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Stanley B. Prusiner - Wikipedia
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vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" 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Prusiner</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 55 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-55" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">55 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B3%D8%AA%D8%A7%D9%86%D9%84%D9%8A_%D8%A8%D8%B1%D9%88%D8%B3%D9%8A%D9%86%D8%B1" title="ستانلي بروسينر – Arabic" lang="ar" hreflang="ar" data-title="ستانلي بروسينر" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Stenli_Pruziner" title="Stenli Pruziner – Azerbaijani" lang="az" hreflang="az" data-title="Stenli Pruziner" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%A7%D8%B3%D8%AA%D9%86%D9%84%DB%8C_%D9%BE%D8%B1%DB%8C%D8%B3%DB%8C%D9%86%D8%B1" title="استنلی پریسینر – South Azerbaijani" lang="azb" hreflang="azb" data-title="استنلی پریسینر" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B8%E0%A7%8D%E0%A6%9F%E0%A6%BE%E0%A6%A8%E0%A6%B2%E0%A6%BF_%E0%A6%AC%E0%A6%BF_%E0%A6%AA%E0%A7%8D%E0%A6%B0%E0%A7%81%E0%A6%B8%E0%A6%BF%E0%A6%A8%E0%A6%BE%E0%A6%B0" title="স্টানলি বি প্রুসিনার – Bangla" lang="bn" hreflang="bn" data-title="স্টানলি বি প্রুসিনার" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A1%D1%82%D1%8D%D0%BD%D0%BB%D1%96_%D0%91%D0%B5%D0%BD_%D0%9F%D1%80%D1%83%D0%B7%D1%96%D0%BD%D0%B5%D1%80" title="Стэнлі Бен Прузінер – Belarusian" lang="be" hreflang="be" data-title="Стэнлі Бен Прузінер" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-be-x-old mw-list-item"><a href="https://be-tarask.wikipedia.org/wiki/%D0%A1%D1%82%D1%8D%D0%BD%D0%BB%D1%96_%D0%9F%D1%80%D1%83%D0%B7%D1%96%D0%BD%D1%8D%D1%80" title="Стэнлі Прузінэр – Belarusian (Taraškievica orthography)" lang="be-tarask" hreflang="be-tarask" data-title="Стэнлі Прузінэр" data-language-autonym="Беларуская (тарашкевіца)" data-language-local-name="Belarusian (Taraškievica orthography)" class="interlanguage-link-target"><span>Беларуская (тарашкевіца)</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D0%B8_%D0%9F%D1%80%D1%83%D1%81%D0%B8%D0%BD%D1%8A%D1%80" title="Стенли Прусинър – Bulgarian" lang="bg" hreflang="bg" data-title="Стенли Прусинър" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Stanley_Ben_Prusiner" title="Stanley Ben Prusiner – Catalan" lang="ca" hreflang="ca" data-title="Stanley Ben Prusiner" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Czech" lang="cs" hreflang="cs" data-title="Stanley B. Prusiner" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – German" lang="de" hreflang="de" data-title="Stanley Prusiner" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Estonian" lang="et" hreflang="et" data-title="Stanley Prusiner" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%A3%CF%84%CE%AC%CE%BD%CE%BB%CE%B5%CF%8B_%CE%A0%CF%81%CE%BF%CF%8D%CE%B6%CE%B9%CE%BD%CE%B5%CF%81" title="Στάνλεϋ Προύζινερ – Greek" lang="el" hreflang="el" data-title="Στάνλεϋ Προύζινερ" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Spanish" lang="es" hreflang="es" data-title="Stanley B. Prusiner" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Basque" lang="eu" hreflang="eu" data-title="Stanley Prusiner" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%A7%D8%B3%D8%AA%D9%86%D9%84%DB%8C_%D9%BE%D8%B1%DB%8C%D8%B3%DB%8C%D9%86%D8%B1" title="استنلی پریسینر – Persian" lang="fa" hreflang="fa" data-title="استنلی پریسینر" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – French" lang="fr" hreflang="fr" data-title="Stanley Prusiner" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Irish" lang="ga" hreflang="ga" data-title="Stanley Prusiner" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Galician" lang="gl" hreflang="gl" data-title="Stanley B. Prusiner" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%8A%A4%ED%83%A0%EB%A6%AC_B._%ED%94%84%EB%A3%A8%EC%8B%9C%EB%84%88" title="스탠리 B. 프루시너 – Korean" lang="ko" hreflang="ko" data-title="스탠리 B. 프루시너" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Croatian" lang="hr" hreflang="hr" data-title="Stanley B. Prusiner" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Stanley_Ben_Prusiner" title="Stanley Ben Prusiner – Ido" lang="io" hreflang="io" data-title="Stanley Ben Prusiner" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Indonesian" lang="id" hreflang="id" data-title="Stanley B. Prusiner" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Italian" lang="it" hreflang="it" data-title="Stanley Prusiner" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A1%D7%98%D7%A0%D7%9C%D7%99_%D7%A4%D7%A8%D7%95%D7%96%D7%99%D7%A0%D7%A8" title="סטנלי פרוזינר – Hebrew" lang="he" hreflang="he" data-title="סטנלי פרוזינר" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D0%B8_%D0%9F%D1%80%D1%83%D0%B7%D0%B8%D0%BD%D0%B5%D1%80" title="Стенли Прузинер – Kazakh" lang="kk" hreflang="kk" data-title="Стенли Прузинер" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Swahili" lang="sw" hreflang="sw" data-title="Stanley Prusiner" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Latin" lang="la" hreflang="la" data-title="Stanley B. Prusiner" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Hungarian" lang="hu" hreflang="hu" data-title="Stanley B. Prusiner" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%B8%E0%B5%8D%E0%B4%B1%E0%B5%8D%E0%B4%B1%E0%B4%BE%E0%B5%BB%E0%B4%B2%E0%B4%BF_%E0%B4%AC%E0%B5%86%E0%B4%9E%E0%B5%8D%E0%B4%9A%E0%B4%AE%E0%B4%BF%E0%B5%BB_%E0%B4%AA%E0%B5%8D%E0%B4%B0%E0%B5%81%E0%B4%B8%E0%B4%BF%E0%B4%A8%E0%B5%BC" title="സ്റ്റാൻലി ബെഞ്ചമിൻ പ്രുസിനർ – Malayalam" lang="ml" hreflang="ml" data-title="സ്റ്റാൻലി ബെഞ്ചമിൻ പ്രുസിനർ" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%B3%D8%AA%D8%A7%D9%86%D9%84%D9%89_%D8%A8%D8%B1%D9%88%D8%B3%D9%8A%D9%86%D8%B1" title="ستانلى بروسينر – Egyptian Arabic" lang="arz" hreflang="arz" data-title="ستانلى بروسينر" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%A7%D8%B3%D8%AA%D9%86%D9%84%DB%8C_%D9%BE%D8%B1%DB%8C%D8%B3%DB%8C%D9%86%D8%B1" title="استنلی پریسینر – Mazanderani" lang="mzn" hreflang="mzn" data-title="استنلی پریسینر" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Malay" lang="ms" hreflang="ms" data-title="Stanley B. Prusiner" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Dutch" lang="nl" hreflang="nl" data-title="Stanley B. Prusiner" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B9%E3%82%BF%E3%83%B3%E3%83%AA%E3%83%BC%E3%83%BBB%E3%83%BB%E3%83%97%E3%83%AB%E3%82%B7%E3%83%8A%E3%83%BC" title="スタンリー・B・プルシナー – Japanese" lang="ja" hreflang="ja" data-title="スタンリー・B・プルシナー" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Stanley B. Prusiner" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Occitan" lang="oc" hreflang="oc" data-title="Stanley B. Prusiner" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%B3%D9%B9%DB%8C%D9%86%D9%84%DB%8C_%D9%BE%D8%B1%D9%88%D8%B3%D9%86%D8%B1" title="سٹینلی پروسنر – Western Punjabi" lang="pnb" hreflang="pnb" data-title="سٹینلی پروسنر" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Stanley_Benjamin_Prusiner" title="Stanley Benjamin Prusiner – Low German" lang="nds" hreflang="nds" data-title="Stanley Benjamin Prusiner" data-language-autonym="Plattdüütsch" data-language-local-name="Low German" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Polish" lang="pl" hreflang="pl" data-title="Stanley Prusiner" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Portuguese" lang="pt" hreflang="pt" data-title="Stanley Prusiner" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Romanian" lang="ro" hreflang="ro" data-title="Stanley B. Prusiner" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9F%D1%80%D1%83%D0%B7%D0%B8%D0%BD%D0%B5%D1%80,_%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D0%B8" title="Прузинер, Стенли – Russian" lang="ru" hreflang="ru" data-title="Прузинер, Стенли" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Simple English" lang="en-simple" hreflang="en-simple" data-title="Stanley B. Prusiner" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Slovak" lang="sk" hreflang="sk" data-title="Stanley B. Prusiner" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Slovenian" lang="sl" hreflang="sl" data-title="Stanley B. Prusiner" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D0%B8_%D0%91._%D0%9F%D1%80%D1%83%D0%B7%D0%B8%D0%BD%D0%B5%D1%80" title="Стенли Б. Прузинер – Serbian" lang="sr" hreflang="sr" data-title="Стенли Б. Прузинер" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Finnish" lang="fi" hreflang="fi" data-title="Stanley Prusiner" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Swedish" lang="sv" hreflang="sv" data-title="Stanley B. Prusiner" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%A1%D1%82%D3%99%D0%BD%D0%BB%D0%B8_%D0%9F%D1%80%D1%83%D0%B7%D0%B8%D0%BD%D0%B5%D1%80" title="Стәнли Прузинер – Tatar" lang="tt" hreflang="tt" data-title="Стәнли Прузинер" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatar" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Turkish" lang="tr" hreflang="tr" data-title="Stanley B. Prusiner" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D1%96_%D0%9F%D1%80%D1%83%D0%B7%D1%96%D0%BD%D0%B5%D1%80" title="Стенлі Прузінер – Ukrainian" lang="uk" hreflang="uk" data-title="Стенлі Прузінер" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%B3%D9%B9%DB%8C%D9%86%D9%84%DB%8C_%D8%A8%DB%8C_%D9%BE%D8%B1%D9%88%D8%B3%DB%8C%D9%86%D8%B1" title="سٹینلی بی پروسینر – Urdu" lang="ur" hreflang="ur" data-title="سٹینلی بی پروسینر" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Vietnamese" lang="vi" hreflang="vi" data-title="Stanley B. Prusiner" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Yoruba" lang="yo" hreflang="yo" data-title="Stanley B. Prusiner" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%8F%B2%E5%9D%A6%E5%88%A9%C2%B7%E5%B8%83%E9%AD%AF%E5%B8%8C%E7%B4%8D" title="史坦利·布魯希納 – Chinese" lang="zh" hreflang="zh" data-title="史坦利·布魯希納" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q155380#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs 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Prusiner">← Previous revision</a> | Latest revision (diff) | Newer revision → (diff)</div></div></div></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">American neurologist and chemist</div> <p class="mw-empty-elt"> </p> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">Stanley Prusiner</div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Stanley_B._Prusiner_in_2024.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Stanley_B._Prusiner_in_2024.jpg/220px-Stanley_B._Prusiner_in_2024.jpg" decoding="async" width="220" height="330" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Stanley_B._Prusiner_in_2024.jpg/330px-Stanley_B._Prusiner_in_2024.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Stanley_B._Prusiner_in_2024.jpg/440px-Stanley_B._Prusiner_in_2024.jpg 2x" data-file-width="5085" data-file-height="7624" /></a></span><div class="infobox-caption">Prusiner in 2024</div></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><div style="display:inline" class="nickname">Stanley Ben Prusiner</div><br /><span style="display:none"> (<span class="bday">1942-05-28</span>) </span>May 28, 1942<span class="noprint ForceAgeToShow"> (age 82)</span><br /><div style="display:inline" class="birthplace"><a href="/wiki/Des_Moines,_Iowa" title="Des Moines, Iowa">Des Moines, Iowa</a>, United States</div></td></tr><tr><th scope="row" class="infobox-label">Nationality</th><td class="infobox-data category">American</td></tr><tr><th scope="row" class="infobox-label">Alma mater</th><td class="infobox-data"><a href="/wiki/University_of_Pennsylvania" title="University of Pennsylvania">University of Pennsylvania</a> (<a href="/wiki/B._A." class="mw-redirect" title="B. A.">BA</a>, <a href="/wiki/M._D." class="mw-redirect" title="M. D.">MD</a>)</td></tr><tr><th scope="row" class="infobox-label">Known for</th><td class="infobox-data"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style><div class="plainlist"> <ul><li><a href="/wiki/Prion" title="Prion">Prions</a></li> <li><a href="/wiki/Transmissible_spongiform_encephalopathy" title="Transmissible spongiform encephalopathy">Transmissible spongiform encephalopathy</a></li> <li><a href="/wiki/Creutzfeldt%E2%80%93Jakob_disease" title="Creutzfeldt–Jakob disease">Creutzfeldt–Jakob disease</a></li></ul> </div></td></tr><tr><th scope="row" class="infobox-label">Spouse</th><td class="infobox-data">Sandy Turk Prusiner<sup id="cite_ref-nobelprize_3-0" class="reference"><a href="#cite_note-nobelprize-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></td></tr><tr><th scope="row" class="infobox-label">Children</th><td class="infobox-data">two<sup id="cite_ref-nobelprize_3-1" class="reference"><a href="#cite_note-nobelprize-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"><div class="plainlist"> <ul><li><a href="/wiki/Potamkin_Prize" title="Potamkin Prize">Potamkin Prize</a> <span style="font-size:85%;">(1991)</span></li> <li><a href="/wiki/Metlife_Foundation_Award_for_Medical_Research_in_Alzheimer%27s_Disease" title="Metlife Foundation Award for Medical Research in Alzheimer's Disease">Metlife Foundation Award for Medical Research in Alzheimer's Disease</a> <span style="font-size:85%;">(1991)</span><sup id="cite_ref-MetLife_1-0" class="reference"><a href="#cite_note-MetLife-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Dickson_Prize" title="Dickson Prize">Dickson Prize</a> <span style="font-size:85%;">(1993)</span></li> <li><a href="/wiki/Richard_Lounsbery_Award" title="Richard Lounsbery Award">Richard Lounsbery Award</a> <span style="font-size:85%;">(1993)</span></li> <li><a href="/wiki/Lasker_Award" title="Lasker Award">Lasker Award</a> <span style="font-size:85%;">(1994)</span></li> <li><a href="/wiki/Keio_Medical_Science_Prize" title="Keio Medical Science Prize">Keio Medical Science Prize</a> <span style="font-size:85%;">(1996)</span></li> <li><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> <span style="font-size:85%;">(1997)</span></li> <li><a href="/wiki/Fellow_of_the_Royal_Society" title="Fellow of the Royal Society">ForMemRS</a> <span style="font-size:85%;">(1997)</span><sup id="cite_ref-frs_2-0" class="reference"><a href="#cite_note-frs-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Sir_Hans_Krebs_Medal" title="Sir Hans Krebs Medal">Sir Hans Krebs Medal</a> <span style="font-size:85%;">(1999)</span></li></ul> </div></td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"><div class="plainlist"> <ul><li><a href="/wiki/Neurology" title="Neurology">Neurology</a></li> <li><a href="/wiki/Infectious_disease" class="mw-redirect" title="Infectious disease">Infectious diseases</a></li></ul> </div></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"><div class="plainlist"> <ul><li><a href="/wiki/University_of_California,_Berkeley" title="University of California, Berkeley">University of California, Berkeley</a></li> <li><a href="/wiki/University_of_California,_San_Francisco" title="University of California, San Francisco">University of California, San Francisco</a></li></ul> </div></td></tr><tr style="display:none"><td colspan="2"> </td></tr><tr><th scope="row" class="infobox-label">Website</th><td class="infobox-data"><span class="url"><a rel="nofollow" class="external text" href="http://ind.ucsf.edu/ind/aboutus/faculty/prusiners">ind<wbr />.ucsf<wbr />.edu<wbr />/ind<wbr />/aboutus<wbr />/faculty<wbr />/prusiners</a></span></td></tr></tbody></table> <p><b>Stanley Ben Prusiner</b> (born May 28, 1942<sup id="cite_ref-nobelprize_3-2" class="reference"><a href="#cite_note-nobelprize-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup>) is an American <a href="/wiki/Neurology" title="Neurology">neurologist</a> and <a href="/wiki/Biochemist" title="Biochemist">biochemist</a>. He is the director of the Institute for Neurodegenerative Diseases at <a href="/wiki/University_of_California,_San_Francisco" title="University of California, San Francisco">University of California, San Francisco</a> (UCSF).<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Prusiner discovered <a href="/wiki/Prion" title="Prion">prions</a>, a class of <a href="/wiki/Infection" title="Infection">infectious</a> <a href="/wiki/Biological_reproduction" class="mw-redirect" title="Biological reproduction">self-reproducing</a> <a href="/wiki/Pathogen" title="Pathogen">pathogens</a> primarily or solely composed of <a href="/wiki/Protein" title="Protein">protein</a>, a scientific theory considered by many as a <a href="#Prion:_A_heretical_idea">heretical idea</a> when first proposed. He received the <a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award for Basic Medical Research</a> in 1994 and the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> in 1997 for research on prion diseases developed by him and his team of experts (<a href="/wiki/QPNC-PAGE#Persons" title="QPNC-PAGE">D. E. Garfin</a>,<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> D. P. Stites, W. J. Hadlow, C. M. Eklund) beginning in the early 1970s.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Early_life,_career_and_research"><span id="Early_life.2C_career_and_research"></span>Early life, career and research</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Stanley_B._Prusiner&action=edit&section=1" title="Edit section: Early life, career and research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>He was born in <a href="/wiki/Des_Moines,_Iowa" title="Des Moines, Iowa">Des Moines, Iowa</a>, into a Jewish<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> family to Miriam (Spigel) and Lawrence Prusiner, an architect. He spent his childhood in Des Moines and <a href="/wiki/Cincinnati" title="Cincinnati">Cincinnati</a>, Ohio, where he attended <a href="/wiki/Walnut_Hills_High_School" title="Walnut Hills High School">Walnut Hills High School</a>, where he was known as "the little genius" for his groundbreaking work on a repellent for <a href="/wiki/Boxelder_bug" title="Boxelder bug">Boxelder bugs</a>. Prusiner received a Bachelor of Arts <a href="/wiki/Academic_degree" title="Academic degree">degree</a> in <a href="/wiki/Chemistry" title="Chemistry">chemistry</a> from the <a href="/wiki/University_of_Pennsylvania" title="University of Pennsylvania">University of Pennsylvania</a> and later received his <a href="/wiki/Doctor_of_Medicine" title="Doctor of Medicine">M.D.</a> from the <a href="/wiki/University_of_Pennsylvania_School_of_Medicine" class="mw-redirect" title="University of Pennsylvania School of Medicine">University of Pennsylvania School of Medicine</a>.<sup id="cite_ref-nobelprize_3-3" class="reference"><a href="#cite_note-nobelprize-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Prusiner then completed an <a href="/wiki/Internship" title="Internship">internship</a> in medicine at the University of California, San Francisco. Later Prusiner moved to the <a href="/wiki/National_Institutes_of_Health" title="National Institutes of Health">National Institutes of Health</a>, where he studied glutaminases in <i><a href="/wiki/Escherichia_coli" title="Escherichia coli">E. coli</a></i> in the laboratory of Earl Stadtman.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (September 2022)">citation needed</span></a></i>]</sup> </p><p>After three years at NIH, Prusiner returned to UCSF to complete a <a href="/wiki/Residency_(medicine)" title="Residency (medicine)">residency</a> in <a href="/wiki/Neurology" title="Neurology">neurology</a>. Upon completion of the residency in 1974, Prusiner joined the faculty of the UCSF neurology department. Since that time, Prusiner has held various faculty and visiting faculty positions at both UCSF and <a href="/wiki/University_of_California,_Berkeley" title="University of California, Berkeley">UC Berkeley</a>.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (September 2022)">citation needed</span></a></i>]</sup> </p><p>Since 1999, Prusiner has been director of the Institute for Neurodegenerative Diseases research laboratory at UCSF, working on prion diseases, <a href="/wiki/Alzheimer%27s_disease" title="Alzheimer's disease">Alzheimer's disease</a> and <a href="/wiki/Tauopathy" title="Tauopathy">tauopathies</a>.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Prion:_A_heretical_idea">Prion: A heretical idea</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Stanley_B._Prusiner&action=edit&section=2" title="Edit section: Prion: A heretical idea"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In his 1998 PNAS review article on Prions, <a href="/wiki/Prusiner" class="mw-redirect" title="Prusiner">Prusiner</a> wrote:<sup id="cite_ref-prions1998_10-0" class="reference"><a href="#cite_note-prions1998-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> "The idea that scrapie prions were composed of an amyloidogenic protein was truly heretical when it was introduced" (by <a href="/wiki/Tikvah_Alper" title="Tikvah Alper">Tikvah Alper</a><sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup>). Encephalopathy was a mysterious disease that attacks the brain, and leaves the brains of its victims full of holes. Scientists did not know what pathogen or disease-causing organism that produced such pattern. Prusiner and his co-workers suggested "One scientific theory, viewed as heretical in that it seems to challenge the role of nucleic acids as the exclusive carriers of genetic information." This theory suggested that this pathogen might be a "deadly variety of a normal protein that has the ability to amplify itself in the brain. The hypothetical protein is called a prion (pronounced PREE-on)."<sup id="cite_ref-prions1998_10-1" class="reference"><a href="#cite_note-prions1998-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Awards_and_honors">Awards and honors</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Stanley_B._Prusiner&action=edit&section=3" title="Edit section: Awards and honors"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Stanley Prusiner was awarded the Nobel Prize in Physiology or Medicine in 1997 for his work in proposing an explanation for the cause of <a href="/wiki/Bovine_spongiform_encephalopathy" title="Bovine spongiform encephalopathy">bovine spongiform encephalopathy</a> ("mad cow disease") and its human equivalent, <a href="/wiki/Creutzfeldt%E2%80%93Jakob_disease" title="Creutzfeldt–Jakob disease">Creutzfeldt–Jakob disease</a>.<sup id="cite_ref-nobelprize_3-4" class="reference"><a href="#cite_note-nobelprize-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> In this work, he coined the term <i>prion</i>, which comes from the words "proteinaceous" and "infectious," in 1982 to refer to a previously undescribed form of infection due to protein misfolding.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p><p>Prusiner was elected to the <a href="/wiki/United_States_National_Academy_of_Sciences" class="mw-redirect" title="United States National Academy of Sciences">National Academy of Science</a> in 1992 and to its governing council in 2007.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> He is also an elected member of <a href="/wiki/The_American_Academy_of_Arts_and_Sciences" class="mw-redirect" title="The American Academy of Arts and Sciences">the American Academy of Arts and Sciences</a> (1993),<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> a <a href="/wiki/List_of_Fellows_of_the_Royal_Society_elected_in_1997" class="mw-redirect" title="List of Fellows of the Royal Society elected in 1997">Foreign Member of the Royal Society (ForMemRS) in 1997</a>,<sup id="cite_ref-frs_2-1" class="reference"><a href="#cite_note-frs-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-frs2_18-0" class="reference"><a href="#cite_note-frs2-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> and the <a href="/wiki/American_Philosophical_Society" title="American Philosophical Society">American Philosophical Society</a> (1998),<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> the <a href="/wiki/Serbian_Academy_of_Sciences_and_Arts" title="Serbian Academy of Sciences and Arts">Serbian Academy of Sciences and Arts</a> (2003), and the <a href="/wiki/Institute_of_Medicine" class="mw-redirect" title="Institute of Medicine">Institute of Medicine</a>. </p> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col" style="column-width: 35em;"> <ul><li><a href="/wiki/Potamkin_Prize" title="Potamkin Prize">Potamkin Prize</a> for Alzheimer's Disease Research from the American Academy of Neurology (1991)</li> <li>The <a href="/wiki/Richard_Lounsbery_Award" title="Richard Lounsbery Award">Richard Lounsbery Award</a> for Extraordinary Scientific Research in Biology and Medicine from the National Academy of Sciences (1993)</li> <li><a href="/wiki/Dickson_Prize" title="Dickson Prize">Dickson Prize</a> (1993)</li> <li>The <a href="/wiki/Gairdner_Foundation_International_Award" class="mw-redirect" title="Gairdner Foundation International Award">Gairdner Foundation International Award</a> (1993)</li> <li>The <a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award for Basic Medical Research</a> (1994)</li> <li>The <a href="/wiki/Paul_Ehrlich_and_Ludwig_Darmstaedter_Prize" title="Paul Ehrlich and Ludwig Darmstaedter Prize">Paul Ehrlich and Ludwig Darmstaedter Prize</a> from the Federal Republic of Germany (1995)</li> <li>The <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a> from the <a href="/wiki/Wolf_Foundation" title="Wolf Foundation">Wolf Foundation</a>/<a href="/wiki/Israel" title="Israel">State of Israel</a> (1996)</li> <li><a href="/wiki/Grand_Prix_Charles-Leopold_Mayer" title="Grand Prix Charles-Leopold Mayer">Grand Prix Charles-Leopold Mayer</a> (1996)</li> <li>The <a href="/wiki/Keio_Medical_Science_Prize" title="Keio Medical Science Prize">Keio International Award for Medical Science</a> (1996)</li> <li>Golden Plate Award of the <a href="/wiki/Academy_of_Achievement" title="Academy of Achievement">American Academy of Achievement</a> (1996)<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup></li> <li>The <a href="/wiki/Louisa_Gross_Horwitz_Prize" title="Louisa Gross Horwitz Prize">Louisa Gross Horwitz Prize</a> from <a href="/wiki/Columbia_University" title="Columbia University">Columbia University</a> (1997)</li> <li>The <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> (1997)<sup id="cite_ref-nobelprize_3-5" class="reference"><a href="#cite_note-nobelprize-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></li> <li>The <a href="/wiki/Benjamin_Franklin_Medal_(Franklin_Institute)" class="mw-redirect" title="Benjamin Franklin Medal (Franklin Institute)">Benjamin Franklin Medal</a> from the <a href="/wiki/Franklin_Institute" title="Franklin Institute">Franklin Institute</a> (1998)<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup></li> <li>Honorary Doctorate from <a href="/wiki/CEU_Cardinal_Herrera_University" title="CEU Cardinal Herrera University">CEU Cardinal Herrera University</a> (2005)</li> <li>The <a href="/wiki/National_Medal_of_Science" title="National Medal of Science">National Medal of Science</a> (2010)<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup></li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Stanley_B._Prusiner&action=edit&section=4" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Frank_Bastian" class="mw-redirect" title="Frank Bastian">Frank Bastian</a></li> <li><a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Carleton Gajdusek</a></li> <li><a href="/wiki/Laura_Manuelidis" title="Laura Manuelidis">Laura Manuelidis</a></li> <li><a href="/wiki/List_of_Jewish_Nobel_laureates" title="List of Jewish Nobel laureates">List of Jewish Nobel laureates</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Stanley_B._Prusiner&action=edit&section=5" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><div class="side-box metadata side-box-right"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Scholia_logo.svg/40px-Scholia_logo.svg.png" decoding="async" width="40" height="39" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Scholia_logo.svg/60px-Scholia_logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Scholia_logo.svg/80px-Scholia_logo.svg.png 2x" data-file-width="107" data-file-height="104" /></span></span></div> <div class="side-box-text plainlist"><a href="https://www.wikidata.org/wiki/Wikidata:Scholia" class="extiw" title="d:Wikidata:Scholia">Scholia</a> has an <i>author</i> profile for <i><b><a href="https://iw.toolforge.org/scholia/author/Q155380" class="extiw" title="toolforge:scholia/author/Q155380">Stanley B. Prusiner</a></b></i>.</div></div> </div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 35em;"> <ol class="references"> <li id="cite_note-MetLife-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-MetLife_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20181013093256/http://mlfawards.afar.org/docs/2016Edition_MetLifeFoundationAwards_PastWinners.pdf">"MetLife Foundation Awards for Medical Research in Alzheimer's Disease"</a> <span class="cs1-format">(PDF)</span>. Archived from <a rel="nofollow" class="external text" href="http://mlfawards.afar.org/docs/2016Edition_MetLifeFoundationAwards_PastWinners.pdf">the original</a> <span class="cs1-format">(PDF)</span> on October 13, 2018.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=MetLife+Foundation+Awards+for+Medical+Research+in+Alzheimer%27s+Disease&rft_id=http%3A%2F%2Fmlfawards.afar.org%2Fdocs%2F2016Edition_MetLifeFoundationAwards_PastWinners.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3AStanley+B.+Prusiner" class="Z3988"></span></span> </li> <li id="cite_note-frs-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-frs_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-frs_2-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20150316060617/https://royalsociety.org/about-us/fellowship/fellows/">"Fellows of the Royal Society"</a>. London: <a href="/wiki/Royal_Society" title="Royal Society">Royal Society</a>. Archived from <a rel="nofollow" class="external text" href="https://royalsociety.org/about-us/fellowship/fellows/">the original</a> on March 16, 2015.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Fellows+of+the+Royal+Society&rft.place=London&rft.pub=Royal+Society&rft_id=https%3A%2F%2Froyalsociety.org%2Fabout-us%2Ffellowship%2Ffellows%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AStanley+B.+Prusiner" class="Z3988"></span></span> </li> <li id="cite_note-nobelprize-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-nobelprize_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-nobelprize_3-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-nobelprize_3-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-nobelprize_3-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-nobelprize_3-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-nobelprize_3-5"><sup><i><b>f</b></i></sup></a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.nobelprize.org/laureate/457">Stanley B. Prusiner</a> on Nobelprize.org <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q155380#P8024" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.researchgate.net/profile/Stanley-Prusiner">Stanley Prusiner</a> on <a href="/wiki/ResearchGate" title="ResearchGate">ResearchGate</a></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFintschenko,_Y,_Salamanzadeh,_A,_Davalos,_R_(2014)" class="citation web cs1">Fintschenko, Y, Salamanzadeh, A, Davalos, R (2014). <a rel="nofollow" class="external text" href="https://www.americanlaboratory.com/166335-AES-2013-Annual-Meeting-of-the-AES-Electrophoresis-Society\">"AES 2013: Annual Meeting of the AES Electrophoresis Society"</a>. American Laboratory<span class="reference-accessdate">. Retrieved <span class="nowrap">April 18,</span> 2024</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=AES+2013%3A+Annual+Meeting+of+the+AES+Electrophoresis+Society&rft.pub=American+Laboratory&rft.au=Fintschenko%2C+Y%2C+Salamanzadeh%2C+A%2C+Davalos%2C+R+%282014%29&rft_id=https%3A%2F%2Fwww.americanlaboratory.com%2F166335-AES-2013-Annual-Meeting-of-the-AES-Electrophoresis-Society%5C&rfr_id=info%3Asid%2Fen.wikipedia.org%3AStanley+B.+Prusiner" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_web" title="Template:Cite web">cite web</a>}}</code>: CS1 maint: multiple names: authors list (<a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">link</a>) CS1 maint: numeric names: authors list (<a href="/wiki/Category:CS1_maint:_numeric_names:_authors_list" title="Category:CS1 maint: numeric names: authors list">link</a>)</span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPrusiner_S._B.1982" class="citation journal cs1">Prusiner S. B. (1982). "Novel proteinaceous infectious particles cause scrapie". <i>Science</i>. <b>216</b> (4542): 136–144. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1982Sci...216..136P">1982Sci...216..136P</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.6801762">10.1126/science.6801762</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/6801762">6801762</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Novel+proteinaceous+infectious+particles+cause+scrapie&rft.volume=216&rft.issue=4542&rft.pages=136-144&rft.date=1982&rft_id=info%3Apmid%2F6801762&rft_id=info%3Adoi%2F10.1126%2Fscience.6801762&rft_id=info%3Abibcode%2F1982Sci...216..136P&rft.au=Prusiner+S.+B.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AStanley+B.+Prusiner" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPrusiner_S._B.1991" class="citation journal cs1">Prusiner S. B. (1991). "Molecular biology of prion diseases". <i>Science</i>. <b>252</b> (5012): 1515–1522. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1991Sci...252.1515P">1991Sci...252.1515P</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1675487">10.1126/science.1675487</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/1675487">1675487</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:22417182">22417182</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Molecular+biology+of+prion+diseases&rft.volume=252&rft.issue=5012&rft.pages=1515-1522&rft.date=1991&rft_id=info%3Adoi%2F10.1126%2Fscience.1675487&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A22417182%23id-name%3DS2CID&rft_id=info%3Apmid%2F1675487&rft_id=info%3Abibcode%2F1991Sci...252.1515P&rft.au=Prusiner+S.+B.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AStanley+B.+Prusiner" class="Z3988"></span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.jinfo.org/Nobels_Medicine.html">"Jewish Nobel Prize Winners in Medicine"</a>. <i>www.jinfo.org</i><span class="reference-accessdate">. Retrieved <span class="nowrap">March 30,</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=www.jinfo.org&rft.atitle=Jewish+Nobel+Prize+Winners+in+Medicine&rft_id=https%3A%2F%2Fwww.jinfo.org%2FNobels_Medicine.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AStanley+B.+Prusiner" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20080828015629/http://prusinerlab.ucsf.edu/people/director.php">"UCSF – Prusiner Laboratory – Stanley B. Prusiner, M.D."</a> August 28, 2008. Archived from <a rel="nofollow" class="external text" href="http://prusinerlab.ucsf.edu/people/director.php">the original</a> on August 28, 2008<span class="reference-accessdate">. Retrieved <span class="nowrap">May 9,</span> 2018</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=UCSF+%E2%80%93+Prusiner+Laboratory+%E2%80%93+Stanley+B.+Prusiner%2C+M.D.&rft.date=2008-08-28&rft_id=http%3A%2F%2Fprusinerlab.ucsf.edu%2Fpeople%2Fdirector.php&rfr_id=info%3Asid%2Fen.wikipedia.org%3AStanley+B.+Prusiner" class="Z3988"></span></span> </li> <li id="cite_note-prions1998-10"><span class="mw-cite-backlink">^ <a href="#cite_ref-prions1998_10-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-prions1998_10-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFStanley_Prusiner1998" class="citation journal cs1">Stanley Prusiner (November 10, 1998). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC33918">"Prions"</a>. <i>PNAS</i>. <b>95</b> (23): 13363–13383. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1998PNAS...9513363P">1998PNAS...9513363P</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.95.23.13363">10.1073/pnas.95.23.13363</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC33918">33918</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/9811807">9811807</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=PNAS&rft.atitle=Prions&rft.volume=95&rft.issue=23&rft.pages=13363-13383&rft.date=1998-11-10&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC33918%23id-name%3DPMC&rft_id=info%3Apmid%2F9811807&rft_id=info%3Adoi%2F10.1073%2Fpnas.95.23.13363&rft_id=info%3Abibcode%2F1998PNAS...9513363P&rft.au=Stanley+Prusiner&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC33918&rfr_id=info%3Asid%2Fen.wikipedia.org%3AStanley+B.+Prusiner" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text">T. 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Prusiner, M.D."</a> Boston University, Amyloidosis Center<span class="reference-accessdate">. Retrieved <span class="nowrap">August 25,</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Stanley+B.+Prusiner%2C+M.D.&rft.pub=Boston+University%2C+Amyloidosis+Center&rft_id=https%3A%2F%2Fwww.bu.edu%2Famyloid%2Fevents%2Fprusiner-bio%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AStanley+B.+Prusiner" class="Z3988"></span></span> </li> <li id="cite_note-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-15">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation magazine cs1"><a rel="nofollow" class="external text" href="http://blog.wired.com/wiredscience/2007/01/what_really_cau.html">"What really causes mad cow disease?"</a>. <i>Wired</i>. 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Prusiner</a> on Nobelprize.org <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q155380#P8024" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output 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.navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Wolf_Prize_in_Medicine" title="Template:Wolf Prize in Medicine"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Wolf_Prize_in_Medicine" title="Template talk:Wolf Prize in Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Wolf_Prize_in_Medicine" title="Special:EditPage/Template:Wolf Prize in Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Wolf_Prize_in_Medicine" style="font-size:114%;margin:0 4em">Laureates of the <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1970s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/Jon_van_Rood" title="Jon van Rood">Jon J. van Rood</a> (1978)</li> <li><a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Oleh_Hornykiewicz" title="Oleh Hornykiewicz">Oleh Hornykiewicz</a> (1979)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1980s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> / <a href="/wiki/Leo_Sachs" title="Leo Sachs">Leo Sachs</a> / <a href="/wiki/James_Learmonth_Gowans" title="James Learmonth Gowans">James L. Gowans</a> (1980)</li> <li><a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a> / <a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a> (1981)</li> <li><a href="/wiki/Jean-Pierre_Changeux" title="Jean-Pierre Changeux">Jean-Pierre Changeux</a> / <a href="/wiki/Solomon_H._Snyder" title="Solomon H. Snyder">Solomon H. Snyder</a> / <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> (1982)</li> <li><a href="/wiki/Donald_F._Steiner" title="Donald F. Steiner">Donald F. Steiner</a> (1984/5)</li> <li><a href="/wiki/Osamu_Hayaishi" title="Osamu Hayaishi">Osamu Hayaishi</a> (1986)</li> <li><a href="/wiki/Pedro_Cuatrecasas" title="Pedro Cuatrecasas">Pedro Cuatrecasas</a> / <a href="/wiki/Meir_Wilchek" title="Meir Wilchek">Meir Wilchek</a> (1987)</li> <li><a href="/wiki/Henri_G._Hers" title="Henri G. Hers">Henri G. Hers</a> / <a href="/wiki/Elizabeth_F._Neufeld" title="Elizabeth F. Neufeld">Elizabeth F. Neufeld</a> (1988)</li> <li><a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> (1989)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1990s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Maclyn_McCarty" title="Maclyn McCarty">Maclyn McCarty</a> (1990)</li> <li><a href="/wiki/Seymour_Benzer" title="Seymour Benzer">Seymour Benzer</a> (1991)</li> <li><a href="/wiki/Judah_Folkman" title="Judah Folkman">Judah Folkman</a> (1992)</li> <li><a href="/wiki/Michael_Berridge" title="Michael Berridge">Michael Berridge</a> / <a href="/wiki/Yasutomi_Nishizuka" title="Yasutomi Nishizuka">Yasutomi Nishizuka</a> (1994/5)</li> <li><a class="mw-selflink selflink">Stanley B. Prusiner</a> (1995/6)</li> <li><a href="/wiki/Mary_F._Lyon" title="Mary F. Lyon">Mary F. Lyon</a> (1996/7)</li> <li><a href="/wiki/Michael_Sela" title="Michael Sela">Michael Sela</a> / <a href="/wiki/Ruth_Arnon" title="Ruth Arnon">Ruth Arnon</a> (1998)</li> <li><a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a> (1999)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2000s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Avram_Hershko" title="Avram Hershko">Avram Hershko</a> / <a href="/wiki/Alexander_Varshavsky" title="Alexander Varshavsky">Alexander Varshavsky</a> (2001)</li> <li><a href="/wiki/Ralph_L._Brinster" title="Ralph L. Brinster">Ralph L. Brinster</a> / <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a> (2002/3)</li> <li><a href="/wiki/Robert_Weinberg_(biologist)" title="Robert Weinberg (biologist)">Robert Weinberg</a> / <a href="/wiki/Roger_Y._Tsien" title="Roger Y. Tsien">Roger Y. Tsien</a> (2004)</li> <li><a href="/wiki/Alexander_Levitzki" title="Alexander Levitzki">Alexander Levitzki</a> / <a href="/wiki/Anthony_R._Hunter" title="Anthony R. Hunter">Anthony R. Hunter</a> / <a href="/wiki/Tony_Pawson_(biochemist)" title="Tony Pawson (biochemist)">Tony Pawson</a> (2005)</li> <li><a href="/wiki/Howard_Cedar" title="Howard Cedar">Howard Cedar</a> / <a href="/wiki/Aharon_Razin" title="Aharon Razin">Aharon Razin</a> (2008)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2010s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Axel_Ullrich" title="Axel Ullrich">Axel Ullrich</a> (2010)</li> <li><a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a> / <a href="/wiki/Rudolf_Jaenisch" title="Rudolf Jaenisch">Rudolf Jaenisch</a> (2011)</li> <li><a href="/wiki/Ronald_M._Evans" title="Ronald M. Evans">Ronald M. Evans</a> (2012)</li> <li><a href="/wiki/Nahum_Sonenberg" title="Nahum Sonenberg">Nahum Sonenberg</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a> / <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> (2014)</li> <li><a href="/wiki/John_Kappler" title="John Kappler">John Kappler</a> / <a href="/wiki/Philippa_Marrack" title="Philippa Marrack">Philippa Marrack</a> / <a href="/wiki/Jeffrey_V._Ravetch" title="Jeffrey V. Ravetch">Jeffrey V. Ravetch</a> (2015)</li> <li><a href="/wiki/Lewis_C._Cantley" title="Lewis C. Cantley">Lewis C. Cantley</a> / <a href="/wiki/C._Ronald_Kahn" title="C. Ronald Kahn">C. Ronald Kahn</a> (2016)</li> <li><a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> (2017)</li> <li><a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey M. Friedman</a> (2019)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2020s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a> / <a href="/wiki/Jennifer_Doudna" title="Jennifer Doudna">Jennifer Doudna</a> (2020)</li> <li><a href="/wiki/Joan_Steitz" class="mw-redirect" title="Joan Steitz">Joan Steitz</a> / <a href="/wiki/Lynne_Maquat" class="mw-redirect" title="Lynne Maquat">Lynne Elizabeth Maquat</a> / <a href="/wiki/Adrian_R._Krainer" class="mw-redirect" title="Adrian R. Krainer">Adrian Krainer</a> (2021)</li> <li><a href="/wiki/Daniel_J._Drucker" title="Daniel J. Drucker">Daniel J. Drucker</a> (2023)</li> <li><a href="/wiki/Botond_Roska" title="Botond Roska">Botond Roska</a> / <a href="/wiki/Jos%C3%A9-Alain_Sahel" title="José-Alain Sahel">José-Alain Sahel</a> (2024)</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r1239334494">@media screen{html.skin-theme-clientpref-night .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"></div><div role="navigation" class="navbox" aria-labelledby="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Template:Nobel Prize in Physiology or Medicine"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Nobel_Prize_in_Physiology_or_Medicine" title="Template talk:Nobel Prize in Physiology or Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Special:EditPage/Template:Nobel Prize in Physiology or Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="font-size:114%;margin:0 4em"><a href="/wiki/List_of_Nobel_laureates_in_Physiology_or_Medicine" title="List of Nobel laureates in Physiology or Medicine">Laureates</a> of the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Emil_von_Behring" title="Emil von Behring">Emil Behring</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ronald Ross</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Niels Finsen</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Ivan Pavlov</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Camillo Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Santiago Ramón y Cajal</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Alphonse Laveran</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/%C3%89lie_Metchnikoff" title="Élie Metchnikoff">Élie Metchnikoff</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Paul Ehrlich</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Emil Kocher</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Albrecht Kossel</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Allvar Gullstrand</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Alexis Carrel</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Charles Richet</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" class="mw-redirect" title="Róbert Bárány">Róbert Bárány</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1915</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1917</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1918</span></span></li> <li><span class="nowrap">1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Jules Bordet</a></span></li> <li><span class="nowrap">1920: <a href="/wiki/August_Krogh" title="August Krogh">August Krogh</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1921</span></span></li> <li><span class="nowrap">1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Archibald Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Otto Meyerhof</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Frederick Banting</a> / <a href="/wiki/John_Macleod_(physiologist)" title="John Macleod (physiologist)">John Macleod</a></span></li> <li><span class="nowrap">1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Willem Einthoven</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1925</span></span></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/80px-Nobel_Prize.png" decoding="async" width="80" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/120px-Nobel_Prize.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/160px-Nobel_Prize.png 2x" data-file-width="500" data-file-height="492" /></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Johannes Fibiger</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Charles Nicolle</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Christiaan Eijkman</a> / <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Frederick Gowland Hopkins</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Karl Landsteiner</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Otto Warburg</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Charles Scott Sherrington</a> / <a href="/wiki/Edgar_Adrian" title="Edgar Adrian">Edgar Adrian</a></span></li> <li><span class="nowrap">1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Morgan</a></span></li> <li><span class="nowrap">1934: <a href="/wiki/George_Whipple" title="George Whipple">George Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">George Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">William Murphy</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Hans Spemann</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Otto Loewi</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Albert Szent-Györgyi</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Corneille Heymans</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Gerhard Domagk</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Henrik Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Edward Doisy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Joseph Erlanger</a> / <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Herbert Gasser</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Alexander Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Ernst Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Howard Florey</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Hermann Muller</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">Carl Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">Gerty Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Bernardo Houssay</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Paul Müller</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Walter Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">António Egas Moniz</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Edward Kendall</a> / <a href="/wiki/Tadeusz_Reichstein" title="Tadeusz Reichstein">Tadeusz Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Philip Hench</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Max Theiler</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Selman Waksman</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hans_Krebs_(biochemist)" title="Hans Krebs (biochemist)">Hans Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Lipmann</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Thomas Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Frederick Robbins</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Hugo Theorell</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">André Cournand</a> / <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Werner Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Dickinson W. Richards</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Daniel Bovet</a></span></li> <li><span class="nowrap">1958: <a href="/wiki/George_Beadle" title="George Beadle">George Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Macfarlane_Burnet" title="Macfarlane Burnet">Frank Burnet</a> / <a href="/wiki/Peter_Medawar" title="Peter Medawar">Peter Medawar</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Georg von Békésy</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> / <a href="/wiki/James_Watson" title="James Watson">James Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Maurice Wilkins</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">John Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Alan Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Andrew Huxley</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Konrad Bloch</a> / <a href="/wiki/Feodor_Lynen" title="Feodor Lynen">Feodor Lynen</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">François Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" title="André Michel Lwoff">André Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Jacques Monod</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Charles B. Huggins</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Ragnar Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Haldan Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">George Wald</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Robert W. Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall Nirenberg</a></span></li> <li><span class="nowrap">1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Max Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Bernard Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf von Euler</a> / <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Julius Axelrod</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Earl Sutherland Jr.</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Gerald Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" class="mw-redirect" title="Rodney Robert Porter">Rodney Porter</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> / <a href="/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen">Nikolaas Tinbergen</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Albert Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Palade</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Renato Dulbecco</a> / <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Baruch Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Andrew Schally</a> / <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Yalow</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> / <a href="/wiki/Hamilton_O._Smith" title="Hamilton O. Smith">Hamilton O. Smith</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Allan_MacLeod_Cormack" title="Allan MacLeod Cormack">Allan Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Godfrey Hounsfield</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/David_H._Hubel" title="David H. Hubel">David H. Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Torsten Wiesel</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Sune Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" title="Bengt I. Samuelsson">Bengt I. Samuelsson</a> / <a href="/wiki/John_Vane" title="John Vane">John Vane</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Georges Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Michael Brown</a> / <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Joseph L. Goldstein</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Stanley_Cohen_(biochemist)" title="Stanley Cohen (biochemist)">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> / <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Gertrude B. Elion</a> / <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">George H. Hitchings</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">J. Michael Bishop</a> / <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Harold E. Varmus</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Joseph_Murray" class="mw-redirect" title="Joseph Murray">Joseph Murray</a> / <a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">E. Donnall Thomas</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Edmond Fischer</a> / <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">Edwin G. Krebs</a></span></li> <li><span class="nowrap">1993: <a href="/wiki/Richard_J._Roberts" title="Richard J. Roberts">Richard J. Roberts</a> / <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Sharp</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Alfred G. Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Martin Rodbell</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Christiane Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Eric F. Wieschaus</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Peter_C._Doherty" class="mw-redirect" title="Peter C. Doherty">Peter C. Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Rolf M. Zinkernagel</a></span></li> <li><span class="nowrap">1997: <a class="mw-selflink selflink">Stanley B. Prusiner</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Robert F. Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Louis Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Paul Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Leland H. Hartwell</a> / <a href="/wiki/Tim_Hunt" title="Tim Hunt">Tim Hunt</a> / <a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> / <a href="/wiki/John_Sulston" title="John Sulston">John E. Sulston</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Barry Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Robin Warren</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Andrew Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Craig Mello</a></span></li> <li><span class="nowrap">2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> / <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> / <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Carol W. Greider</a> / <a href="/wiki/Jack_W._Szostak" title="Jack W. Szostak">Jack W. Szostak</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Robert_Edwards_(physiologist)" title="Robert Edwards (physiologist)">Robert G. Edwards</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> / <a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules A. Hoffmann</a> / <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Ralph M. Steinman</a> (<i>posthumously</i>)</span></li> <li><span class="nowrap">2012: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/James_Rothman" title="James Rothman">James Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Randy Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Thomas C. Südhof</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/John_O%27Keefe_(neuroscientist)" title="John O'Keefe (neuroscientist)">John O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">May-Britt Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">Edvard Moser</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/William_C._Campbell_(scientist)" title="William C. Campbell (scientist)">William C. Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Tu Youyou</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Gregg L. Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">William Kaelin Jr.</a></span></li> <li><span class="nowrap"><a href="/wiki/2020_Nobel_Prize_in_Physiology_or_Medicine" title="2020 Nobel Prize in Physiology or Medicine">2020</a>: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Harvey J. Alter</a> / <a href="/wiki/Michael_Houghton_(virologist)" class="mw-redirect" title="Michael Houghton (virologist)">Michael Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Charles M. Rice</a></span></li> <li><span class="nowrap"><a href="/wiki/2021_Nobel_Prize_in_Physiology_or_Medicine" title="2021 Nobel Prize in Physiology or Medicine">2021</a>: <a href="/wiki/David_Julius" title="David Julius">David Julius</a> / <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Ardem Patapoutian</a></span></li> <li><span class="nowrap"><a href="/wiki/2022_Nobel_Prize_in_Physiology_or_Medicine" title="2022 Nobel Prize in Physiology or Medicine">2022</a>: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a></span></li> <li><span class="nowrap"><a href="/wiki/2023_Nobel_Prize_in_Physiology_or_Medicine" title="2023 Nobel Prize in Physiology or Medicine">2023</a>: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style></div><div role="navigation" class="navbox" aria-labelledby="1997_Nobel_Prize_laureates" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:1997_Nobel_Prize_winners" title="Template:1997 Nobel Prize winners"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:1997_Nobel_Prize_winners" title="Template talk:1997 Nobel Prize winners"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:1997_Nobel_Prize_winners" title="Special:EditPage/Template:1997 Nobel Prize winners"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="1997_Nobel_Prize_laureates" style="font-size:114%;margin:0 4em">1997 <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Paul_D._Boyer" title="Paul D. Boyer">Paul D. Boyer</a> (United States)</li> <li><a href="/wiki/John_E._Walker" title="John E. Walker">John E. Walker</a> (United Kingdom)</li> <li><a href="/wiki/Jens_Christian_Skou" title="Jens Christian Skou">Jens C. Skou</a> (Denmark)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a> (<a href="/wiki/1997_Nobel_Prize_in_Literature" title="1997 Nobel Prize in Literature">1997</a>)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><a href="/wiki/Dario_Fo" title="Dario Fo">Dario Fo</a> (Italy)</div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Peace_Prize" title="Nobel Peace Prize">Peace</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/International_Campaign_to_Ban_Landmines" title="International Campaign to Ban Landmines">International Campaign to Ban Landmines</a> (Switzerland)</li> <li><a href="/wiki/Jody_Williams" title="Jody Williams">Jody Williams</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Steven_Chu" title="Steven Chu">Steven Chu</a> (United States)</li> <li><a href="/wiki/Claude_Cohen-Tannoudji" title="Claude Cohen-Tannoudji">Claude Cohen-Tannoudji</a> (France)</li> <li><a href="/wiki/William_Daniel_Phillips" title="William Daniel Phillips">William Daniel Phillips</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><a class="mw-selflink selflink">Stanley B. Prusiner</a> (United States)</div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economic Sciences</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Robert_C._Merton" title="Robert C. Merton">Robert C. Merton</a> (United States)</li> <li><a href="/wiki/Myron_Scholes" title="Myron Scholes">Myron Scholes</a> (Canada/United States)</li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <dl><dt><span class="nobold"><a href="/wiki/List_of_Nobel_laureates" title="List of Nobel laureates">Nobel Prize recipients</a></span></dt> <dd><a href="/wiki/Template:1992_Nobel_Prize_winners" title="Template:1992 Nobel Prize winners">1992</a></dd> <dd><a href="/wiki/Template:1993_Nobel_Prize_winners" title="Template:1993 Nobel Prize winners">1993</a></dd> <dd><a href="/wiki/Template:1994_Nobel_Prize_winners" title="Template:1994 Nobel Prize winners">1994</a></dd> <dd><a href="/wiki/Template:1995_Nobel_Prize_winners" title="Template:1995 Nobel Prize winners">1995</a></dd> <dd><a href="/wiki/Template:1996_Nobel_Prize_winners" title="Template:1996 Nobel Prize winners">1996</a></dd> <dd><b>1997 </b></dd> <dd><a href="/wiki/Template:1998_Nobel_Prize_winners" title="Template:1998 Nobel Prize winners">1998</a></dd> <dd><a href="/wiki/Template:1999_Nobel_Prize_winners" title="Template:1999 Nobel Prize winners">1999</a></dd> <dd><a href="/wiki/Template:2000_Nobel_Prize_winners" title="Template:2000 Nobel Prize winners">2000</a></dd> <dd><a href="/wiki/Template:2001_Nobel_Prize_winners" title="Template:2001 Nobel Prize winners">2001</a></dd> <dd><a href="/wiki/Template:2002_Nobel_Prize_winners" title="Template:2002 Nobel Prize winners">2002</a></dd></dl> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Fellows_of_the_Royal_Society_elected_in_1997" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:FRS_1997" title="Template:FRS 1997"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:FRS_1997" title="Template talk:FRS 1997"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:FRS_1997" title="Special:EditPage/Template:FRS 1997"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Fellows_of_the_Royal_Society_elected_in_1997" style="font-size:114%;margin:0 4em"><a href="/wiki/Fellow_of_the_Royal_Society" title="Fellow of the Royal Society">Fellows</a> of the <a href="/wiki/Royal_Society" title="Royal Society">Royal Society</a> elected <a href="/wiki/List_of_Fellows_of_the_Royal_Society_elected_in_1997" class="mw-redirect" title="List of Fellows of the Royal Society elected in 1997">in 1997</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Fellows</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mike_Bate" title="Mike Bate">Mike Bate</a></li> <li><a href="/wiki/J._Michael_Brady" class="mw-redirect" title="J. Michael Brady">J. Michael Brady</a></li> <li><a href="/wiki/Michael_Bulmer" title="Michael Bulmer">Michael Bulmer</a></li> <li><a href="/wiki/John_Burland" title="John Burland">John Burland</a></li> <li><a href="/wiki/Richard_Dickinson_Chambers" title="Richard Dickinson Chambers">Richard Dickinson Chambers</a></li> <li><a href="/wiki/Colin_W._Clark" title="Colin W. Clark">Colin W. Clark</a></li> <li><a href="/wiki/David_Clary" title="David Clary">David Clary</a></li> <li><a href="/wiki/Laurence_Eaves" title="Laurence Eaves">Laurence Eaves</a></li> <li><a href="/wiki/Richard_Fortey" title="Richard Fortey">Richard Fortey</a></li> <li><a href="/wiki/David_Garner_(chemist)" title="David Garner (chemist)">David Garner</a></li> <li><a href="/wiki/Douglas_Gough" title="Douglas Gough">Douglas Gough</a></li> <li><a href="/wiki/John_Hinch_(mathematician)" title="John Hinch (mathematician)">John Hinch</a></li> <li><a href="/wiki/Julian_Jack" title="Julian Jack">Julian Jack</a></li> <li><a href="/wiki/Paul_Gordon_Jarvis" title="Paul Gordon Jarvis">Paul Gordon Jarvis</a></li> <li><a href="/wiki/Charles_K._Kao" title="Charles K. Kao">Charles K. Kao</a></li> <li><a href="/wiki/Barry_Keverne" title="Barry Keverne">Barry Keverne</a></li> <li><a href="/wiki/Philip_Kocienski" title="Philip Kocienski">Philip Kocienski</a></li> <li><a href="/wiki/Peter_B._Kronheimer" title="Peter B. Kronheimer">Peter B. Kronheimer</a></li> <li><a href="/wiki/Philippa_Marrack" title="Philippa Marrack">Philippa Marrack</a></li> <li><i><a href="https://www.wikidata.org/wiki/Q21165382" class="extiw" title="d:Q21165382">James R. Maxwell</a></i></li> <li><a href="/wiki/Tim_Mitchison" title="Tim Mitchison">Tim Mitchison</a></li> <li><a href="/wiki/Richard_G._Morris" title="Richard G. Morris">Richard G. Morris</a></li> <li><a href="/wiki/Chris_Perrins" title="Chris Perrins">Chris Perrins</a></li> <li><a href="/wiki/George_Pickett_(physicist)" title="George Pickett (physicist)">George Pickett</a></li> <li><a href="/wiki/George_Poste" title="George Poste">George Poste</a></li> <li><i><a href="https://www.wikidata.org/wiki/Q21166479" class="extiw" title="d:Q21166479">Kenneth Reid</a></i></li> <li><i><a href="https://www.wikidata.org/wiki/Q21165006" class="extiw" title="d:Q21165006">Alan Rickinson</a></i></li> <li><a href="/wiki/Leo_Sachs" title="Leo Sachs">Leo Sachs</a></li> <li><a href="/wiki/Giacinto_Scoles" title="Giacinto Scoles">Giacinto Scoles</a></li> <li><a href="/wiki/James_Scott_(cardiologist)" title="James Scott (cardiologist)">James Scott</a></li> <li><a href="/wiki/Wilson_Sibbett" title="Wilson Sibbett">Wilson Sibbett</a></li> <li><a href="/wiki/Bernard_Silverman" title="Bernard Silverman">Bernard Silverman</a></li> <li><a href="/wiki/Richard_Sykes_(biochemist)" class="mw-redirect" title="Richard Sykes (biochemist)">Richard Sykes</a></li> <li><a href="/wiki/Richard_E._Taylor" title="Richard E. Taylor">Richard E. Taylor</a></li> <li><a href="/wiki/Neil_Trudinger" title="Neil Trudinger">Neil Trudinger</a></li> <li><a href="/wiki/Robin_Weiss" title="Robin Weiss">Robin Weiss</a></li> <li><a href="/wiki/Simon_White" title="Simon White">Simon White</a></li> <li><a href="/wiki/Alan_Windle" title="Alan Windle">Alan Windle</a></li> <li><a href="/wiki/Richard_D._Wood" class="mw-redirect" title="Richard D. Wood">Richard D. Wood</a></li> <li><a href="/wiki/Graham_Charles_Wood" title="Graham Charles Wood">Graham C. Wood</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Foreign</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Thomas_Eisner" title="Thomas Eisner">Thomas Eisner</a></li> <li><a href="/wiki/Walter_Jakob_Gehring" title="Walter Jakob Gehring">Walter Jakob Gehring</a></li> <li><a href="/wiki/Roy_J._Glauber" title="Roy J. Glauber">Roy J. Glauber</a></li> <li><a href="/wiki/Martin_David_Kruskal" title="Martin David Kruskal">Martin David Kruskal</a></li> <li><a href="/wiki/George_Andrew_Olah" title="George Andrew Olah">George Andrew Olah</a></li> <li><a class="mw-selflink selflink">Stanley B. Prusiner</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-labelledby="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q155380#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q155380#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a> <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q155380#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">International</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://isni.org/isni/0000000081802961">ISNI</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/112781106">VIAF</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://id.worldcat.org/fast/42373/">FAST</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.oclc.org/worldcat/entity/E39PBJqfCbvb9g8vJFhp8cvJXd">WorldCat</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/14029905X">Germany</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/n79103845">United States</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb134865075">France</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb134865075">BnF data</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=mub2013787423&CON_LNG=ENG">Czech Republic</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p068197888">Netherlands</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://dbn.bn.org.pl/descriptor-details/9810651893105606">Poland</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://olduli.nli.org.il/F/?func=find-b&local_base=NLX10&find_code=UID&request=987007274386005171">Israel</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://opac.kbr.be/LIBRARY/doc/AUTHORITY/14378459">Belgium</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Academics</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://orcid.org/0000-0003-1955-5498">ORCID</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.scopus.com/authid/detail.uri?authorId=56822190400">Scopus</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">People</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.deutsche-biographie.de/pnd14029905X.html?language=en">Deutsche Biographie</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/076631087">IdRef</a></span></li></ul></div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐f69cdc8f6‐xz5ct Cached time: 20241122143927 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 0.641 seconds Real time usage: 0.810 seconds Preprocessor visited node count: 4493/1000000 Post‐expand include size: 145497/2097152 bytes Template argument size: 10140/2097152 bytes Highest expansion depth: 12/100 Expensive parser function count: 8/500 Unstrip 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