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emergence</span> </div> </a> <ul id="toc-Causes_of_emergence-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Taxonomy" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Taxonomy"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>Taxonomy</span> </div> </a> <ul id="toc-Taxonomy-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>References</span> </div> </a> <ul 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mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B9%E0%A7%87%E0%A6%A8%E0%A6%BF%E0%A6%AA%E0%A6%BE%E0%A6%AD%E0%A6%BE%E0%A6%87%E0%A6%B0%E0%A6%BE%E0%A6%B8" title="হেনিপাভাইরাস – Bangla" lang="bn" hreflang="bn" data-title="হেনিপাভাইরাস" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-bh mw-list-item"><a href="https://bh.wikipedia.org/wiki/%E0%A4%B9%E0%A5%87%E0%A4%A8%E0%A4%BF%E0%A4%AA%E0%A4%BE%E0%A4%B5%E0%A4%BE%E0%A4%87%E0%A4%B0%E0%A4%B8" title="हेनिपावाइरस – Bhojpuri" lang="bh" hreflang="bh" data-title="हेनिपावाइरस" data-language-autonym="भोजपुरी" data-language-local-name="Bhojpuri" class="interlanguage-link-target"><span>भोजपुरी</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A5%D0%B5%D0%BD%D0%B8%D0%BF%D0%B0%D0%B2%D0%B8%D1%80%D1%83%D1%81%D0%B8" title="Хенипавируси – Bulgarian" lang="bg" hreflang="bg" data-title="Хенипавируси" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Henipavirus" title="Henipavirus – Catalan" lang="ca" hreflang="ca" data-title="Henipavirus" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Henipavirus" title="Henipavirus – German" lang="de" hreflang="de" data-title="Henipavirus" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Henipavirus" title="Henipavirus – Spanish" lang="es" hreflang="es" data-title="Henipavirus" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Henipabirus" title="Henipabirus – Basque" lang="eu" hreflang="eu" data-title="Henipabirus" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%87%D9%86%DB%8C%D9%BE%D8%A7%D9%88%DB%8C%D8%B1%D9%88%D8%B3" title="هنیپاویروس – Persian" lang="fa" hreflang="fa" data-title="هنیپاویروس" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Henipavirus" title="Henipavirus – French" lang="fr" hreflang="fr" data-title="Henipavirus" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%B9%E0%A5%87%E0%A4%A8%E0%A4%BF%E0%A4%AA%E0%A4%BE%E0%A4%B5%E0%A4%BE%E0%A4%87%E0%A4%B0%E0%A4%B8" title="हेनिपावाइरस – Hindi" lang="hi" hreflang="hi" data-title="हेनिपावाइरस" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Henipavirus" title="Henipavirus – Indonesian" lang="id" hreflang="id" data-title="Henipavirus" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Henipavirus" title="Henipavirus – Italian" lang="it" hreflang="it" data-title="Henipavirus" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/Henipavirus" title="Henipavirus – Hebrew" lang="he" hreflang="he" data-title="Henipavirus" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Henipav%C3%ADrus" title="Henipavírus – Hungarian" lang="hu" hreflang="hu" data-title="Henipavírus" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%A8%E0%B4%BF%E0%B4%AA%E0%B4%BE_%E0%B4%B5%E0%B5%88%E0%B4%B1%E0%B4%B8%E0%B5%8D" title="നിപാ വൈറസ് – Malayalam" lang="ml" hreflang="ml" data-title="നിപാ വൈറസ്" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%81%D9%8A%D8%B1%D9%88%D8%B3_%D9%87%D9%8A%D9%86%D9%8A%D8%A8%D8%A7" title="فيروس هينيبا – Egyptian Arabic" lang="arz" hreflang="arz" data-title="فيروس هينيبا" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Henipavirus" title="Henipavirus – Dutch" lang="nl" hreflang="nl" data-title="Henipavirus" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Henipavirus" title="Henipavirus – Romanian" lang="ro" hreflang="ro" data-title="Henipavirus" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/Henipavirus" title="Henipavirus – Russian" lang="ru" hreflang="ru" data-title="Henipavirus" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Henipavirus" title="Henipavirus – Slovenian" lang="sl" hreflang="sl" data-title="Henipavirus" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" 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class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Genus of RNA viruses</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)"><i>Henipavirus</i> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:CSIRO_ScienceImage_1718_The_Hendra_Virus.jpg" class="mw-file-description"><img alt="Colored transmission electron micrograph of a "Hendra henipavirus" virion (ca. 300 nm length)" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e0/CSIRO_ScienceImage_1718_The_Hendra_Virus.jpg/220px-CSIRO_ScienceImage_1718_The_Hendra_Virus.jpg" decoding="async" width="220" height="257" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e0/CSIRO_ScienceImage_1718_The_Hendra_Virus.jpg/330px-CSIRO_ScienceImage_1718_The_Hendra_Virus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e0/CSIRO_ScienceImage_1718_The_Hendra_Virus.jpg/440px-CSIRO_ScienceImage_1718_The_Hendra_Virus.jpg 2x" data-file-width="1517" data-file-height="1772" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Colored <a href="/wiki/Transmission_electron_micrograph" class="mw-redirect" title="Transmission electron micrograph">transmission electron micrograph</a> of a <i>Hendra henipavirus</i> virion (ca. 300 nm length) </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(250,250,190)"><a href="/wiki/Virus_classification" title="Virus classification">Virus classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Henipavirus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>(unranked): </td> <td><a href="/wiki/Virus" title="Virus">Virus</a> </td></tr> <tr> <td><i>Realm</i>: </td> <td><a href="/wiki/Riboviria" title="Riboviria"><i>Riboviria</i></a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Orthornavirae" title="Orthornavirae"><i>Orthornavirae</i></a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Negarnaviricota" class="mw-redirect" title="Negarnaviricota"><i>Negarnaviricota</i></a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Monjiviricetes" title="Monjiviricetes"><i>Monjiviricetes</i></a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Mononegavirales" title="Mononegavirales"><i>Mononegavirales</i></a> </td></tr> <tr> <td>Family: </td> <td><a href="/wiki/Paramyxoviridae" title="Paramyxoviridae"><i>Paramyxoviridae</i></a> </td></tr> <tr> <td>Subfamily: </td> <td><a href="/wiki/Orthoparamyxovirinae" title="Orthoparamyxovirinae"><i>Orthoparamyxovirinae</i></a> </td></tr> <tr> <td>Genus: </td> <td><b><i>Henipavirus</i></b> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)">Species </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><i><a href="/wiki/Cedar_henipavirus" class="mw-redirect" title="Cedar henipavirus">Cedar henipavirus</a></i></li> <li><i><a href="/wiki/Ghanaian_bat_henipavirus" title="Ghanaian bat henipavirus">Ghanaian bat henipavirus</a></i></li> <li><i><a href="/wiki/Hendra_henipavirus" class="mw-redirect" title="Hendra henipavirus">Hendra henipavirus</a></i></li> <li><i><a href="/wiki/Langya_henipavirus" title="Langya henipavirus">Langya henipavirus</a></i></li> <li><i><a href="/wiki/Mojiang_henipavirus" class="mw-redirect" title="Mojiang henipavirus">Mojiang henipavirus</a></i></li> <li><i><a href="/wiki/Nipah_henipavirus" class="mw-redirect" title="Nipah henipavirus">Nipah henipavirus</a></i></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Henipavirus</b></i> is a <a href="/wiki/Genus" title="Genus">genus</a> of <a href="/wiki/Negative-strand_RNA_virus" title="Negative-strand RNA virus">negative-strand RNA viruses</a> in the family <i><a href="/wiki/Paramyxoviridae" title="Paramyxoviridae">Paramyxoviridae</a></i>, order <i><a href="/wiki/Mononegavirales" title="Mononegavirales">Mononegavirales</a></i> containing six established species,<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> and numerous others still under study.<sup id="cite_ref-Wu-2014_3-0" class="reference"><a href="#cite_note-Wu-2014-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Henipaviruses are naturally harboured by several species of small mammals, notably <a href="/wiki/Pteropus" title="Pteropus">pteropid</a> <a href="/wiki/Fruit_bat" class="mw-redirect" title="Fruit bat">fruit bats</a> (flying foxes), <a href="/wiki/Microbats" class="mw-redirect" title="Microbats">microbats</a> of several species,<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> and <a href="/wiki/Shrew" title="Shrew">shrews</a>.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Zhang-2022_6-0" class="reference"><a href="#cite_note-Zhang-2022-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> Henipaviruses are characterised by long <a href="/wiki/Genome" title="Genome">genomes</a> and a wide host range. Their recent emergence as <a href="/wiki/Zoonosis" title="Zoonosis">zoonotic</a> pathogens capable of causing illness and death in <a href="/wiki/Domestic_animal" class="mw-redirect" title="Domestic animal">domestic animals</a> and humans is a cause of concern.<sup id="cite_ref-sobrino5_7-0" class="reference"><a href="#cite_note-sobrino5-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p>In 2009, RNA sequences of three novel viruses in phylogenetic relationship to known henipaviruses were detected in African straw-colored fruit bats (<i><a href="/wiki/Eidolon_helvum" class="mw-redirect" title="Eidolon helvum">Eidolon helvum</a></i>) in <a href="/wiki/Ghana" title="Ghana">Ghana</a>. The finding of these novel henipaviruses outside Australia and Asia indicates that the region of potential endemicity of henipaviruses may be worldwide.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> These African henipaviruses are slowly being characterised.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p><p><i><a href="/wiki/Nipah_virus" title="Nipah virus">Nipah</a></i> and <i><a href="/wiki/Hendra_virus" title="Hendra virus">Hendra</a> henipaviruses</i> are both considered category C (USDA-HHS overlap) <a href="/wiki/Select_agent" title="Select agent">select agents</a>.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Structure">Structure</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Henipavirus&action=edit&section=1" title="Edit section: Structure"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Henipavirus_structure.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/86/Henipavirus_structure.svg/220px-Henipavirus_structure.svg.png" decoding="async" width="220" height="159" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/86/Henipavirus_structure.svg/330px-Henipavirus_structure.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/86/Henipavirus_structure.svg/440px-Henipavirus_structure.svg.png 2x" data-file-width="442" data-file-height="320" /></a><figcaption>Structure of henipaviruses</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Henipavirus_genome.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/31/Henipavirus_genome.png/220px-Henipavirus_genome.png" decoding="async" width="220" height="96" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/3/31/Henipavirus_genome.png 1.5x" data-file-width="325" data-file-height="142" /></a><figcaption>The henipavirus genome (3’ to 5’ orientation) and products of the P gene</figcaption></figure> <p>Henipavirions are <a href="/wiki/Pleomorphism_(microbiology)" title="Pleomorphism (microbiology)">pleomorphic</a> (variably shaped), ranging in size from 40 to 600 nm in diameter.<sup id="cite_ref-hyatt_12-0" class="reference"><a href="#cite_note-hyatt-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> They possess a <a href="/wiki/Lipid" title="Lipid">lipid</a> membrane overlying a shell of viral <a href="/wiki/Matrix_protein" class="mw-redirect" title="Matrix protein">matrix protein</a>. At the core is a single helical strand of genomic <a href="/wiki/RNA" title="RNA">RNA</a> tightly bound to N (<a href="/wiki/Nucleocapsid" class="mw-redirect" title="Nucleocapsid">nucleocapsid</a>) protein and associated with the L (large) and P (phosphoprotein) proteins, which provide <a href="/wiki/RNA_polymerase" title="RNA polymerase">RNA polymerase</a> activity during replication. </p><p>Embedded within the lipid membrane are spikes of F (fusion) protein trimers and G (attachment) protein tetramers. The function of the G protein (except in the case of MojV-G) is to attach the virus to the surface of a host cell via <a href="/wiki/EFNB2" class="mw-redirect" title="EFNB2">Ephrin B1, B2, or B3</a>, a family of highly <a href="/wiki/Conserved_sequence" title="Conserved sequence">conserved</a> mammalian proteins.<sup id="cite_ref-bonaparte_13-0" class="reference"><a href="#cite_note-bonaparte-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-negrete_14-0" class="reference"><a href="#cite_note-negrete-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> The structure of the attachment glycoprotein has been determined by X-ray crystallography.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> The F protein fuses the viral membrane with the host cell membrane, releasing the virion contents into the cell. It also causes infected cells to fuse with neighbouring cells to form large, multinucleated <a href="/wiki/Syncytia" class="mw-redirect" title="Syncytia">syncytia</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Genome">Genome</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Henipavirus&action=edit&section=2" title="Edit section: Genome"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Journal.pone.0199534.g001.A.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/37/Journal.pone.0199534.g001.A.png/220px-Journal.pone.0199534.g001.A.png" decoding="async" width="220" height="172" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/37/Journal.pone.0199534.g001.A.png/330px-Journal.pone.0199534.g001.A.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/37/Journal.pone.0199534.g001.A.png/440px-Journal.pone.0199534.g001.A.png 2x" data-file-width="2824" data-file-height="2202" /></a><figcaption>Nipah virus (NiV) replication cycle</figcaption></figure> <p>As all mononegaviral genomes, Hendra virus and Nipah virus genomes are non-segmented, single-stranded <a href="/wiki/Sense_(molecular_biology)" title="Sense (molecular biology)">negative-sense</a> RNA. Both genomes are 18.2 kb in length and contain six <a href="/wiki/Gene" title="Gene">genes</a> corresponding to six structural proteins.<sup id="cite_ref-wang_17-0" class="reference"><a href="#cite_note-wang-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p><p>In common with other members of the <i><a href="/wiki/Paramyxoviridae" title="Paramyxoviridae">Paramyxoviridae</a></i> family, the number of <a href="/wiki/Nucleotide" title="Nucleotide">nucleotides</a> in the henipavirus genome is a multiple of six, consistent with what is known as the '<a href="/wiki/Rule_of_six_(viruses)" title="Rule of six (viruses)">rule of six</a>'.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> Deviation from the rule of six, through mutation or incomplete genome synthesis, leads to inefficient viral replication, probably due to structural constraints imposed by the binding between the RNA and the N protein. </p><p>Three additional protein products are produced from the henipavirus P gene: V, W, and C. The V and W proteins are generated through an unusual process called <a href="/wiki/RNA_editing" title="RNA editing">RNA editing</a>. This specific process in henipaviruses involves the insertion of extra <a href="/wiki/Guanosine" title="Guanosine">guanosine</a> residues into the P gene <a href="/wiki/MRNA" class="mw-redirect" title="MRNA">mRNA</a> prior to <a href="/wiki/Translation_(genetics)" class="mw-redirect" title="Translation (genetics)">translation</a>. The addition of a single guanosine results in production of V, and the addition of two guanosines residues produces W.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> The C protein is not produced through RNA editing but instead by <a href="/wiki/Leaky_scanning" title="Leaky scanning">leaky scanning</a> of the host cell ribosome during translation of viral mRNA. P, V, and W possess an alternate <a href="/wiki/Open_reading_frame" title="Open reading frame">open reading frame</a> which results in production of C. P, V, W, and C are known to disrupt the host innate antiviral immune response through several different mechanisms.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> P, V, and W contain <a href="/wiki/STAT1" title="STAT1">STAT1</a> binding domains, and act as <a href="/wiki/Interferon" title="Interferon">interferon</a> antagonists by sequestering STAT1 in the nucleus and cytoplasm.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> The C protein controls the early pro-inflammatory response and is also known to promote the viral budding process via a <a href="/wiki/ESCRT" title="ESCRT">ESCRT</a>-dependent pathway.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Life_cycle">Life cycle</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Henipavirus&action=edit&section=3" title="Edit section: Life cycle"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Cell receptor ephrin-B2, which is located on epithelial cells around smaller arteries, neurons, and smooth muscle cells, is targeted by the viral protein G.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> Once the protein G binds to ephrin-B2, the viral protein F facilitates fusion with the host cell membrane and releases viral RNA into the host cell cytoplasm.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup> Upon entry, transcription of viral mRNA takes place using the viral RNA as a template. This process is started and stopped by the polymerase complex. Viral proteins are gathering in the cell as transcription occurs until the polymerase complex stops transcription and starts genome replication. Transcription of the viral RNA makes positive sense strands of RNA, which are then used as templates to make more negative sense viral RNA . Genome replication is halted before the viral particles can assemble to make a virion. Once the cell membrane is ready, new virions exit the host cell through budding.<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Vaccine">Vaccine</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Henipavirus&action=edit&section=4" title="Edit section: Vaccine"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Henipaviruses have high mortality rates in mammalian hosts, both human and animal. Because of this, there is a need for immunization against HeV and NiV. The <a href="/wiki/World_Health_Organization" title="World Health Organization">World Health Organization</a> has classified henipaviral agents as R&D Blueprint Priority Pathogens, indicating that they pose a significant risk due to their epidemic potential.<sup id="cite_ref-28" class="reference"><a href="#cite_note-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> The broad species tropism of NiV and HeV have resulted in mortality in livestock species in addition to humans, and as a result veterinary vaccines are in various stages of development or licensure. EquiVac HeV, a veterinary vaccine for horses was licensed in Australia in 2012.<sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup> A number of experimental vaccines designed for humans are in preclinical development, but none have yet been licensed. A soluble HeV attachment glycoprotein vaccine designed to protect against NiV completed a <a href="/wiki/Phase_I_clinical_trial" class="mw-redirect" title="Phase I clinical trial">phase I clinical trial</a> in November 2022, but results have not yet been published.<sup id="cite_ref-31" class="reference"><a href="#cite_note-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup> </p><p>The primary mechanism of protection against NiV and HeV induced by vaccination is thought to be neutralizing antibodies.<sup id="cite_ref-32" class="reference"><a href="#cite_note-32"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup> However, a number of preclinical vaccine studies in animal models of disease have identified that the cell-mediated immune response including CD8+ and CD4+ T-cells may play a role in protection.<sup id="cite_ref-33" class="reference"><a href="#cite_note-33"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Causes_of_emergence">Causes of emergence</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Henipavirus&action=edit&section=5" title="Edit section: Causes of emergence"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The emergence of henipaviruses parallels the emergence of other <a href="/wiki/Zoonosis" title="Zoonosis">zoonotic</a> viruses in recent decades. <a href="/wiki/SARS_coronavirus" class="mw-redirect" title="SARS coronavirus">SARS coronavirus</a>, <a href="/wiki/Australian_bat_lyssavirus" title="Australian bat lyssavirus">Australian bat lyssavirus</a>, <a href="/wiki/Menangle_virus" class="mw-redirect" title="Menangle virus">Menangle virus</a>, <a href="/wiki/Marburg_virus" title="Marburg virus">Marburg virus</a>, COVID 19 and possibly <a href="/wiki/Ebola" title="Ebola">Ebola</a> viruses are also harboured by bats, and are capable of infecting a variety of other species. The emergence of each of these viruses has been linked to an increase in contact between bats and humans, sometimes involving an intermediate domestic animal host. The increased contact is driven both by human encroachment into the bats' territory (in the case of Nipah, specifically pigpens in said territory) and by movement of bats towards human populations due to changes in food distribution and loss of habitat. </p><p>There is evidence that habitat loss for flying foxes, both in South Asia and Australia (particularly along the east coast) as well as encroachment of human dwellings and agriculture into the remaining habitats, is creating greater overlap of human and flying fox distributions.<sup id="cite_ref-34" class="reference"><a href="#cite_note-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Taxonomy">Taxonomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Henipavirus&action=edit&section=6" title="Edit section: Taxonomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <table class="sortable wikitable"> <caption>Genus <i>Henipavirus</i>: species and their viruses<sup id="cite_ref-35" class="reference"><a href="#cite_note-35"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup> </caption> <tbody><tr> <td><b>Genus</b> </td> <td><b>Species</b> </td> <td><b>Virus (Abbreviation)</b> </td></tr> <tr valign="TOP"> <td rowspan="6"><i>Henipavirus</i> </td> <td rowspan="1"><i><a href="/wiki/Cedar_henipavirus" class="mw-redirect" title="Cedar henipavirus">Cedar henipavirus</a></i> </td> <td rowspan="1"><a href="/wiki/Cedar_virus" title="Cedar virus">Cedar virus</a> (CedV) </td></tr> <tr valign="TOP"> <td rowspan="1"><i><a href="/wiki/Kumasi_virus" class="mw-redirect" title="Kumasi virus">Ghanaian bat henipavirus</a></i> </td> <td rowspan="1"><a href="/wiki/Kumasi_virus" class="mw-redirect" title="Kumasi virus">Kumasi virus</a> (KV) </td></tr> <tr valign="TOP"> <td><i><a href="/wiki/Hendra_henipavirus" class="mw-redirect" title="Hendra henipavirus">Hendra henipavirus</a></i> </td> <td rowspan="1"><a href="/wiki/Hendra_virus" title="Hendra virus">Hendra virus</a> (HeV) </td></tr> <tr> <td><i><a href="/wiki/Mojiang_henipavirus" class="mw-redirect" title="Mojiang henipavirus">Mojiang henipavirus</a></i> </td> <td rowspan="1"><a href="/wiki/M%C3%B2ji%C4%81ng_virus" title="Mòjiāng virus">Mòjiāng virus</a> (MojV)<sup id="cite_ref-Wu-2014_3-1" class="reference"><a href="#cite_note-Wu-2014-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </td></tr> <tr valign="TOP"> <td rowspan="1"><i><a href="/wiki/Nipah_henipavirus" class="mw-redirect" title="Nipah henipavirus">Nipah henipavirus</a></i> </td> <td rowspan="1"><a href="/wiki/Nipah_virus" title="Nipah virus">Nipah virus</a> (NiV) </td></tr> <tr valign="TOP"> <td rowspan="1"><i><a href="/wiki/Langya_henipavirus" title="Langya henipavirus">Langya henipavirus</a></i> </td> <td rowspan="1"><a href="/wiki/Langya_virus" class="mw-redirect" title="Langya virus">Langya virus</a> (LayV)<sup id="cite_ref-Zhang-2022_6-1" class="reference"><a href="#cite_note-Zhang-2022-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-36" class="reference"><a href="#cite_note-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </td></tr></tbody></table> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Henipavirus&action=edit&section=7" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1266661725">.mw-parser-output .portalbox{padding:0;margin:0.5em 0;display:table;box-sizing:border-box;max-width:175px;list-style:none}.mw-parser-output .portalborder{border:1px solid 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no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output 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href="/wiki/Template_talk:Zoonotic_viral_diseases" title="Template talk:Zoonotic viral diseases"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Zoonotic_viral_diseases" title="Special:EditPage/Template:Zoonotic viral diseases"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Zoonotic_viral_diseases_(A80–B34,_042–079)127" style="font-size:114%;margin:0 4em"><a href="/wiki/Zoonosis" title="Zoonosis">Zoonotic</a> <a href="/wiki/Viral_disease" title="Viral disease">viral diseases</a> (A80–B34, <a href="/wiki/042%E2%80%93079" class="mw-redirect" title="042–079">042–079</a>)</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Arthropod" title="Arthropod">Arthropod<br />-borne</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Mosquito-borne_disease" title="Mosquito-borne disease">Mosquito<br />-borne</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bunyavirales" title="Bunyavirales">Bunyavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arbovirus_encephalitis" class="mw-redirect" title="Arbovirus encephalitis">Arbovirus encephalitides</a>: <a href="/wiki/Batai_virus" class="mw-redirect" title="Batai virus">Batai virus</a> <ul><li>BATV</li></ul></li> <li><a href="/wiki/Bwamba_orthobunyavirus" title="Bwamba orthobunyavirus">Bwamba Fever</a> <ul><li>BWAV</li></ul></li> <li><a href="/wiki/California_encephalitis_orthobunyavirus" title="California encephalitis orthobunyavirus">California encephalitis</a> <ul><li>CEV</li></ul></li> <li><a href="/wiki/Jamestown_Canyon_encephalitis" title="Jamestown Canyon encephalitis">Jamestown Canyon encephalitis</a></li> <li><a href="/wiki/La_Crosse_encephalitis" title="La Crosse encephalitis">La Crosse encephalitis</a> <ul><li>LACV</li></ul></li> <li><a href="/wiki/Tahyna_virus" class="mw-redirect" title="Tahyna virus">Tahyna virus</a> <ul><li>TAHV</li></ul></li> <li><a href="/wiki/Tete_virus" class="mw-redirect" title="Tete virus">Tete virus</a></li></ul> <ul><li><a href="/wiki/Viral_hemorrhagic_fever" title="Viral hemorrhagic fever">Viral hemorrhagic fevers</a>: <a href="/wiki/Bunyamwera_orthobunyavirus" title="Bunyamwera orthobunyavirus">Bunyamwera fever</a> <ul><li>BUNV</li></ul></li> <li><a href="/wiki/Ngari_virus" title="Ngari virus">Ngari virus</a> <ul><li>NRIV</li></ul></li> <li><a href="/wiki/Rift_Valley_fever" title="Rift Valley fever">Rift Valley fever</a> <ul><li>RVFV</li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Flaviviridae" title="Flaviviridae">Flaviviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arbovirus_encephalitis" class="mw-redirect" title="Arbovirus encephalitis">Arbovirus encephalitides</a>: <a href="/wiki/Japanese_encephalitis" title="Japanese encephalitis">Japanese encephalitis</a> <ul><li><a href="/wiki/Japanese_encephalitis_virus" class="mw-redirect" title="Japanese encephalitis virus">JEV</a></li></ul></li> <li><a href="/wiki/Murray_Valley_encephalitis_virus" title="Murray Valley encephalitis virus">Australian encephalitis</a> <ul><li><a href="/wiki/Murray_Valley_encephalitis_virus" title="Murray Valley encephalitis virus">MVEV</a></li> <li><a href="/wiki/Kunjin_virus" title="Kunjin virus">KUNV</a></li></ul></li> <li><a href="/wiki/Saint_Louis_encephalitis" title="Saint Louis encephalitis">Saint Louis encephalitis</a> <ul><li>SLEV</li></ul></li> <li><a href="/wiki/Usutu_virus" title="Usutu virus">Usutu virus</a> <ul><li>USUV</li></ul></li> <li><a href="/wiki/West_Nile_fever" title="West Nile fever">West Nile fever</a> <ul><li><a href="/wiki/West_Nile_virus" title="West Nile virus">WNV</a></li></ul></li></ul> <ul><li><a href="/wiki/Viral_hemorrhagic_fever" title="Viral hemorrhagic fever">Viral hemorrhagic fevers</a>: <a href="/wiki/Dengue_fever" title="Dengue fever">Dengue fever</a> <ul><li><a href="/wiki/Dengue_virus" title="Dengue virus">DENV-1-4</a></li></ul></li> <li><a href="/wiki/Yellow_fever" title="Yellow fever">Yellow fever</a> <ul><li><a href="/wiki/Yellow_fever_virus" class="mw-redirect" title="Yellow fever virus">YFV</a></li></ul></li> <li><a href="/wiki/Zika_fever" title="Zika fever">Zika fever</a> <ul><li><a href="/wiki/ZIKV" class="mw-redirect" title="ZIKV">ZIKV</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Togaviridae" class="mw-redirect" title="Togaviridae">Togaviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arbovirus_encephalitis" class="mw-redirect" title="Arbovirus encephalitis">Arbovirus encephalitides</a>: <a href="/wiki/Eastern_equine_encephalitis" title="Eastern equine encephalitis">Eastern equine encephalomyelitis</a> <ul><li>EEEV</li></ul></li> <li><a href="/wiki/Western_equine_encephalitis_virus" title="Western equine encephalitis virus">Western equine encephalomyelitis</a> <ul><li>WEEV</li></ul></li> <li><a href="/wiki/Venezuelan_equine_encephalitis_virus" title="Venezuelan equine encephalitis virus">Venezuelan equine encephalomyelitis</a> <ul><li>VEEV</li></ul></li> <li><a href="/wiki/Chikungunya" title="Chikungunya">Chikungunya</a> <ul><li><a href="/wiki/Chikungunya_virus" class="mw-redirect" title="Chikungunya virus">CHIKV</a></li></ul></li> <li><a href="/wiki/O%27nyong%27nyong_virus" title="O'nyong'nyong virus">O'nyong'nyong fever</a> <ul><li>ONNV</li></ul></li> <li><a href="/wiki/Pogosta_disease" title="Pogosta disease">Pogosta disease</a> <ul><li><a href="/wiki/Sindbis_virus" title="Sindbis virus">Sindbis virus</a></li></ul></li> <li><a href="/wiki/Ross_River_fever" title="Ross River fever">Ross River fever</a> <ul><li><a href="/wiki/Ross_River_virus" title="Ross River virus">RRV</a></li></ul></li> <li><a href="/wiki/Semliki_Forest_virus" title="Semliki Forest virus">Semliki Forest virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Reoviridae" class="mw-redirect" title="Reoviridae">Reoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Banna_virus" title="Banna virus">Banna virus encephalitis</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tick-borne_disease#Viral" title="Tick-borne disease">Tick<br />-borne</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bunyavirales" title="Bunyavirales">Bunyavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Viral_hemorrhagic_fever" title="Viral hemorrhagic fever">Viral hemorrhagic fevers</a>: <a href="/wiki/Crimean%E2%80%93Congo_hemorrhagic_fever" title="Crimean–Congo hemorrhagic fever">Crimean–Congo hemorrhagic fever</a> (CCHFV)</li> <li><a href="/wiki/Heartland_virus" class="mw-redirect" title="Heartland virus">Heartland virus</a></li> <li><a href="/wiki/Severe_fever_with_thrombocytopenia_syndrome" title="Severe fever with thrombocytopenia syndrome">Severe fever with thrombocytopenia syndrome</a> (<a href="/wiki/Huaiyangshan_banyangvirus" class="mw-redirect" title="Huaiyangshan banyangvirus">Huaiyangshan banyangvirus</a>)</li> <li><a href="/wiki/Tete_virus" class="mw-redirect" title="Tete virus">Tete virus</a></li> <li><a href="/wiki/Arbovirus_encephalitis" class="mw-redirect" title="Arbovirus encephalitis">Arbovirus encephalitides</a>: <a href="/wiki/Bhanja_virus" title="Bhanja virus">Bhanja virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Flaviviridae" title="Flaviviridae">Flaviviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arbovirus_encephalitis" class="mw-redirect" title="Arbovirus encephalitis">Arbovirus encephalitides</a>: <a href="/wiki/Tick-borne_encephalitis" title="Tick-borne encephalitis">Tick-borne encephalitis</a> <ul><li><a href="/wiki/Tick-borne_encephalitis_virus" title="Tick-borne encephalitis virus">TBEV</a></li></ul></li> <li><a href="/wiki/Powassan_encephalitis" title="Powassan encephalitis">Powassan encephalitis</a> <ul><li><a href="/wiki/Powassan_virus" title="Powassan virus">POWV</a></li></ul></li> <li><a href="/wiki/Viral_hemorrhagic_fever" title="Viral hemorrhagic fever">Viral hemorrhagic fevers</a>: <a href="/wiki/Omsk_hemorrhagic_fever" title="Omsk hemorrhagic fever">Omsk hemorrhagic fever</a> <ul><li>OHFV</li></ul></li> <li><a href="/wiki/Kyasanur_Forest_disease" title="Kyasanur Forest disease">Kyasanur Forest disease</a> <ul><li>KFDV</li> <li><a href="/wiki/Alkhurma_virus" title="Alkhurma virus">AHFV</a></li></ul></li> <li><a href="/wiki/Langat_virus" title="Langat virus">Langat virus</a> <ul><li>LGTV</li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthomyxoviridae" title="Orthomyxoviridae">Orthomyxoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bourbon_virus" title="Bourbon virus">Bourbon virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Reoviridae" class="mw-redirect" title="Reoviridae">Reoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Colorado_tick_fever" title="Colorado tick fever">Colorado tick fever</a> <ul><li>CTFV</li></ul></li> <li><a href="/wiki/Kemerovo_tickborne_viral_fever" title="Kemerovo tickborne viral fever">Kemerovo tickborne viral fever</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Sandfly#Viruses" title="Sandfly">Sandfly<br />-borne</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bunyavirales" title="Bunyavirales">Bunyavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Adria_virus" title="Adria virus">Adria virus</a> (ADRV)</li> <li><a href="/wiki/Oropouche_fever" title="Oropouche fever">Oropouche fever</a> <ul><li><a href="/wiki/Oropouche_virus" class="mw-redirect" title="Oropouche virus">Oropouche virus</a></li></ul></li> <li><a href="/wiki/Pappataci_fever" title="Pappataci fever">Pappataci fever</a> <ul><li><a href="/wiki/Toscana_virus" class="mw-redirect" title="Toscana virus">Toscana virus</a></li></ul></li> <li><a href="/wiki/Sandfly_fever_Naples_phlebovirus" class="mw-redirect" title="Sandfly fever Naples phlebovirus">Sandfly fever Naples virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Rhabdoviridae" title="Rhabdoviridae">Rhabdoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Chandipura_vesiculovirus" title="Chandipura vesiculovirus">Chandipura virus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Mammal" title="Mammal">Mammal<br />-borne</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Rodent" title="Rodent">Rodent<br />-borne</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Arenavirus" title="Arenavirus">Arenaviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Viral_hemorrhagic_fever" title="Viral hemorrhagic fever">Viral hemorrhagic fevers</a>: <a href="/wiki/Lassa_fever" title="Lassa fever">Lassa fever</a> <ul><li><a href="/wiki/Lassa_virus" class="mw-redirect" title="Lassa virus">LASV</a></li></ul></li> <li><a href="/wiki/Venezuelan_hemorrhagic_fever" title="Venezuelan hemorrhagic fever">Venezuelan hemorrhagic fever</a> <ul><li>GTOV</li></ul></li> <li><a href="/wiki/Argentine_hemorrhagic_fever" title="Argentine hemorrhagic fever">Argentine hemorrhagic fever</a> <ul><li><a href="/wiki/Argentinian_mammarenavirus" title="Argentinian mammarenavirus">JUNV</a></li></ul></li> <li><a href="/wiki/Brazilian_hemorrhagic_fever" title="Brazilian hemorrhagic fever">Brazilian hemorrhagic fever</a> <ul><li><a href="/wiki/Brazilian_hemorrhagic_fever" title="Brazilian hemorrhagic fever">SABV</a></li></ul></li> <li><a href="/wiki/Bolivian_hemorrhagic_fever" title="Bolivian hemorrhagic fever">Bolivian hemorrhagic fever</a> <ul><li>MACV</li></ul></li> <li><a href="/wiki/Lujo_mammarenavirus" title="Lujo mammarenavirus">LUJV</a></li> <li><a href="/wiki/Chapare_virus" class="mw-redirect" title="Chapare virus">CHPV</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bunyavirales" title="Bunyavirales">Bunyavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hantavirus_hemorrhagic_fever_with_renal_syndrome" title="Hantavirus hemorrhagic fever with renal syndrome">Hemorrhagic fever with renal syndrome</a> <ul><li><a href="/wiki/Dobrava-Belgrade_orthohantavirus" class="mw-redirect" title="Dobrava-Belgrade orthohantavirus">DOBV</a></li> <li><a href="/wiki/Hantaan_orthohantavirus" class="mw-redirect" title="Hantaan orthohantavirus">HTNV</a></li> <li><a href="/wiki/Puumala_orthohantavirus" class="mw-redirect" title="Puumala orthohantavirus">PUUV</a></li> <li><a href="/wiki/Seoul_orthohantavirus" class="mw-redirect" title="Seoul orthohantavirus">SEOV</a></li> <li><a href="/wiki/Amur_virus" title="Amur virus">AMRV</a></li> <li><a href="/wiki/Thailand_virus" title="Thailand virus">THAIV</a></li></ul></li> <li><a href="/wiki/Hantavirus_pulmonary_syndrome" title="Hantavirus pulmonary syndrome">Hantavirus pulmonary syndrome</a> <ul><li><a href="/wiki/Andes_orthohantavirus" class="mw-redirect" title="Andes orthohantavirus">ANDV</a></li> <li><a href="/wiki/Sin_Nombre_orthohantavirus" class="mw-redirect" title="Sin Nombre orthohantavirus">SNV</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Herpesviridae" title="Herpesviridae">Herpesviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Murid_gammaherpesvirus_4" title="Murid gammaherpesvirus 4">Murid gammaherpesvirus 4</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Bat#Disease_transmission" title="Bat">Bat<br />-borne</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Filoviridae" title="Filoviridae">Filoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ebola_virus_disease" class="mw-redirect" title="Ebola virus disease">Ebola virus disease</a> <ul><li><a href="/wiki/Bundibugyo_virus" class="mw-redirect" title="Bundibugyo virus">BDBV</a></li> <li><a href="/wiki/Sudan_virus" class="mw-redirect" title="Sudan virus">SUDV</a></li> <li><a href="/wiki/Ta%C3%AF_Forest_virus" class="mw-redirect" title="Taï Forest virus">TAFV</a></li> <li><a href="/wiki/Zaire_ebolavirus" title="Zaire ebolavirus">ZEBOV</a></li></ul></li> <li><a href="/wiki/Marburg_virus_disease" title="Marburg virus disease">Marburg virus disease</a> <ul><li><a href="/wiki/Marburg_virus" title="Marburg virus">MARV</a></li> <li><a href="/wiki/Ravn_virus" title="Ravn virus">RAVV</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Rhabdoviridae" title="Rhabdoviridae">Rhabdoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rabies" title="Rabies">Rabies</a> <ul><li><a href="/wiki/Australian_bat_lyssavirus" title="Australian bat lyssavirus">ABLV</a></li> <li><a href="/wiki/Mokola_lyssavirus" title="Mokola lyssavirus">MOKV</a></li> <li><a href="/wiki/Duvenhage_lyssavirus" title="Duvenhage lyssavirus">DUVV</a></li> <li><a href="/wiki/Lagos_bat_lyssavirus" title="Lagos bat lyssavirus">LBV</a></li></ul></li> <li><a href="/wiki/Chandipura_vesiculovirus" title="Chandipura vesiculovirus">CHPV</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Paramyxoviridae" title="Paramyxoviridae">Paramyxoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Henipavirus encephalitis</a> <ul><li><a href="/wiki/Hendra_virus" title="Hendra virus">HeV</a></li> <li><a href="/wiki/Nipah_virus" title="Nipah virus">NiV</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Coronaviridae" title="Coronaviridae">Coronaviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Middle_East_respiratory_syndrome%E2%80%93related_coronavirus" class="mw-redirect" title="Middle East respiratory syndrome–related coronavirus">MERS-CoV</a></li> <li><a href="/wiki/SARS-related_coronavirus" title="SARS-related coronavirus">SARS-related coronavirus</a> <ul><li><a href="/wiki/Severe_acute_respiratory_syndrome_coronavirus" class="mw-redirect" title="Severe acute respiratory syndrome coronavirus">SARS-CoV</a></li> <li><a href="/wiki/SARS-CoV-2" title="SARS-CoV-2">SARS-CoV-2</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Primate" title="Primate">Primate<br />-borne</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Herpesviridae" title="Herpesviridae">Herpesviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/B_virus" title="B virus">B virus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Retrovirus" title="Retrovirus">Retroviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Simian_foamy_virus" class="mw-redirect" title="Simian foamy virus">Simian foamy virus</a></li> <li><a href="/wiki/Human_T-lymphotropic_virus_1" title="Human T-lymphotropic virus 1">HTLV-1</a></li> <li><a href="/wiki/Human_T-lymphotropic_virus_2" title="Human T-lymphotropic virus 2">HTLV-2</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Poxviridae" title="Poxviridae">Poxviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Tanapox" title="Tanapox">Tanapox</a></li> <li><a href="/wiki/Yaba_monkey_tumor_virus" title="Yaba monkey tumor virus">Yaba monkey tumor virus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Multiple<br />vectors</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Rhabdoviridae" title="Rhabdoviridae">Rhabdoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rabies" title="Rabies">Rabies</a> <ul><li><a href="/wiki/Rabies_virus" title="Rabies virus">RABV</a></li></ul></li> <li><a href="/wiki/Mokola_lyssavirus" title="Mokola lyssavirus">Mokola virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Poxviridae" title="Poxviridae">Poxviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Monkeypox" class="mw-redirect" title="Monkeypox">Monkeypox</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers1426" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers1426" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Henipavirus</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q1481398" class="extiw" title="wikidata:Q1481398">Q1481398</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Henipavirus" class="extiw" title="wikispecies:Henipavirus">Henipavirus</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/62VCD">62VCD</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/80652">80652</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/10589448">10589448</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=1014047">1014047</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=260964">260964</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.marinespecies.org/aphia.php?p=taxdetails&id=600284">600284</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r1038841319">.mw-parser-output .tooltip-dotted{border-bottom:1px dotted;cursor:help}</style><link 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typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q1481398#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" 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