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Available in 24 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-24" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">24 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B7%D8%AD%D8%A7%D9%84%D8%A8_%D9%85%D8%AA%D8%B1%D8%A7%D8%A8%D8%B7%D8%A9" title="طحالب مترابطة – Arabic" lang="ar" hreflang="ar" data-title="طحالب مترابطة" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Zygnematophyceae" title="Zygnematophyceae – Azerbaijani" lang="az" hreflang="az" data-title="Zygnematophyceae" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Zignematof%C3%ADcies" title="Zignematofícies – Catalan" lang="ca" hreflang="ca" data-title="Zignematofícies" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Sp%C3%A1jivky" title="Spájivky – Czech" lang="cs" hreflang="cs" data-title="Spájivky" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Schmuckalgen" title="Schmuckalgen – German" lang="de" hreflang="de" data-title="Schmuckalgen" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Ikkesvetikad" title="Ikkesvetikad – Estonian" lang="et" hreflang="et" data-title="Ikkesvetikad" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Zygnematophyceae" title="Zygnematophyceae – Spanish" lang="es" hreflang="es" data-title="Zygnematophyceae" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DB%8C%D9%88%D8%BA%DB%8C%E2%80%8C%D8%AC%D9%84%D8%A8%DA%A9%D8%A7%D9%86" title="یوغی‌جلبکان – Persian" lang="fa" hreflang="fa" data-title="یوغی‌جلبکان" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Zygnematophyceae" title="Zygnematophyceae – French" lang="fr" hreflang="fr" data-title="Zygnematophyceae" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%A0%91%ED%95%A9%EC%A1%B0%EA%B0%95" title="접합조강 – Korean" lang="ko" hreflang="ko" data-title="접합조강" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Zygnematophyceae" title="Zygnematophyceae – Indonesian" lang="id" hreflang="id" data-title="Zygnematophyceae" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Zygnematophyceae" title="Zygnematophyceae – Hungarian" lang="hu" hreflang="hu" data-title="Zygnematophyceae" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%B7%D8%AD%D8%A7%D9%84%D8%A8_%D9%85%D8%AA%D8%B1%D8%A7%D8%A8%D8%B7%D9%87" title="طحالب مترابطه – Egyptian Arabic" lang="arz" hreflang="arz" data-title="طحالب مترابطه" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E6%8E%A5%E5%90%88%E8%97%BB" title="接合藻 – Japanese" lang="ja" hreflang="ja" data-title="接合藻" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Sprz%C4%99%C5%BCnice" title="Sprzężnice – Polish" lang="pl" hreflang="pl" data-title="Sprzężnice" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Zygnematophyceae" title="Zygnematophyceae – Portuguese" lang="pt" hreflang="pt" data-title="Zygnematophyceae" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9A%D0%BE%D0%BD%D1%8A%D1%8E%D0%B3%D0%B0%D1%82%D1%8B" title="Конъюгаты – Russian" lang="ru" hreflang="ru" data-title="Конъюгаты" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Zygnematophyceae" title="Zygnematophyceae – Simple English" lang="en-simple" hreflang="en-simple" data-title="Zygnematophyceae" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Sp%C3%A1javky" title="Spájavky – Slovak" lang="sk" hreflang="sk" data-title="Spájavky" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Konjugatalger" title="Konjugatalger – Swedish" lang="sv" hreflang="sv" data-title="Konjugatalger" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%97%D0%B8%D0%B3%D0%BD%D0%B5%D0%BC%D0%B0%D1%82%D0%BE%D1%84%D1%96%D1%86%D1%96%D1%94%D0%B2%D1%96" title="Зигнематофіцієві – Ukrainian" lang="uk" hreflang="uk" data-title="Зигнематофіцієві" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/L%E1%BB%9Bp_Song_tinh_t%E1%BA%A3o" title="Lớp Song tinh tảo – Vietnamese" lang="vi" hreflang="vi" data-title="Lớp Song tinh tảo" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E6%8E%A5%E5%90%88%E8%97%BB%E7%B6%B1" title="接合藻綱 – Cantonese" lang="yue" hreflang="yue" data-title="接合藻綱" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%8F%8C%E6%98%9F%E8%97%BB%E7%BA%B2" title="双星藻纲 – Chinese" lang="zh" hreflang="zh" data-title="双星藻纲" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet 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<div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Class of algae</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(180,250,180)">Zygnematophyceae </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Spirogyra_3.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Spirogyra_3.jpg/220px-Spirogyra_3.jpg" decoding="async" width="220" height="232" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Spirogyra_3.jpg/330px-Spirogyra_3.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Spirogyra_3.jpg/440px-Spirogyra_3.jpg 2x" data-file-width="1135" data-file-height="1197" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Conjugation in <i><a href="/wiki/Spirogyra" title="Spirogyra">Spirogyra</a></i>. </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(180,250,180)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Zygnematophyceae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Viridiplantae" title="Viridiplantae">Viridiplantae</a> </td></tr> <tr> <td>(unranked): </td> <td><a href="/wiki/Charophyta" title="Charophyta">Charophyta</a> </td></tr> <tr> <td>Class: </td> <td><a class="mw-selflink selflink">Zygnematophyceae</a><br /><small>Round, 1971</small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(180,250,180)">Subclasses </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li>Subclass Zygnematophycidae <ul><li>Order <a href="/wiki/Zygnematales" title="Zygnematales">Zygnematales</a></li> <li>Order <a href="/wiki/Desmidiales" title="Desmidiales">Desmidiales</a></li> <li>Order <a href="/w/index.php?title=Spirogyrales&action=edit&redlink=1" class="new" title="Spirogyrales (page does not exist)">Spirogyrales</a></li> <li>Order <a href="/w/index.php?title=Serritaeniales&action=edit&redlink=1" class="new" title="Serritaeniales (page does not exist)">Serritaeniales</a></li></ul></li> <li>Subclass Spirogloeophycidae <ul><li>Order <a href="/w/index.php?title=Spirogloeales&action=edit&redlink=1" class="new" title="Spirogloeales (page does not exist)">Spirogloeales</a></li></ul></li></ul> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(180,250,180)"><a href="/wiki/Synonym_(taxonomy)" title="Synonym (taxonomy)">Synonyms</a><sup id="cite_ref-Guiry_1-0" class="reference"><a href="#cite_note-Guiry-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li>Akontae <small>Blackman & Tansley, 1902</small></li> <li>Conjugatophyceae <small>Engler, 1892</small></li> <li>'Conjugatae' <small>Engler, 1892</small></li> <li>Gamophyceae</li> <li>Saccodermae <small>Pascher ex Kossinskaja, 1952</small></li> <li>Zygnemaphyceae <small>Round 1971</small></li> <li>Zygophyceae <small>Widder, 1960</small></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><b>Zygnematophyceae</b> (or <b>Conjugatophyceae</b>) is a class of <a href="/wiki/Green_algae" title="Green algae">green algae</a> in the paraphylum streptophyte algae, also referred to as <a href="/wiki/Charophyta" title="Charophyta">Charophyta</a>,<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> consisting of more than 4000 described species.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> The Zygnematophyceae are the sister clade of the <a href="/wiki/Embryophyta" class="mw-redirect" title="Embryophyta">Embryophyta</a> (land plants).<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>Common members of the Zygnematophyceae include the filamentous algae <i><a href="/wiki/Spirogyra" title="Spirogyra">Spirogyra</a></i> and <i><a href="/wiki/Mougeotia" title="Mougeotia">Mougeotia</a></i>, as well as <a href="/wiki/Desmid" class="mw-redirect" title="Desmid">desmids</a>, which are microscopic algae characterized by symmetrical and elaborately ornate cells.<sup id="cite_ref-Hall_7-0" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Morphology">Morphology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Zygnematophyceae&action=edit&section=1" title="Edit section: Morphology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The body plan of Zygnematophyceae is simple, and the group appears to have gone through a secondary loss of morphological complexity.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> The most basal members are unicellular, but filamentous species have evolved at least five times,<sup id="cite_ref-Hess2022_9-0" class="reference"><a href="#cite_note-Hess2022-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> and a few species form colonies.<sup id="cite_ref-Hall_7-1" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> Each cell contains a single <a href="/wiki/Cell_nucleus" title="Cell nucleus">nucleus</a>.<sup id="cite_ref-Hall_7-2" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p>The <a href="/wiki/Chloroplast" title="Chloroplast">chloroplasts</a> of the Zygnematophyceae are large, typically axile but sometimes parietally located, and contain one or several <a href="/wiki/Pyrenoid" title="Pyrenoid">pyrenoids</a>. One or several chloroplasts may be present in a single cell. The shape of the chloroplast may be star-shaped (in <i><a href="/wiki/Zygnema" title="Zygnema">Zygnema</a></i>), ribbon-shaped (in <i><a href="/wiki/Spirogyra" title="Spirogyra">Spirogyra</a></i>), or elaborately lobed and dissected. In some taxa, particularly <i><a href="/wiki/Mougeotia" title="Mougeotia">Mougeotia</a></i>, the chloroplast is able to move in response to different light conditions.<sup id="cite_ref-Hall_7-3" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p>In the Zygnematophyceae, cell walls are composed of three layers: one outer layer consisting of mucus, a primary wall consisting of <a href="/wiki/Microfibril" title="Microfibril">microfibrils</a>, and an innermost layer of cellulosic <a href="/wiki/Microfibrils" class="mw-redirect" title="Microfibrils">microfibrils</a>. Some species shed their primary wall and retain only the innermost layer. The cell wall may be variously decorated with striations, granules, or spines. In the Desmidiales, there are pores in the cell wall which allow the cell to extrude a layer of <a href="/wiki/Mucilage" title="Mucilage">mucilage</a> for protection.<sup id="cite_ref-Hall_7-4" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p>Several Zygnematophyceae contain genes involved in protection from desiccation that appear to have been derived by <a href="/wiki/Horizontal_gene_transfer" title="Horizontal gene transfer">horizontal gene transfer</a> from bacteria; the genes are found in plants, Zygnematophyceae, and <a href="/wiki/Bacteria" title="Bacteria">bacteria</a>, but no other organisms. The genes may have helped to enable plants to make the transition to life on land.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-ChengXian2019_11-0" class="reference"><a href="#cite_note-ChengXian2019-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p><p>A new subclass called Spirogloeophycidae, represented by the species <i>Spirogloea muscicola</i>, was established after the unicellular <a href="/wiki/Subaerial" title="Subaerial">subaerial</a> algae, resembling a "gelatinous blob", was rediscovered on a rock close to a river bank near <a href="/wiki/Cologne" title="Cologne">Cologne</a> in 2006, after first being discovered in France in 1845.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Reproduction">Reproduction</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Zygnematophyceae&action=edit&section=2" title="Edit section: Reproduction"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The Zygnematophyceae are able to reproduce both asexually and sexually. Asexual reproduction takes place via fragmentation, <a href="/wiki/Cell_division" title="Cell division">cell division</a>, akinete formation, or parthenospores.<sup id="cite_ref-Hall_7-5" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p><a href="/wiki/Sexual_reproduction" title="Sexual reproduction">Sexual reproduction</a> in the Zygnematophyceae takes place through a process called <i><a href="/wiki/Isogamy" title="Isogamy">conjugation</a></i>.<sup id="cite_ref-pmid17272304_13-0" class="reference"><a href="#cite_note-pmid17272304-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> Unusually, the gametes lack <a href="/wiki/Flagella" class="mw-redirect" title="Flagella">flagella</a>,<sup id="cite_ref-Hall_7-6" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> and they also lack <a href="/wiki/Centriole" title="Centriole">centrioles</a> which suggest that flagella were not secondarily lost. This lack of flagella sets the Zygnematophyceae apart from nearly all groups of algae, except for <a href="/wiki/Red_algae" title="Red algae">red algae</a> and <a href="/wiki/Cyanobacteria" title="Cyanobacteria">cyanobacteria</a> (also known as blue-green algae, if they are classified as such).<sup id="cite_ref-Guiry_1-1" class="reference"><a href="#cite_note-Guiry-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> The gametes are scarcely differentiated from vegetative cells.<sup id="cite_ref-Hall_7-7" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p>In conjugation, cells or filaments of opposite sex line up, and tubes form between corresponding cells. The male cells then become amoeboid and crawl across the female, or sometimes both cells crawl into the connecting tube. The cells then meet and fuse to form a <a href="/wiki/Zygote" title="Zygote">zygote</a>, which later undergoes <a href="/wiki/Meiosis" title="Meiosis">meiosis</a> to produce new cells or filaments. As in land plants, only the female passes its <a href="/wiki/Chloroplast" title="Chloroplast">chloroplasts</a> on to the offspring.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Taxonomy">Taxonomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Zygnematophyceae&action=edit&section=3" title="Edit section: Taxonomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The Zygnematophyceae is <a href="/wiki/Monophyletic" class="mw-redirect" title="Monophyletic">monophyletic</a>. Traditionally, it has been known to contain two orders, the <a href="/wiki/Zygnematales" title="Zygnematales">Zygnematales</a> and <a href="/wiki/Desmidiales" title="Desmidiales">Desmidiales</a>.<sup id="cite_ref-Hall_7-8" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> However, the Zygnematales were found to be <a href="/wiki/Paraphyletic" class="mw-redirect" title="Paraphyletic">paraphyletic</a> with respect to the Desmidiales, and so it was split up. As of 2023<sup class="plainlinks noexcerpt noprint asof-tag update" style="display:none;"><a class="external text" href="https://en.wikipedia.org/w/index.php?title=Zygnematophyceae&action=edit">[update]</a></sup>, five orders and two subclasses are recognized: the Spirogloeales in Spirogloeophycidae, and the Serritaeniales, the <a href="/wiki/Zygnematales" title="Zygnematales">Zygnematales</a>, the Spirogyrales, and the <a href="/wiki/Desmidiales" title="Desmidiales">Desmidiales</a> in Zygnematophycidae.<sup id="cite_ref-Hess2022_9-1" class="reference"><a href="#cite_note-Hess2022-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p><p>Class Zygnematophyceae<sup id="cite_ref-AlgaeBase_15-0" class="reference"><a href="#cite_note-AlgaeBase-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p> <ul><li>Subclass <a href="/w/index.php?title=Spirogloeophycidae&action=edit&redlink=1" class="new" title="Spirogloeophycidae (page does not exist)">Spirogloeophycidae</a> <small>Melkonian, Gontcharov & Marin 2019</small><sup id="cite_ref-ChengXian2019_11-1" class="reference"><a href="#cite_note-ChengXian2019-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> <ul><li>Order <a href="/w/index.php?title=Spirogloeales&action=edit&redlink=1" class="new" title="Spirogloeales (page does not exist)">Spirogloeales</a> <small>Melkonian, Gontcharov & Marin 2019</small> <ul><li>Family <a href="/w/index.php?title=Spirogloeaceae&action=edit&redlink=1" class="new" title="Spirogloeaceae (page does not exist)">Spirogloeaceae</a> <small>Melkonian, Gontcharov & Marin 2019</small></li></ul></li></ul></li> <li>Subclass <a href="/w/index.php?title=Zygnematophycidae&action=edit&redlink=1" class="new" title="Zygnematophycidae (page does not exist)">Zygnematophycidae</a> <small>Melkonian, Gontcharov & Marin 2019</small> <ul><li>Order <a href="/w/index.php?title=Serritaeniales&action=edit&redlink=1" class="new" title="Serritaeniales (page does not exist)">Serritaeniales</a> <small>Hess & de Vries 2022</small> <ul><li>Family <a href="/w/index.php?title=Serritaeniaceae&action=edit&redlink=1" class="new" title="Serritaeniaceae (page does not exist)">Serritaeniaceae</a> <small>Hess & de Vries 2022</small></li></ul></li> <li>Order <a href="/wiki/Zygnematales" title="Zygnematales">Zygnematales</a> <small>Borge & Pascher 1931</small> <ul><li>Family <a href="/wiki/Zygnemataceae" title="Zygnemataceae">Zygnemataceae</a> <small>(Meneghini) Kützing 1843</small></li> <li>Family <a href="/wiki/Mesotaeniaceae" title="Mesotaeniaceae">Mesotaeniaceae</a> <small>Oltmans 1904</small> (paraphyletic)</li></ul></li> <li>Order <a href="/w/index.php?title=Spirogyrales&action=edit&redlink=1" class="new" title="Spirogyrales (page does not exist)">Spirogyrales</a> <small>Hess & de Vries 2022</small> <ul><li>Family <a href="/w/index.php?title=Spirogyraceae&action=edit&redlink=1" class="new" title="Spirogyraceae (page does not exist)">Spirogyraceae</a> <small>Palla 1894</small></li></ul></li> <li>Order <a href="/wiki/Desmidiales" title="Desmidiales">Desmidiales</a> <small>Krieger 1933</small> <ul><li>Family <a href="/wiki/Gonatozygaceae" title="Gonatozygaceae">Gonatozygaceae</a> <small>Fritsch 1927</small></li> <li>Family <a href="/wiki/Closteriaceae" title="Closteriaceae">Closteriaceae</a> <small>Bessey 1907 ex Pritchard 1852</small></li> <li>Family <a href="/wiki/Peniaceae" class="mw-redirect" title="Peniaceae">Peniaceae</a> <small>Haeckel 1894</small></li> <li>Family <a href="/wiki/Desmidiaceae" title="Desmidiaceae">Desmidiaceae</a> <small>Kützing 1833b ex Ralfs 1845</small></li></ul></li></ul></li></ul> <div class="mw-heading mw-heading3"><h3 id="Phylogeny">Phylogeny</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Zygnematophyceae&action=edit&section=4" title="Edit section: Phylogeny"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>A phylogeny of the families of Zygnematophyceae is presented below:<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-ChengXian2019_11-2" class="reference"><a href="#cite_note-ChengXian2019-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1258728058">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="font-size:90%;line-height:80%;width:400px;"> <tbody><tr> <td class="clade-label first"><b>Zygnematophyceae</b> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Spirogloeophycidae&action=edit&redlink=1" class="new" title="Spirogloeophycidae (page does not exist)">Spirogloeophycidae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Spirogloeales&action=edit&redlink=1" class="new" title="Spirogloeales (page does not exist)">Spirogloeales</a> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/w/index.php?title=Spirogloeaceae&action=edit&redlink=1" class="new" title="Spirogloeaceae (page does not exist)">Spirogloeaceae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Zygnematophycidae&action=edit&redlink=1" class="new" title="Zygnematophycidae (page does not exist)">Zygnematophycidae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Serritaeniales&action=edit&redlink=1" class="new" title="Serritaeniales (page does not exist)">Serritaeniales</a> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/w/index.php?title=Serritaeniaceae&action=edit&redlink=1" class="new" title="Serritaeniaceae (page does not exist)">Serritaeniaceae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Zygnematales" title="Zygnematales">Zygnematales</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Zygnemataceae" title="Zygnemataceae">Zygnemataceae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Mesotaeniaceae" title="Mesotaeniaceae">Mesotaeniaceae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Spirogyrales&action=edit&redlink=1" class="new" title="Spirogyrales (page does not exist)">Spirogyrales</a> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/w/index.php?title=Spirogyraceae&action=edit&redlink=1" class="new" title="Spirogyraceae (page does not exist)">Spirogyraceae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Desmidiales" title="Desmidiales">Desmidiales</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Gonatozygaceae" title="Gonatozygaceae">Gonatozygaceae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Closteriaceae" title="Closteriaceae">Closteriaceae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Peniaceae" class="mw-redirect" title="Peniaceae">Peniaceae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Desmidiaceae" title="Desmidiaceae">Desmidiaceae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <div class="mw-heading mw-heading2"><h2 id="Habitat">Habitat</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Zygnematophyceae&action=edit&section=5" title="Edit section: Habitat"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Members of the Zygnematophyceae are common in nearly all freshwater habitats, particularly filamentous genera such as <i><a href="/wiki/Spirogyra" title="Spirogyra">Spirogyra</a></i> and <i><a href="/wiki/Mougeotia" title="Mougeotia">Mougeotia</a></i>. Some <i>Spirogyra</i> species can tolerate disturbed habitats. On the other hand, desmids (the Desmidiales) often prefer bogs, peatlands, and lakes.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Fossil_record">Fossil record</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Zygnematophyceae&action=edit&section=6" title="Edit section: Fossil record"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Vegetative cells of the Zygnematophyceae are fragile and usually not preserved, but the zygospores are resistant to decay and can become fossilized. The Zygnematophyceae have been recorded from at least the middle <a href="/wiki/Devonian" title="Devonian">Devonian</a> period, and before the <a href="/wiki/Carboniferous" title="Carboniferous">Carboniferous</a> period, all the major groups had diverged. Fossils of zygospores are indicators of warming spring conditions and shallow, stagnant mesotrophic habitats.<sup id="cite_ref-Hall_7-9" class="reference"><a href="#cite_note-Hall-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Genomes">Genomes</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Zygnematophyceae&action=edit&section=7" title="Edit section: Genomes"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The first genomes published for Zygnematophyceae were <i>Mesotaenium endlicherianum</i> and <i>Spirogloea muscicola</i>.<sup id="cite_ref-ChengXian2019_11-3" class="reference"><a href="#cite_note-ChengXian2019-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Since then, genomes have been published on <i>Penium margaritaceum</i>,<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> <i>Zygnema</i> spp.,<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> and the <i>Closterium peracerosum–strigosum–littorale</i> complex.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> Furthermore, the gene models of <i>Mesotaenium endlicherianum</i> have been updated.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Zygnematophyceae&action=edit&section=8" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-Guiry-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-Guiry_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Guiry_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFGuiry2013" class="citation journal cs1">Guiry, Michael D. 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"}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Plant_classification" title="Template:Plant classification"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Plant_classification" title="Template talk:Plant classification"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Plant_classification" title="Special:EditPage/Template:Plant classification"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Classification_of_Archaeplastida_or_Plantae_s.l." style="font-size:114%;margin:0 4em">Classification of <a href="/wiki/Archaeplastida" title="Archaeplastida">Archaeplastida</a> or <a href="/wiki/Plant" title="Plant">Plantae <span class="rt-commentedText tooltip tooltip-dotted" title="sensu lato; in the broad sense"><i>s.l.</i></span></a></div></th></tr><tr><td class="navbox-abovebelow" colspan="2" style="background: #C3EEC3;"><div><div class="hlist"> <dl><dt><span class="nobold"><a href="/wiki/Domain_(biology)" title="Domain (biology)">Domain</a></span></dt> <dd><a href="/wiki/Template:Archaea_classification" title="Template:Archaea classification">Archaea</a></dd> <dd><a href="/wiki/Template:Bacteria_classification" title="Template:Bacteria classification">Bacteria</a></dd> <dd><a href="/wiki/Template:Eukaryota" title="Template:Eukaryota">Eukaryota</a></dd> <dt><span class="nobold">(major groups</span></dt> <dd><a href="/wiki/Template:Excavata" title="Template:Excavata">Excavata</a></dd> <dd><a href="/wiki/Diaphoretickes" title="Diaphoretickes">Diaphoretickes</a> <dl><dd><a href="/wiki/Template:Hacrobia" title="Template:Hacrobia">Hacrobia</a> <dl><dd><a href="/wiki/Template:Cryptista" title="Template:Cryptista">Cryptista</a></dd></dl></dd> <dd><a href="/wiki/Template:Rhizaria" title="Template:Rhizaria">Rhizaria</a></dd> <dd><a href="/wiki/Template:Alveolata" title="Template:Alveolata">Alveolata</a></dd> <dd><a href="/wiki/Template:Stramenopile" title="Template:Stramenopile">Stramenopiles</a></dd> <dd><a href="/wiki/Template:Plant_classification" title="Template:Plant classification">Plants</a></dd></dl></dd> <dd><a href="/wiki/Amorphea" title="Amorphea">Amorphea</a> <dl><dd><a href="/wiki/Template:Amoebozoa" title="Template:Amoebozoa">Amoebozoa</a></dd> <dd><a href="/wiki/Template:Opisthokont_protists" title="Template:Opisthokont protists">Opisthokonta</a> <dl><dd><a href="/wiki/Template:Animalia" title="Template:Animalia">Animals</a></dd> <dd><a href="/wiki/Template:Fungi_classification" title="Template:Fungi classification">Fungi</a></dd> <dd><a href="/wiki/Template:Mesomycetozoea" title="Template:Mesomycetozoea">Mesomycetozoea</a>)</dd></dl></dd></dl></dd></dl> </div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Archaeplastida" scope="row" class="navbox-group" style="width:1%;line-height:1.0em; background: #D4EED4;"><span style="margin:-40px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg);transform: rotate(270deg);"><a href="/wiki/Archaeplastida" title="Archaeplastida">Archaeplastida</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Picozoa" title="Picozoa">Picozoa</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Picomonas" class="mw-redirect" title="Picomonas">Picomonas</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Rhodelphis" title="Rhodelphis">Rhodelphidia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Rhodelphis" title="Rhodelphis">Rhodelphis</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Red_algae" title="Red algae">Rhodophyta</a><br /><span class="nobold">(red algae)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cyanidiophyceae" title="Cyanidiophyceae">Cyanidiophyceae</a></li> <li><a href="/wiki/Porphyridiophyceae" title="Porphyridiophyceae">Porphyridiophyceae</a></li> <li><a href="/wiki/Stylonematophyceae" title="Stylonematophyceae">Stylonematophyceae</a></li> <li><a href="/wiki/Compsopogonophyceae" title="Compsopogonophyceae">Compsopogonophyceae</a></li> <li><a href="/wiki/Rhodellophyceae" title="Rhodellophyceae">Rhodellophyceae</a></li> <li><a href="/wiki/Bangiophyceae" title="Bangiophyceae">Bangiophyceae</a></li> <li><a href="/wiki/Florideophyceae" title="Florideophyceae">Florideophyceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Glaucophyte" title="Glaucophyte">Glaucophyta</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Glaucophyte" title="Glaucophyte">Glaucocystophyceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><i>incertae sedis</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Algospongia" title="Algospongia">Algospongia</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Viridiplantae" title="Viridiplantae">Viridiplantae</a> <br /> <span class="nobold">or <a href="/wiki/Plant" title="Plant">Plantae <span class="rt-commentedText tooltip tooltip-dotted" title="sensu stricto; in the strict sense"><i>s.s.</i></span></a><br />(<a href="/wiki/Green_algae" title="Green algae">green algae</a> &<br /> land plants)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Prasinodermophyta" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Prasinodermophyta" title="Prasinodermophyta">Prasinodermophyta</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Prasinoderma" title="Prasinoderma">Prasinodermophyceae</a></li> <li><a href="/wiki/Palmophyllophyceae" title="Palmophyllophyceae">Palmophyllophyceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin:-29px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);"> <a href="/wiki/Chlorophyta" title="Chlorophyta">Chlorophyta</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;">Prasinophytina</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mamiellophyceae" title="Mamiellophyceae">Mamiellophyceae</a></li> <li><a href="/wiki/Pyramimonadophyceae" title="Pyramimonadophyceae">Pyramimonadophyceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;">Chlorophytina</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nephroselmidaceae" title="Nephroselmidaceae">Nephroselmidophyceae</a></li> <li><a href="/wiki/Picocystis" title="Picocystis">Picocystophyceae</a></li> <li><a href="/wiki/Chloropicophyceae" class="mw-redirect" title="Chloropicophyceae">Chloropicophyceae</a></li> <li><a href="/wiki/Pedinophyceae" title="Pedinophyceae">Pedinophyceae</a></li> <li><a href="/wiki/Chlorodendrales" title="Chlorodendrales">Chlorodendrophyceae</a></li> <li><a href="/wiki/UTC_clade" title="UTC clade">UTC clade</a> <ul><li><a href="/wiki/Ulvophyceae" title="Ulvophyceae">Ulvophyceae</a></li> <li><a href="/wiki/Trebouxiophyceae" title="Trebouxiophyceae">Trebouxiophyceae</a></li> <li><a href="/wiki/Chlorophyceae" title="Chlorophyceae">Chlorophyceae</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin:-29px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);"><a href="/wiki/Streptophyta" title="Streptophyta">Streptophyta</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mesostigmatophyceae" title="Mesostigmatophyceae">Mesostigmatophyceae</a></li> <li><a href="/wiki/Chlorokybophyceae" class="mw-redirect" title="Chlorokybophyceae">Chlorokybophyceae</a></li> <li><i><a href="/wiki/Spirotaenia" title="Spirotaenia">Spirotaenia</a></i></li> <li><a href="/wiki/Klebsormidiophyceae" class="mw-redirect" title="Klebsormidiophyceae">Klebsormidiophyceae</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Phragmoplastophyta" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Phragmoplastophyta" title="Phragmoplastophyta">Phragmoplastophyta</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Charophyceae" title="Charophyceae">Charophyceae</a></li> <li><a href="/wiki/Coleochaetophyceae" title="Coleochaetophyceae">Coleochaetophyceae</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;">Anydrophyta</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Zygnematophyceae</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Embryophyta" class="mw-redirect" title="Embryophyta">Embryophyta</a><br /><span class="nobold">(land plants)</span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>(see below↓)</li></ul> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background: #C3EEC3;"><div id="Embryophytes_or_Plantae_sensu_strictissimo" style="font-size:114%;margin:0 4em"><a href="/wiki/Embryophyte" title="Embryophyte">Embryophytes</a> or <a href="/wiki/Plant" title="Plant">Plantae <span class="rt-commentedText tooltip tooltip-dotted" title="in the strictest sense"><i>sensu strictissimo</i></span></a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Bryophytes" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -25px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"><a href="/wiki/Bryophyte" title="Bryophyte">Bryophytes</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Marchantiophyta" title="Marchantiophyta">Marchantiophyta</a><br /><span class="nobold">(liverworts)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Haplomitriopsida" title="Haplomitriopsida">Haplomitriopsida</a></li> <li><a href="/wiki/Marchantiopsida" title="Marchantiopsida">Marchantiopsida</a></li> <li><a href="/wiki/Jungermanniopsida" title="Jungermanniopsida">Jungermanniopsida</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Hornwort" title="Hornwort">Anthocerotophyta</a><br /><span class="nobold">(hornworts)</span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Leiosporoceros" title="Leiosporoceros">Leiosporocerotopsida</a></li> <li><a href="/wiki/Hornwort" title="Hornwort">Anthocerotopsida</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Moss" title="Moss">Bryophyta</a><br /><span class="nobold">(mosses)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Takakia" title="Takakia">Takakiopsida</a></li> <li><a href="/wiki/Sphagnopsida" title="Sphagnopsida">Sphagnopsida</a></li> <li><a href="/wiki/Andreaeobryum" title="Andreaeobryum">Andreaeobryopsida</a></li> <li><a href="/wiki/Andreaeaceae" title="Andreaeaceae">Andreaeopsida</a></li> <li><a href="/wiki/Oedipodium" title="Oedipodium">Oedipodiopsida</a></li> <li><a href="/wiki/Tetraphidaceae" title="Tetraphidaceae">Tetraphidopsida</a></li> <li><a href="/wiki/Polytrichaceae" title="Polytrichaceae">Polytrichopsida</a></li> <li><a href="/wiki/Bryopsida" title="Bryopsida">Bryopsida</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -55px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"> <a href="/wiki/Polysporangiophyte" title="Polysporangiophyte">Polysporangiophytes</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="†Protracheophytes*" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span class="nobold">†</span>Protracheophytes*</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/wiki/Horneophytopsida" title="Horneophytopsida">Horneophytopsida</a></li> <li>†<i><a href="/wiki/Aglaophyton" title="Aglaophyton">Aglaophyton</a></i></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -25px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"><a href="/wiki/Vascular_plant" title="Vascular plant">Tracheophytes</a><br /><span class="nobold">(vascular plants)</span></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="†Paratracheophytes*" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span class="nobold">†</span>Paratracheophytes*</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/w/index.php?title=Cooksoniopsida&action=edit&redlink=1" class="new" title="Cooksoniopsida (page does not exist)">Cooksoniopsida</a></li> <li>†<a href="/wiki/Rhyniophyte" title="Rhyniophyte">Rhyniopsida</a></li> <li>†<a href="/w/index.php?title=Renaliales&action=edit&redlink=1" class="new" title="Renaliales (page does not exist)">Renaliales</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -40px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);">Eutracheophytes</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Lycophytes" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Lycophyte" title="Lycophyte">Lycophytes</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/wiki/Barinophyte" title="Barinophyte">Barinophytopsida</a></li> <li>†<a href="/wiki/Zosterophyll" title="Zosterophyll">Zosterophyllopsida</a></li> <li><a href="/wiki/Lycopodiopsida" title="Lycopodiopsida">Lycopodiopsida</a> (<a href="/wiki/Lycopodiaceae" title="Lycopodiaceae">clubmosses</a>, <a href="/wiki/Selaginella" title="Selaginella">spikemosses</a> & <a href="/wiki/Isoetes" title="Isoetes">quillworts</a>)</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -35px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"><a href="/wiki/Euphyllophyte" title="Euphyllophyte">Euphyllophytes</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/wiki/Eophyllophyton" title="Eophyllophyton">Eophyllophytopsida</a></li> <li>†<a href="/wiki/Trimerophytopsida" title="Trimerophytopsida">Trimerophytopsida</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Moniliformopses" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;">Moniliformopses</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/w/index.php?title=Ibykales&action=edit&redlink=1" class="new" title="Ibykales (page does not exist)">Ibykales</a></li> <li>†<a href="/wiki/Cladoxylopsida" title="Cladoxylopsida">Cladoxylopsida</a></li> <li><a href="/wiki/Fern" title="Fern">Polypodiopsida</a> (ferns and <a href="/wiki/Equisetum" title="Equisetum">horsetails</a>)</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -30px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);">Lignophytes</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="†Progymnosperms*" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span class="nobold">†</span><a href="/wiki/Progymnosperm" title="Progymnosperm">Progymnosperms</a>*</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/w/index.php?title=Noeggerathiopsida&action=edit&redlink=1" class="new" title="Noeggerathiopsida (page does not exist)">Noeggerathiopsida</a></li> <li>†<a href="/w/index.php?title=Aneurophytopsida&action=edit&redlink=1" class="new" title="Aneurophytopsida (page does not exist)">Aneurophytopsida</a></li> <li>†<a href="/w/index.php?title=Archaeopteridopsida&action=edit&redlink=1" class="new" title="Archaeopteridopsida (page does not exist)">Archaeopteridopsida</a></li> <li>†<a href="/w/index.php?title=Protopityales&action=edit&redlink=1" class="new" title="Protopityales (page does not exist)">Protopityales</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -25px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"><a href="/wiki/Spermatophyte" class="mw-redirect" title="Spermatophyte">Spermatophytes</a><br /><span class="nobold">(seed plants)</span></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span class="nobold">†</span><a href="/wiki/Pteridospermatophyta" title="Pteridospermatophyta">Pteridosperms</a><span class="nobold">*</span><br /><span class="nobold">(seed ferns)<br />and other extinct<br /> seed plant groups</span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/wiki/Bennettitales" title="Bennettitales">Bennettitales</a></li> <li>†<a href="/wiki/Calamopityales" class="mw-redirect" title="Calamopityales">Calamopityales</a></li> <li>†<a href="/wiki/Callistophytales" title="Callistophytales">Callistophytales</a></li> <li>†<a href="/wiki/Caytoniales" title="Caytoniales">Caytoniales</a></li> <li>†<a href="/wiki/Cordaitales" title="Cordaitales">Cordaitales</a></li> <li>†<a href="/wiki/Corystospermaceae" title="Corystospermaceae">Corystospermales</a></li> <li>†<a href="/wiki/Czekanowskiales" title="Czekanowskiales">Czekanowskiales</a></li> <li>†<a href="/wiki/Gigantopteridales" class="mw-redirect" title="Gigantopteridales">Gigantopteridales</a></li> <li>†<a href="/wiki/Glossopteridales" title="Glossopteridales">Glossopteridales</a></li> <li>†<a href="/wiki/Lyginopteridales" title="Lyginopteridales">Lyginopteridopsida</a></li> <li>†<a href="/wiki/Medullosales" title="Medullosales">Medullosales</a></li> <li>†<a href="/wiki/Peltaspermales" title="Peltaspermales">Peltaspermopsida</a></li> <li>†<a href="/wiki/Pentoxylales" title="Pentoxylales">Pentoxylopsida</a></li> <li>†<a href="/w/index.php?title=Petriellales&action=edit&redlink=1" class="new" title="Petriellales (page does not exist)">Petriellales</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Gymnosperm" title="Gymnosperm">Acrogymnospermae</a><br /><span class="nobold">(living gymnosperms)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cycad" title="Cycad">Cycadopsida (cycads)</a></li> <li><a href="/wiki/Ginkgoales" title="Ginkgoales">Ginkgoales (<i>Ginkgo</i>)</a></li> <li><a href="/wiki/Conifer" title="Conifer">Pinopsida (conifers)</a></li> <li><a href="/wiki/Gnetophyta" title="Gnetophyta">Gnetopsida</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Flowering_plant" title="Flowering plant">Angiospermae</a><br /><span class="nobold">(flowering plants)</span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Basal_angiosperms" title="Basal angiosperms">Basal angiosperms</a></li> <li><a href="/wiki/Mesangiospermae" title="Mesangiospermae">Core angiosperms</a> <ul><li><a href="/wiki/Magnoliids" title="Magnoliids">Magnoliids</a></li> <li><a href="/wiki/Monocotyledon" title="Monocotyledon">Monocots</a></li> <li><a href="/wiki/Eudicots" title="Eudicots">Eudicots</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="background: #C3EEC3;"><div> <ul><li>*<a href="/wiki/Paraphyletic" class="mw-redirect" title="Paraphyletic">paraphyletic</a> groups</li> <li>† = <a href="/wiki/Extinction" title="Extinction">extinct</a></li> <li><span class="noviewer" typeof="mw:File"><span title="List-Class article"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/d/db/Symbol_list_class.svg/16px-Symbol_list_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/d/db/Symbol_list_class.svg/23px-Symbol_list_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/d/db/Symbol_list_class.svg/31px-Symbol_list_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/List_of_plant_orders" title="List of plant orders">List of plant orders</a></li> <li><a href="/wiki/Plant#Current_definitions_of_Plantae" title="Plant">Current definitions of Plantae</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Zygnematophyceae</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q1151744" class="extiw" title="wikidata:Q1151744">Q1151744</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Zygnematophyceae" class="extiw" title="wikispecies:Zygnematophyceae">Zygnematophyceae</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/AlgaeBase" title="AlgaeBase">AlgaeBase</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.algaebase.org/browse/taxonomy/detail/?taxonid=113648">113648</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Barcode_of_Life_Data_System" title="Barcode of Life Data System">BOLD</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.boldsystems.org/index.php/TaxBrowser_TaxonPage?taxid=413966">413966</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/4053">4053</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/EPPO_Code" title="EPPO Code">EPPO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://gd.eppo.int/taxon/1ZGNC">1ZGNC</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/333">333</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/123848">123848</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=954910">954910</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=131209">131209</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=322824">322824</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=55101">55101</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Tropicos" title="Tropicos">Tropicos</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://legacy.tropicos.org/Name/100370504">100370504</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.marinespecies.org/aphia.php?p=taxdetails&id=162685">162685</a></span></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Conjugatophyceae</i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q110567253" class="extiw" title="wikidata:Q110567253">Q110567253</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/622C9">622C9</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/9748626">9748626</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/774724">774724</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=322824">322824</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.marinespecies.org/aphia.php?p=taxdetails&id=886290">886290</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1038841319"></div><div role="navigation" class="navbox authority-control" aria-label="Navbox" style="padding:3px"><table class="nowraplinks hlist navbox-inner" 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