="Rhizoid">Rhizoid</a></li> <li><a href="/wiki/Rhizome" title="Rhizome">Rhizome</a></li> <li><a href="/wiki/Shoot_(botany)" title="Shoot (botany)">Shoot</a> <ul><li><a href="/wiki/Bud" title="Bud">Bud</a></li> <li><a href="/wiki/Leaf" title="Leaf">Leaf</a> <ul><li><a href="/wiki/Cataphyll" title="Cataphyll">Cataphyll</a></li> <li><a href="/wiki/Petiole_(botany)" title="Petiole (botany)">Petiole</a></li></ul></li> <li><a href="/wiki/Sessility_(botany)" title="Sessility (botany)">Sessility</a></li> <li><a href="/wiki/Plant_stem" title="Plant stem">Stem</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%"><a href="/wiki/Plant_reproductive_morphology" title="Plant reproductive morphology">Reproductive</a><br />(incl. Flower)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archegonium" title="Archegonium">Archegonium</a></li> <li><a href="/wiki/Antheridium" title="Antheridium">Antheridium</a></li> <li><a href="/wiki/Stamen" title="Stamen">Androecium</a> <ul><li><a href="/wiki/Pollen" title="Pollen">Pollen</a></li> <li><a href="/wiki/Stamen" title="Stamen">Stamen</a> <ul><li><a href="/wiki/Anther" class="mw-redirect" title="Anther">Anther</a></li> <li><a href="/wiki/Stamen#Morphology_and_terminology" title="Stamen">Filament</a></li></ul></li> <li><a href="/wiki/Staminode" title="Staminode">Staminode</a></li> <li><a href="/wiki/Tapetum_(botany)" title="Tapetum (botany)">Tapetum</a></li></ul></li> <li><a href="/wiki/Flower" title="Flower">Flower</a> <ul><li><a href="/wiki/Aestivation_(botany)" title="Aestivation (botany)">Aestivation</a></li> <li><a href="/wiki/ABC_model_of_flower_development" title="ABC model of flower development">Flower development</a></li> <li><a href="/wiki/Floral_diagram" title="Floral diagram">Floral diagram</a></li> <li><a href="/wiki/Floral_formula" title="Floral formula">Floral formula</a></li> <li><a href="/wiki/Floral_symmetry" title="Floral symmetry">Floral symmetry</a></li> <li><a href="/wiki/Whorl_(botany)" title="Whorl (botany)">Whorl</a></li></ul></li> <li><a href="/wiki/Fruit" title="Fruit">Fruit</a> <ul><li><a href="/wiki/Fruit_anatomy" class="mw-redirect" title="Fruit anatomy">Anatomy</a></li> <li><a href="/wiki/Berry_(botany)" title="Berry (botany)">Berry</a></li> <li><a href="/wiki/Capsule_(fruit)" title="Capsule (fruit)">Capsule</a></li> <li><a href="/wiki/Nut_(fruit)" title="Nut (fruit)">Nut</a></li> <li><a href="/wiki/Pyrena" title="Pyrena">Pyrena</a></li> <li><a href="/wiki/Seed" title="Seed">Seed</a> <ul><li><a href="/wiki/Seed_dispersal" title="Seed dispersal">Dispersal</a></li> <li><a href="/wiki/Endosperm" title="Endosperm">Endosperm</a></li></ul></li></ul></li> <li><a href="/wiki/Gametophyte" title="Gametophyte">Gametophyte</a></li> <li><a href="/wiki/Column_(botany)" title="Column (botany)">Gynandrium</a></li> <li><a href="/wiki/Gynoecium" title="Gynoecium">Gynoecium</a> <ul><li><a href="/wiki/Carpel" class="mw-redirect" title="Carpel">Carpel</a> <ul><li><a href="/wiki/Ovary_(botany)" title="Ovary (botany)">Ovary</a> <ul><li><a href="/wiki/Locule" title="Locule">Locule</a></li> <li><a href="/wiki/Ovule" title="Ovule">Ovule</a></li></ul></li> <li><a href="/wiki/Stigma_(botany)" title="Stigma (botany)">Stigma</a></li> <li><a href="/wiki/Style_(botany)" title="Style (botany)">Style</a></li></ul></li></ul></li> <li><a href="/wiki/Hypanthium" title="Hypanthium">Hypanthium (Floral cup)</a></li> <li><a href="/wiki/Inflorescence" title="Inflorescence">Inflorescence</a> <ul><li><a href="/wiki/Bract" title="Bract">Bract</a></li> <li><a href="/wiki/Pedicel_(botany)" title="Pedicel (botany)">Pedicellate</a></li> <li><a href="/wiki/Raceme" title="Raceme">Raceme</a></li> <li><a href="/wiki/Umbel" title="Umbel">Umbel</a></li></ul></li> <li><a href="/wiki/Perianth" title="Perianth">Perianth</a> <ul><li><a href="/wiki/Tepal" title="Tepal">Tepal</a></li> <li><a href="/wiki/Petal" title="Petal">Petal</a></li> <li><a href="/wiki/Sepal" title="Sepal">Sepal</a></li></ul></li> <li><a href="/wiki/Embryo#Plant_embryos" title="Embryo">Plant embryo</a></li> <li><a href="/wiki/Receptacle_(botany)" title="Receptacle (botany)">Receptacle</a></li> <li><a href="/wiki/Sporophyll" title="Sporophyll">Sporophyll</a></li> <li><a href="/wiki/Sporophyte" title="Sporophyte">Sporophyte</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%">Surface structures</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Plant_cuticle" title="Plant cuticle">Cuticle</a></li> <li><a href="/wiki/Epicuticular_wax" title="Epicuticular wax">Epicuticular wax</a></li> <li><a href="/wiki/Epidermis_(botany)" title="Epidermis (botany)">Epidermis</a></li> <li><a href="/wiki/Nectar" title="Nectar">Nectar</a></li> <li><a href="/wiki/Stoma" title="Stoma">Stoma</a></li> <li><a href="/wiki/Thorns,_spines,_and_prickles" title="Thorns, spines, and prickles">Thorns, spines, and prickles</a></li> <li><a href="/wiki/Trichome" title="Trichome">Trichome</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant_physiology" title="Plant physiology">Plant physiology</a><br />Materials</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aleurone" title="Aleurone">Aleurone</a></li> <li><a href="/wiki/Apical_dominance" title="Apical dominance">Apical dominance</a></li> <li><a href="/wiki/Bulk_movement" title="Bulk movement">Bulk flow</a></li> <li><a href="/wiki/Cellulose" title="Cellulose">Cellulose</a></li> <li><a href="/wiki/Plant_nutrition" title="Plant nutrition">Nutrition</a></li> <li><a href="/wiki/Photosynthesis" title="Photosynthesis">Photosynthesis</a> <ul><li><a href="/wiki/Chlorophyll" title="Chlorophyll">Chlorophyll</a></li></ul></li> <li><a href="/wiki/Phytomelanin" title="Phytomelanin">Phytomelanin</a></li> <li><a href="/wiki/Plant_hormone" title="Plant hormone">Plant hormones</a></li> <li><a href="/wiki/Respiratory_system#Plants" title="Respiratory system">Respiration</a> <ul><li><a href="/wiki/Gas_exchange#Plants" title="Gas exchange">Gas Exchange</a></li> <li><a href="/wiki/Cellular_respiration" title="Cellular respiration">Cellular respiration</a></li></ul></li> <li><a href="/wiki/Sap" title="Sap">Sap</a></li> <li><a href="/wiki/Starch" title="Starch">Starch</a></li> <li><a href="/wiki/Sugar" title="Sugar">Sugar</a></li> <li><a href="/wiki/Transpiration" title="Transpiration">Transpiration</a></li> <li><a href="/wiki/Turgor_pressure" title="Turgor pressure">Turgor pressure</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%">Plant growth<br />and habit</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Habit_(biology)#Structure" title="Habit (biology)">Habit</a> <ul><li><a href="/wiki/Cushion_plant" title="Cushion plant">Cushion plants</a></li> <li><a href="/wiki/Rosette_(botany)" title="Rosette (botany)">Rosettes</a></li> <li><a href="/wiki/Shrub" title="Shrub">Shrubs</a> <ul><li><a href="/wiki/Prostrate_shrub" title="Prostrate shrub">Prostrate shrubs</a></li> <li><a href="/wiki/Subshrub" title="Subshrub">Subshrubs</a></li></ul></li> <li><a href="/wiki/Succulent_plant" title="Succulent plant">Succulent plants</a></li> <li><a href="/wiki/Tree" title="Tree">Trees</a></li> <li><a href="/wiki/Vine" title="Vine">Vines</a> <ul><li><a href="/wiki/Liana" title="Liana">Lianas</a></li></ul></li></ul></li> <li><a href="/wiki/Herbaceous_plant" title="Herbaceous plant">Herbaceous plants</a></li> <li><a href="/wiki/Secondary_growth" title="Secondary growth">Secondary growth</a></li> <li><a href="/wiki/Woody_plant" title="Woody plant">Woody plants</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant_reproduction" title="Plant reproduction">Reproduction</a> <div class="hlist"><ul><li><a href="/wiki/Plant_evolution" title="Plant evolution">Evolution</a></li><li><a href="/wiki/Plant_ecology" title="Plant ecology">Ecology</a></li></ul></div></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Alternation_of_generations" title="Alternation of generations">Alternation of generations</a></li> <li><a href="/wiki/Double_fertilization" title="Double fertilization">Double fertilization</a></li> <li><a href="/wiki/Plant_evolutionary_developmental_biology" title="Plant evolutionary developmental biology">Evolutionary development</a></li> <li><a href="/wiki/Evolutionary_history_of_plants" title="Evolutionary history of plants">Evolutionary history</a> <ul><li><a href="/wiki/Timeline_of_plant_evolution" title="Timeline of plant evolution">timeline</a></li></ul></li> <li><a href="/wiki/Flora" title="Flora">Flora</a></li> <li><a href="/wiki/Germination" title="Germination">Germination</a></li> <li><a href="/wiki/Pollination" title="Pollination">Pollination</a> <ul><li><a href="/wiki/Artificial_pollination" class="mw-redirect" title="Artificial pollination">Artificial</a></li> <li><a href="/wiki/Pollinator" title="Pollinator">Pollinators</a></li> <li><a href="/wiki/Pollen_tube" title="Pollen tube">Pollen tube</a></li> <li><a href="/wiki/Self-pollination" title="Self-pollination">Self</a></li></ul></li> <li><a href="/wiki/Sporangium" title="Sporangium">Sporangium</a> <ul><li><a href="/wiki/Microsporangia" class="mw-redirect" title="Microsporangia">Microsporangia</a> <ul><li><a href="/wiki/Microspore" title="Microspore">Microspore</a></li></ul></li> <li><a href="/wiki/Sporangium" title="Sporangium">Megasporangium</a> <ul><li><a href="/wiki/Megaspore" title="Megaspore">Megaspore</a></li></ul></li> <li><a href="/wiki/Spore" title="Spore">Spore</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant_taxonomy" title="Plant taxonomy">Plant taxonomy</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Biological classification</a></li> <li><a href="/wiki/Botanical_nomenclature" title="Botanical nomenclature">Botanical nomenclature</a> <ul><li><a href="/wiki/Botanical_name" title="Botanical name">Botanical name</a></li> <li><a href="/wiki/Correct_name" title="Correct name">Correct name</a></li> <li><a href="/wiki/Author_citation_(botany)" title="Author citation (botany)">Author citation</a></li> <li><a href="/wiki/International_Code_of_Nomenclature_for_algae,_fungi,_and_plants" title="International Code of Nomenclature for algae, fungi, and plants">International Code of Nomenclature (ICN)</a></li> <li><a href="/wiki/International_Code_of_Nomenclature_for_Cultivated_Plants" title="International Code of Nomenclature for Cultivated Plants">ICN for Cultivated Plants (ICNCP)</a></li></ul></li> <li><a href="/wiki/Cultivated_plant_taxonomy" title="Cultivated plant taxonomy">Cultivated plant taxonomy</a> <ul><li><a href="/wiki/Citrus_taxonomy" title="Citrus taxonomy">Citrus taxonomy</a></li> <li><a href="/wiki/Cultigen" title="Cultigen">Cultigen</a> <ul><li><a href="/wiki/Cultivar" title="Cultivar">Cultivar</a></li> <li><a href="/wiki/Cultivar_group" title="Cultivar group">Group</a></li> <li><a href="/wiki/Grex_(horticulture)" title="Grex (horticulture)">Grex</a></li></ul></li></ul></li> <li><a href="/wiki/History_of_plant_systematics" title="History of plant systematics">History of plant systematics</a></li> <li><a href="/wiki/Herbarium" title="Herbarium">Herbarium</a></li> <li><a href="/wiki/International_Association_for_Plant_Taxonomy" title="International Association for Plant Taxonomy">International Association for Plant Taxonomy</a> (IAPT)</li> <li><a href="/wiki/List_of_systems_of_plant_taxonomy" title="List of systems of plant taxonomy">Plant taxonomy systems</a></li> <li><a href="/wiki/Taxonomic_rank" title="Taxonomic rank">Taxonomic rank</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%">Practice</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agronomy" title="Agronomy">Agronomy</a></li> <li><a href="/wiki/Floriculture" title="Floriculture">Floriculture</a></li> <li><a href="/wiki/Forestry" title="Forestry">Forestry</a></li> <li><a href="/wiki/Horticulture" title="Horticulture">Horticulture</a></li> <li><a href="/wiki/Phytochemical" title="Phytochemical">Phytochemical</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><div class="hlist"><ul><li>Lists</li><li>Related topics</li></ul></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Glossary_of_botanical_terms" title="Glossary of botanical terms">Botanical terms</a></li> <li><a href="/wiki/List_of_botanists" title="List of botanists">Botanists</a> <ul><li><a href="/wiki/List_of_botanists_by_author_abbreviation_(W%E2%80%93Z)" title="List 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