CINXE.COM
Just a moment...
<!DOCTYPE html><html lang="en-US"><head><title>Just a moment...</title><meta http-equiv="Content-Type" content="text/html; charset=UTF-8"><meta http-equiv="X-UA-Compatible" content="IE=Edge"><meta name="robots" content="noindex,nofollow"><meta name="viewport" content="width=device-width,initial-scale=1"><style>*{box-sizing:border-box;margin:0;padding:0}html{line-height:1.15;-webkit-text-size-adjust:100%;color:#313131;font-family:system-ui,-apple-system,BlinkMacSystemFont,Segoe UI,Roboto,Helvetica Neue,Arial,Noto Sans,sans-serif,Apple Color Emoji,Segoe UI Emoji,Segoe UI Symbol,Noto Color Emoji}body{display:flex;flex-direction:column;height:100vh;min-height:100vh}.main-content{margin:8rem auto;max-width:60rem;padding-left:1.5rem}@media (width <= 720px){.main-content{margin-top:4rem}}.h2{font-size:1.5rem;font-weight:500;line-height:2.25rem}@media (width <= 720px){.h2{font-size:1.25rem;line-height:1.5rem}}#challenge-error-text{background-image:url(data:image/svg+xml;base64,PHN2ZyB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciIHdpZHRoPSIzMiIgaGVpZ2h0PSIzMiIgZmlsbD0ibm9uZSI+PHBhdGggZmlsbD0iI0IyMEYwMyIgZD0iTTE2IDNhMTMgMTMgMCAxIDAgMTMgMTNBMTMuMDE1IDEzLjAxNSAwIDAgMCAxNiAzbTAgMjRhMTEgMTEgMCAxIDEgMTEtMTEgMTEuMDEgMTEuMDEgMCAwIDEtMTEgMTEiLz48cGF0aCBmaWxsPSIjQjIwRjAzIiBkPSJNMTcuMDM4IDE4LjYxNUgxNC44N0wxNC41NjMgOS41aDIuNzgzem0tMS4wODQgMS40MjdxLjY2IDAgMS4wNTcuMzg4LjQwNy4zODkuNDA3Ljk5NCAwIC41OTYtLjQwNy45ODQtLjM5Ny4zOS0xLjA1Ny4zODktLjY1IDAtMS4wNTYtLjM4OS0uMzk4LS4zODktLjM5OC0uOTg0IDAtLjU5Ny4zOTgtLjk4NS40MDYtLjM5NyAxLjA1Ni0uMzk3Ii8+PC9zdmc+);background-repeat:no-repeat;background-size:contain;padding-left:34px}@media (prefers-color-scheme:dark){body{background-color:#222;color:#d9d9d9}}</style><meta http-equiv="refresh" content="390"></head><body class="no-js"><div class="main-wrapper" role="main"><div class="main-content"><noscript><div class="h2"><span id="challenge-error-text">Enable JavaScript and cookies to continue</span></div></noscript></div></div><script>(function(){window._cf_chl_opt={cvId: '3',cZone: "www.science.org",cType: 'managed',cRay: '8e94aa263e9b4c9b',cH: 'hVvSIueplFxObXxVz5SJfDk6nEM.XVDplUZmmDLwZ.I-1732737209-1.2.1.1-DX5pDnLuC4KPjDA75sYzOAbweWyttTEZ3Y0g9BswKTO4xSeex4PJobL7V4lOEA6y',cUPMDTk: "\/doi\/10.1126\/sciadv.aba2578?__cf_chl_tk=2.zW5Ku3vmdXWZTChqoTzgma0DCggq.8rpmsqTfy5Ss-1732737209-1.0.1.1-CPWZK3H2DabvnrK0qaXJ0KOxij2VSlNMo1qMp7.bEug",cFPWv: 'b',cITimeS: '1732737209',cTTimeMs: '1000',cMTimeMs: '390000',cTplC: 0,cTplV: 5,cTplB: 'cf',cK: "",fa: "\/doi\/10.1126\/sciadv.aba2578?__cf_chl_f_tk=2.zW5Ku3vmdXWZTChqoTzgma0DCggq.8rpmsqTfy5Ss-1732737209-1.0.1.1-CPWZK3H2DabvnrK0qaXJ0KOxij2VSlNMo1qMp7.bEug",md: "Sq.BnvxHCMu3SMqo36JVoQAdkuwqxXCUYF.QCXJo0LU-1732737209-1.2.1.1-kpyKyxt5ka_X8vlo9x5ApacQUg4bbXtzKWgu8Pn3CuREWlQQCP4tGW5jkp4Yafhe3YarD6xp3Kdm0cKHMGs1Et22X0zpJ8Gc5xzzqMBm32U0rDORnrAb_lvlovFR3Whx7Q.cVr3OLHTmUU9zHfSaVwJsr2x2Jxir39UVxFYlzqbrU3vCXZOJwKMtyN8cUaZ_zSSw.W2wl0.x2rUF4YsbUBaP58o..Ryg_zQXdYVUIPT_gHNwVvI7fJnFjjCB7BCd74p0x8ToGzpNiBqIkvyRCJYDLz13GSeiakIkkXivcnJGKnFcqlSZ_IsQGhj3n0VWSDhB1l6av5clYlae1VeF.4TfUsuQgW_GQ2SBbalfzmPIXMNmaSevwDyRBKuG2gYTegibHrJ5QFC6JlxPVilEdAIhGHAgq0eUlNf0HXkSnJBsCp7RTqbd9XfrHpXiiEcAqvdq2Lk5Y5MRbxQX72I6dsPbY7pKhe3UkNoPgCf5i1sgBWmRJ6PONl6eUvHooL9U.ibPaCXn8QvciDlRVz8sWLrA4rO4nN9yV3keozBicQjsKBfXDLa91JS60QCstYUCyjRDDh93mA1OFj.oCS2_s8vYqkCFbrBJbL0JOx3V6pJRTzxINhxfeNR4ROGtneG0qVcg.xxk9fyc.As5ICY6rnR__jiMqdBStXCc9obdxRHKW83UKyOJUkAl8nA7_vz_VsaR.yg1n0AXh2_b.sstzXvZQNEQ9FfZyVsaoAcxtVps9wAbtrRLLvRZGdJMlYuTcrQdwRh2Am1U1QjjmcCvwD6oCcT12ySZtKU.QRZOqOessCLYGoXT6n7lCdfMb3qbvoqIAiiiqyo4rmPeJP8kiAtlVukEwB_pfI2nQLla5aaYLMNcdlI.gHdEvyNgtpgRlqTghz47W1ukxkaSgNPTLneSOiZYFlrLx7BS.K_CTbu1cX156Xv2NnKR7MWqhY79MAvxxXFdaPQvjPc9y_JD7jpWr.GB_261wgR15wN8YWQlC3NX_rxw1xRDkgsKuRE6H.oZ3Te8kUM9xwtXFHeIGX.aIMMYYVOF.tx9J18.JCQ3BLsnNqR4iFCqDS_tRrKQE4yUKcj9zLC1sWIZ5jXI8VQA1eK1yioSAUMqkYqS3tw2hivN2zqw_QjQynEVyAg6NR4FhDP.oTq_rt7BqrCLtuei9Si4.iF21gXYRjP4MQEvBZdLP_w4.n1KSHzhg1exLSj4x.1uoB678LKO0B2tvJ8HMXCR7ObUP5vEdXWbZNHlvBMCltBuzGcY6MO4WB8.d93Zt0Vt4xqT1MC3txx5Ca49MEKeMNlpo.tNUI942LQxck5q6u3yrPF.WtlOZAYxRSFBmoOFhFDbruc0Uqum0GtvPZmSZkX6cP5CRwDJNLIaA7poCH3ZWF9ncIMO9TxFSISzxqyeHJ4EfsjVADQZSVzmVXETUlME8LS9obtp9Bd1rtR.WkNFW1t3CJcIFCHVNjj2l4kk5BQBPeAL7wZavlYVpnP1XQ.AAuG.AtxuujZVxU4ijYj2uL7h1mC3cCxE45f3KEijCVKvYjhg4Iwe7TsoqEHy_9tkhoJhYEh0IrMNZs8gaUAql4hXrVSlGN2eOQ3TQvCfy7ReZgDDygFI95kCkB6uGA3NAXN6W6sYYVvbuSa6wiZ9uDze_tG9QoR5q9XN_HdD7oKzIexgVBydP33JrBtENG5bgRDvdzfduOL.HsuUOCJu3Lcd6hpGOP0F6p2nSf6Ws9wAWd8F1G1yMQJMS70ZQVns3EpWxrjHOIkngyhnx2TW6kimw7VJwsxgm9aU5BaFo9tQCwdE4X6xSda8k_IQz_JDlr2aCM9NkLcKAWeEENyngvtHxrFE_rFwUxMO47oaqct7Bs_WXuC.nVUltaQYRkRU3sILIgdxA5s.jCaRFeWHzFy1Pvm34zhvCOHEylq.BnvuvikROFXx_4_hATbImMSQBLyAcp1WFr6LUbmDodoUSVdSwR3CvufctJvaajkTIpbGAI8kQrFQlct_ak8qY56Z052Nu439kAF62yzz4B_oS.otbSggvlwbzB_nGzkOVNxCNwEi8jfv4yRgvF6KkZ8IM0T8i5fxDuUn4OIAIa5x0yu4.p0ilE68ilemwhs8I92a6b2sPupMAkenMaHTJFWSSwBNPI8gYecVlPgwUeb.BpuM9gu95bb1FGcZMvyi4SWmYfPjQDyEixuA3ypgwr7206A0Mq2k2xrCuuxi57fz5kDkprwpFrQqn9iPQyqGSWt4JTa1aC25H1r_AO0j9kbAlhggl.l6AUp6qp9KIyx7syzbt07MRZ_l4h0OxMujcOoGZNxwTiwcONnW233oDbFchTTWimnA4KGov4c9xnyR6gHTuNLsvRaq3x8tQVTxAIf__q1b6QQJ5ZkbbAtyj8OJQjpuZZfKs.u7WPTducYujtWniJ4yQkKccHrLV4Ph_tSxw0jglYDskhE2q.x8F2NlXfD0cFCdN6h0U07BhYSgafpynBg41enzmFqp2.d9s4hOzVcwduDD5.N8.Pa6ochi_U_NoQm2.UK1g6NqKhw1IlZOvWrlGLy5wINi3ZdmqBcWwTOQ2o4iVjmoi3yclzaBTOWMQV7gCcaCIKdVxSDVOf8E84A01UL3K2LW_6ebiV8lN7ZM06wt64F1UZS5Z5_qVjNp9ZL.koFgC.Fd3zb7z8co1QGCl3rYNSfEOyG8XaUmZy3UgDKX.ATi9lpB_nYPfHz.Efj6TqXzI1TKIujkKTFUFNEFOR1l9R96ZkqDYSGAs2avy046XdPj.JcuiBwiVehCYTB1QX5XBGf74a5Qb05T4AsapfHDi_md8hq513ES_N1epW2W1wgN8jU36nnuNglHdhG5m.VnuTlHMOiOVLN4bAJHfOShdA_ja2LRqhmxikrUxrXSE16tNMSjYabKjnG0OSSqYg.i8UIYvGZFDZtCrn05acWFmH.lyyV2NSqf5Q1biIILZs9AlYzz4kFA4bcYXnzwJz8",mdrd: "deg5VWDNV5uIueoOj0hfBZCcrtTWsBZg0N_032OtlLU-1732737209-1.2.1.1-TWzGPwA9ZmEEniQW3aK665VgPmIa9.UOa5pYobXZoUzwuULeczByxCrhrp1HyfnslpMnxUB9MXkkZ05qcvtU1w7pfLHC2teuKT59vdImxHadH3MzcBUMNHNedO8s5F3IkPwujqn0hgtkDXE9twMgIBGeEB.s6A.ohjQ6mZBBGfyrySgfQZ1fqlsc9S36jNQkRGKHpUzrPRIPiy8LTRDlQznsjvcW4utMCIzHbxZcmDzwG5nTd4nsenxBws5k24fcLRj799E7mW2CSmYb0fQ5sSqVjtCbdISK9fvfB4qOhO1PV1C3Bs4C4eve2QV98W8LhwFYsPwoKyRkXdezxOUoK3jqMiunwJ57NjP6bLscVJnRA87tzEBLHl7Hp1l_K..JEW9cMeLt5.NxPdRE5krKPiXMGj6FGxg2iN3c5qEir0PjZFvThYlR0vYd_B9WCDK17h8NBGTecfBJt07KoeTVYFoluGeDBett_5p7qEXS77sZu2EIhZq3mgvNJPpwu6vTFw9tdnkE9DCDcFa2OMGhKsvnswT9OPtyBNWG7zEb2xzz1J_w5DsakwfH4.uPNU0HpWD_CAkumio8Iqvp3lhSpT15U5zEqxwm3J9pSUrMJ9GLJ19wxyVyHIblxJ.FMxQQpUwY4qpMVPTn3qGckuD8Gd_j897msmVM_tirWCycQu6b8QBC7qDj7PKr6kW5fEY8X4wqqR_zeFeKFHHtpjIYxEXSoEQKsBfsU.7lMyh5ZuYDzWD3qZNUxbl6M5SqZnktJetng_bjOKYmTXpBr6foGMspcCdXkWoQQCcOCHvdTMKXGFooHKFxuY80nHv0cXhc2emTEyjdhB1HjeENVgCLi9M02q1mW2o2dIOWZPV4E7bzM0xUArwfoNtQPPJXVJz8Xl2swjhsLcHSvaPwZw4BJxR_pCZ_njybF2ZIqtwH02aTw8mLg7C5PBwkbEM5IdsHP6MM_ofj6Dcc70Xav7z2JOJxJaKGOI_pkfjZbOo0QJhzD191fUbBQ2qyzc1ngECRAKwlztFhhooy__Rx1RPhubUpx22nvUL.f1NerplXmxTjpH37ix1Sbs6J3nKGrtoBL32UCy7uCC22FPnwnE5OrlwBJnDjD.zj9n0iy7Zdap0w6tw9DiBfUY2Zz4AgyBkmNPCgBxiaG.FQsIYBfuL2Xy4ZjEUMQymZ_skE.IOR9N7oYEzPXU7mLjr0cv0f89Rkv.p5PntLVUahWT.DIS25oQmSLgXfK3CCP4AEz316rAO.Pv7BwS_u.INYjGwvE1gudVQAKMfu6eze5C5htgMs.eimwX6HKpXCXvbneUa74EQK8KNi.vdNN2bwjuEyebfCA2cd7dPP0GuRKpNzJZYBV7wPM0XI9MBGgS3Oay4mK40kvJVCdUcpxnjInfgQ0FCitXlc0lumoTOPT1iImyJhByJLCi0aPt7hq5k3FZhY6UehxQiV8s3CNgWCnIRESf6CagOAoEsTY_tUqwolWt7U9uR_mW2TtaxHCtw.o.t7WFdGZKzeKccv_oHW7fQLRhEcJrHf3ebjzUkRxFmbRPvnAznoghzCbV0F71kzYBQrW59ACyIWh7QymQRq400Tdgx2jExl6tltfJkYGkrA6AiCFLqUIxFqxvCYvavMV_HZAsB4W6o6ezthgwDMGwF8.30GC7ME85qvTdOiJM.f.gickeVrosYRDTG2Jhrh3brb5ILF0.e645MCxuaIcTRvAq212IbRplGNi1IBsSXjshdtpnF3f9zTvQb0RILv7v594PC9tLRWPp_8_ev7ySKah52NL6Qn3y.kpYOyvoK2SXEQG0JA_wkiyPdvXK7Ic0z7FVRz6Gk.CXxGR1baEppnP5upvNfMNjs.Wjd2zrEQcBVPzbxa6nbcMW29UkWi_AtLwiBY_EybL4R_ymBy6h8s3wWUQG_p.tOZgv8qB5TnXy_Y9vCCpR_EF7Hg45ZoyhuR2VmBgL3wS6jTpKCLN665V4F1"};var cpo = document.createElement('script');cpo.src = '/cdn-cgi/challenge-platform/h/b/orchestrate/chl_page/v1?ray=8e94aa263e9b4c9b';window._cf_chl_opt.cOgUHash = location.hash === '' && location.href.indexOf('#') !== -1 ? '#' : location.hash;window._cf_chl_opt.cOgUQuery = location.search === '' && location.href.slice(0, location.href.length - window._cf_chl_opt.cOgUHash.length).indexOf('?') !== -1 ? '?' : location.search;if (window.history && window.history.replaceState) {var ogU = location.pathname + window._cf_chl_opt.cOgUQuery + window._cf_chl_opt.cOgUHash;history.replaceState(null, null, "\/doi\/10.1126\/sciadv.aba2578?__cf_chl_rt_tk=2.zW5Ku3vmdXWZTChqoTzgma0DCggq.8rpmsqTfy5Ss-1732737209-1.0.1.1-CPWZK3H2DabvnrK0qaXJ0KOxij2VSlNMo1qMp7.bEug" + window._cf_chl_opt.cOgUHash);cpo.onload = function() {history.replaceState(null, null, ogU);}}document.getElementsByTagName('head')[0].appendChild(cpo);}());</script></body></html>