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Chitin - Wikipedia
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<li id="toc-Plants" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Plants"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.2</span> <span>Plants</span> </div> </a> <ul id="toc-Plants-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Fossil_record" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Fossil_record"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Fossil record</span> </div> </a> <ul id="toc-Fossil_record-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Uses" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Uses"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Uses</span> </div> </a> <button aria-controls="toc-Uses-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span 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</div> </a> <ul id="toc-Research-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" title="Table of Contents" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Chitin</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 68 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-68" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">68 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%83%D8%A7%D9%8A%D8%AA%D9%8A%D9%86" title="كايتين – Arabic" lang="ar" hreflang="ar" data-title="كايتين" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Quitina" title="Quitina – Asturian" lang="ast" hreflang="ast" data-title="Quitina" data-language-autonym="Asturianu" data-language-local-name="Asturian" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Xitin" title="Xitin – Azerbaijani" lang="az" hreflang="az" data-title="Xitin" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%95%E0%A6%BE%E0%A6%87%E0%A6%9F%E0%A6%BF%E0%A6%A8" title="কাইটিন – Bangla" lang="bn" hreflang="bn" data-title="কাইটিন" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ba mw-list-item"><a href="https://ba.wikipedia.org/wiki/%D0%A5%D0%B8%D1%82%D0%B8%D0%BD" title="Хитин – Bashkir" lang="ba" hreflang="ba" data-title="Хитин" data-language-autonym="Башҡортса" data-language-local-name="Bashkir" class="interlanguage-link-target"><span>Башҡортса</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A5%D1%96%D1%86%D1%96%D0%BD" title="Хіцін – Belarusian" lang="be" hreflang="be" data-title="Хіцін" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-be-x-old mw-list-item"><a href="https://be-tarask.wikipedia.org/wiki/%D0%A5%D1%96%D1%82%D1%8B%D0%BD" title="Хітын – Belarusian (Taraškievica orthography)" lang="be-tarask" hreflang="be-tarask" data-title="Хітын" data-language-autonym="Беларуская (тарашкевіца)" data-language-local-name="Belarusian (Taraškievica orthography)" class="interlanguage-link-target"><span>Беларуская (тарашкевіца)</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A5%D0%B8%D1%82%D0%B8%D0%BD" title="Хитин – Bulgarian" lang="bg" hreflang="bg" data-title="Хитин" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Hitin" title="Hitin – Bosnian" lang="bs" hreflang="bs" data-title="Hitin" data-language-autonym="Bosanski" data-language-local-name="Bosnian" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Quitina" title="Quitina – Catalan" lang="ca" hreflang="ca" data-title="Quitina" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Chitin" title="Chitin – Czech" lang="cs" hreflang="cs" data-title="Chitin" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-cy mw-list-item"><a href="https://cy.wikipedia.org/wiki/Citin" title="Citin – Welsh" lang="cy" hreflang="cy" data-title="Citin" data-language-autonym="Cymraeg" data-language-local-name="Welsh" class="interlanguage-link-target"><span>Cymraeg</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Kitin" title="Kitin – Danish" lang="da" hreflang="da" data-title="Kitin" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Chitin" title="Chitin – German" lang="de" hreflang="de" data-title="Chitin" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Kitiin" title="Kitiin – Estonian" lang="et" hreflang="et" data-title="Kitiin" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%A7%CE%B9%CF%84%CE%AF%CE%BD%CE%B7" title="Χιτίνη – Greek" lang="el" hreflang="el" data-title="Χιτίνη" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Quitina" title="Quitina – Spanish" lang="es" hreflang="es" data-title="Quitina" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/%C4%A4itino" title="Ĥitino – Esperanto" lang="eo" hreflang="eo" data-title="Ĥitino" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Kitina" title="Kitina – Basque" lang="eu" hreflang="eu" data-title="Kitina" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DA%A9%DB%8C%D8%AA%DB%8C%D9%86" title="کیتین – Persian" lang="fa" hreflang="fa" data-title="کیتین" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Chitine" title="Chitine – French" lang="fr" hreflang="fr" data-title="Chitine" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Citin" title="Citin – Irish" lang="ga" hreflang="ga" data-title="Citin" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Quitina" title="Quitina – Galician" lang="gl" hreflang="gl" data-title="Quitina" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%82%A4%ED%8B%B4" title="키틴 – Korean" lang="ko" hreflang="ko" data-title="키틴" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D4%BD%D5%AB%D5%BF%D5%AB%D5%B6" title="Խիտին – Armenian" lang="hy" hreflang="hy" data-title="Խիտին" data-language-autonym="Հայերեն" data-language-local-name="Armenian" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%95%E0%A4%BE%E0%A4%87%E0%A4%9F%E0%A4%BF%E0%A4%A8" title="काइटिन – Hindi" lang="hi" hreflang="hi" data-title="काइटिन" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Hitin" title="Hitin – Croatian" lang="hr" hreflang="hr" data-title="Hitin" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Chitino" title="Chitino – Ido" lang="io" hreflang="io" data-title="Chitino" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Kitin" title="Kitin – Indonesian" lang="id" hreflang="id" data-title="Kitin" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/K%C3%ADt%C3%ADn" title="Kítín – Icelandic" lang="is" hreflang="is" data-title="Kítín" data-language-autonym="Íslenska" data-language-local-name="Icelandic" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Chitina" title="Chitina – Italian" lang="it" hreflang="it" data-title="Chitina" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%9B%D7%99%D7%98%D7%99%D7%9F" title="כיטין – Hebrew" lang="he" hreflang="he" data-title="כיטין" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-jv mw-list-item"><a href="https://jv.wikipedia.org/wiki/Kitin" title="Kitin – Javanese" lang="jv" hreflang="jv" data-title="Kitin" data-language-autonym="Jawa" data-language-local-name="Javanese" class="interlanguage-link-target"><span>Jawa</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%95%E0%B3%88%E0%B2%9F%E0%B2%BF%E0%B2%A8%E0%B3%8D" title="ಕೈಟಿನ್ – Kannada" lang="kn" hreflang="kn" data-title="ಕೈಟಿನ್" data-language-autonym="ಕನ್ನಡ" data-language-local-name="Kannada" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A5%D0%B8%D1%82%D0%B8%D0%BD" title="Хитин – Kazakh" lang="kk" hreflang="kk" data-title="Хитин" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Khitini" title="Khitini – Swahili" lang="sw" hreflang="sw" data-title="Khitini" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ht mw-list-item"><a href="https://ht.wikipedia.org/wiki/Kitin" title="Kitin – Haitian Creole" lang="ht" hreflang="ht" data-title="Kitin" data-language-autonym="Kreyòl ayisyen" data-language-local-name="Haitian Creole" class="interlanguage-link-target"><span>Kreyòl ayisyen</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Chitinum" title="Chitinum – Latin" lang="la" hreflang="la" data-title="Chitinum" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Hit%C4%ABns" title="Hitīns – Latvian" lang="lv" hreflang="lv" data-title="Hitīns" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Chitinas" title="Chitinas – Lithuanian" lang="lt" hreflang="lt" data-title="Chitinas" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Kitin" title="Kitin – Hungarian" lang="hu" hreflang="hu" data-title="Kitin" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Kitin" title="Kitin – Malay" lang="ms" hreflang="ms" data-title="Kitin" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Chitine" title="Chitine – Dutch" lang="nl" hreflang="nl" data-title="Chitine" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%AD%E3%83%81%E3%83%B3" title="キチン – Japanese" lang="ja" hreflang="ja" data-title="キチン" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Kitin" title="Kitin – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Kitin" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Kitin" title="Kitin – Norwegian Nynorsk" lang="nn" hreflang="nn" data-title="Kitin" data-language-autonym="Norsk nynorsk" data-language-local-name="Norwegian Nynorsk" class="interlanguage-link-target"><span>Norsk nynorsk</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Xitin" title="Xitin – Uzbek" lang="uz" hreflang="uz" data-title="Xitin" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Chityna" title="Chityna – Polish" lang="pl" hreflang="pl" data-title="Chityna" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Quitina" title="Quitina – Portuguese" lang="pt" hreflang="pt" data-title="Quitina" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Chitin%C4%83" title="Chitină – Romanian" lang="ro" hreflang="ro" data-title="Chitină" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A5%D0%B8%D1%82%D0%B8%D0%BD" title="Хитин – Russian" lang="ru" hreflang="ru" data-title="Хитин" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Chitin" title="Chitin – Simple English" lang="en-simple" hreflang="en-simple" data-title="Chitin" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Chit%C3%ADn" title="Chitín – Slovak" lang="sk" hreflang="sk" data-title="Chitín" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Hitin" title="Hitin – Slovenian" lang="sl" hreflang="sl" data-title="Hitin" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%A5%D0%B8%D1%82%D0%B8%D0%BD" title="Хитин – Serbian" lang="sr" hreflang="sr" data-title="Хитин" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Hitin" title="Hitin – Serbo-Croatian" lang="sh" hreflang="sh" data-title="Hitin" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbo-Croatian" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-su mw-list-item"><a href="https://su.wikipedia.org/wiki/Kitin" title="Kitin – Sundanese" lang="su" hreflang="su" data-title="Kitin" data-language-autonym="Sunda" data-language-local-name="Sundanese" class="interlanguage-link-target"><span>Sunda</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Kitiini" title="Kitiini – Finnish" lang="fi" hreflang="fi" data-title="Kitiini" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Kitin" title="Kitin – Swedish" lang="sv" hreflang="sv" data-title="Kitin" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%95%E0%AF%88%E0%AE%9F%E0%AF%8D%E0%AE%9F%E0%AE%BF%E0%AE%A9%E0%AF%8D" title="கைட்டின் – Tamil" lang="ta" hreflang="ta" data-title="கைட்டின்" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B9%84%E0%B8%84%E0%B8%97%E0%B8%B4%E0%B8%99" title="ไคทิน – Thai" lang="th" hreflang="th" data-title="ไคทิน" data-language-autonym="ไทย" data-language-local-name="Thai" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Kitin" title="Kitin – Turkish" lang="tr" hreflang="tr" data-title="Kitin" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A5%D1%96%D1%82%D0%B8%D0%BD" title="Хітин – Ukrainian" lang="uk" hreflang="uk" data-title="Хітин" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%B5%D9%84%D8%A8%DB%8C%D9%86" title="صلبین – Urdu" lang="ur" hreflang="ur" data-title="صلبین" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Kitin" title="Kitin – Vietnamese" lang="vi" hreflang="vi" data-title="Kitin" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E7%94%B2%E5%A3%B3%E7%B4%A0" title="甲壳素 – Wu" lang="wuu" hreflang="wuu" data-title="甲壳素" data-language-autonym="吴语" data-language-local-name="Wu" class="interlanguage-link-target"><span>吴语</span></a></li><li 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</div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Long-chain polymer of a N-acetylglucosamine</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">For other uses, see <a href="/wiki/Chitin_(disambiguation)" class="mw-disambig" title="Chitin (disambiguation)">Chitin (disambiguation)</a>.</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">Not to be confused with <a href="/wiki/Chiton" title="Chiton">chiton</a> or <a href="/wiki/Keratin" title="Keratin">keratin</a>.</div> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Chitin.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/13/Chitin.svg/250px-Chitin.svg.png" decoding="async" width="250" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/13/Chitin.svg/375px-Chitin.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/13/Chitin.svg/500px-Chitin.svg.png 2x" data-file-width="512" data-file-height="302" /></a><figcaption>Structure of the chitin molecule, showing two of the <a href="/wiki/N-Acetylglucosamine" title="N-Acetylglucosamine"><i>N</i>-acetylglucosamine</a> units that repeat to form long chains in β-(1→4)-linkage.</figcaption></figure> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Haworth_projection_of_chitin.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7e/Haworth_projection_of_chitin.svg/250px-Haworth_projection_of_chitin.svg.png" decoding="async" width="250" height="176" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7e/Haworth_projection_of_chitin.svg/375px-Haworth_projection_of_chitin.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7e/Haworth_projection_of_chitin.svg/500px-Haworth_projection_of_chitin.svg.png 2x" data-file-width="512" data-file-height="360" /></a><figcaption><a href="/wiki/Haworth_projection" title="Haworth projection">Haworth projection</a> of the chitin molecule.</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Glanzkaefer.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8b/Glanzkaefer.jpg/220px-Glanzkaefer.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8b/Glanzkaefer.jpg/330px-Glanzkaefer.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8b/Glanzkaefer.jpg/440px-Glanzkaefer.jpg 2x" data-file-width="800" data-file-height="600" /></a><figcaption>A close-up of the wing of a <a href="/wiki/Leafhopper" title="Leafhopper">leafhopper</a>; the wing is composed of chitin.</figcaption></figure> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Lyristes_plebejus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/71/Lyristes_plebejus.jpg/220px-Lyristes_plebejus.jpg" decoding="async" width="220" height="210" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/71/Lyristes_plebejus.jpg/330px-Lyristes_plebejus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/71/Lyristes_plebejus.jpg/440px-Lyristes_plebejus.jpg 2x" data-file-width="1842" data-file-height="1758" /></a><figcaption>A <a href="/wiki/Cicada" title="Cicada">cicada</a> emerges from its nymphal exoskeleton; the shed exoskeleton is mostly modified chitin (<a href="/wiki/Sclerotin" title="Sclerotin">sclerotin</a>) but the wings and much of the adult body are still unsclerotized chitin at this stage</figcaption></figure> <p><b>Chitin</b> (<a href="/wiki/Carbon" title="Carbon">C</a><sub>8</sub><a href="/wiki/Hydrogen" title="Hydrogen">H</a><sub>13</sub><a href="/wiki/Oxygen" title="Oxygen">O</a><sub>5</sub><a href="/wiki/Nitrogen" title="Nitrogen">N</a>)<sub>n</sub> (<span class="rt-commentedText nowrap"><span class="IPA nopopups noexcerpt" lang="en-fonipa"><a href="/wiki/Help:IPA/English" title="Help:IPA/English">/<span style="border-bottom:1px dotted"><span title="/ˈ/: primary stress follows">ˈ</span><span title="'k' in 'kind'">k</span><span title="/aɪ/: 'i' in 'tide'">aɪ</span><span title="'t' in 'tie'">t</span><span title="/ɪ/: 'i' in 'kit'">ɪ</span><span title="'n' in 'nigh'">n</span></span>/</a></span></span> <a href="/wiki/Help:Pronunciation_respelling_key" title="Help:Pronunciation respelling key"><i title="English pronunciation respelling"><span style="font-size:90%">KY</span>-tin</i></a>) is a long-chain <a href="/wiki/Polymer" title="Polymer">polymer</a> of <a href="/wiki/N-Acetylglucosamine" title="N-Acetylglucosamine"><i>N</i>-acetylglucosamine</a>, an <a href="/wiki/Amide" title="Amide">amide</a> derivative of <a href="/wiki/Glucose" title="Glucose">glucose</a>. Chitin is the second most abundant <a href="/wiki/Polysaccharide" title="Polysaccharide">polysaccharide</a> in nature (behind only <a href="/wiki/Cellulose" title="Cellulose">cellulose</a>); an estimated 1 billion tons of chitin are produced each year in the <a href="/wiki/Biosphere" title="Biosphere">biosphere</a>.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> It is a primary component of <a href="/wiki/Cell_wall" title="Cell wall">cell walls</a> in <a href="/wiki/Fungi" class="mw-redirect" title="Fungi">fungi</a> (especially filamentous and mushroom-forming fungi), the <a href="/wiki/Exoskeleton" title="Exoskeleton">exoskeletons</a> of <a href="/wiki/Arthropod" title="Arthropod">arthropods</a> such as crustaceans and insects, the <a href="/wiki/Radula" title="Radula">radulae</a>, <a href="/wiki/Cephalopod_beak" title="Cephalopod beak">cephalopod beaks</a> and <a href="/wiki/Gladius_(cephalopod)" title="Gladius (cephalopod)">gladii</a> of <a href="/wiki/Mollusc" class="mw-redirect" title="Mollusc">molluscs</a> and in some nematodes and diatoms.<sup id="cite_ref-:0_2-0" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:1_3-0" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> It is also synthesised by at least some fish and <a href="/wiki/Lissamphibia" title="Lissamphibia">lissamphibians</a>.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Commercially, chitin is extracted from the shells of crabs, shrimps, shellfish and lobsters, which are major by-products of the seafood industry.<sup id="cite_ref-:0_2-1" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:1_3-1" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> The structure of chitin is comparable to cellulose, forming crystalline nanofibrils or whiskers. It is functionally comparable to the protein <a href="/wiki/Keratin" title="Keratin">keratin</a>. Chitin has proved useful for several medicinal, industrial and biotechnological purposes.<sup id="cite_ref-:1_3-2" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Etymology">Etymology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=1" title="Edit section: Etymology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The English word "chitin" comes from the <a href="/wiki/French_language" title="French language">French</a> word <i>chitine</i>, which was derived in 1821 from the <a href="/wiki/Greek_language" title="Greek language">Greek</a> word χιτών (<i>khitōn</i>) meaning covering.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>A similar word, "<a href="/wiki/Chiton#Name" title="Chiton">chiton</a>", refers to a marine animal with a protective shell. </p> <div class="mw-heading mw-heading2"><h2 id="Chemistry,_physical_properties_and_biological_function"><span id="Chemistry.2C_physical_properties_and_biological_function"></span>Chemistry, physical properties and biological function</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=2" title="Edit section: Chemistry, physical properties and biological function"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Chitin_glucose_and_cellulose.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/57/Chitin_glucose_and_cellulose.svg/250px-Chitin_glucose_and_cellulose.svg.png" decoding="async" width="200" height="107" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/57/Chitin_glucose_and_cellulose.svg/330px-Chitin_glucose_and_cellulose.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/57/Chitin_glucose_and_cellulose.svg/500px-Chitin_glucose_and_cellulose.svg.png 2x" data-file-width="512" data-file-height="274" /></a><figcaption>Chemical configurations of the different monosaccharides (glucose and N-acetylglucosamine) and polysaccharides (chitin and cellulose) presented in <a href="/wiki/Haworth_projection" title="Haworth projection">Haworth projection</a></figcaption></figure> <p>The structure of chitin was determined by <a href="/wiki/Albert_Hofmann" title="Albert Hofmann">Albert Hofmann</a> in 1929. Hofmann hydrolyzed chitin using a crude preparation of the enzyme chitinase, which he obtained from the snail <i>Helix pomatia</i>.<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p><p>Chitin is a modified <a href="/wiki/Polysaccharide" title="Polysaccharide">polysaccharide</a> that contains nitrogen; it is <a href="/wiki/Biosynthesis" title="Biosynthesis">synthesized</a> from units of <a href="/wiki/N-acetylglucosamine" class="mw-redirect" title="N-acetylglucosamine"><i>N</i>-acetyl-<small>D</small>-glucosamine</a> (to be precise, 2-(acetylamino)-2-deoxy-<small>D</small>-glucose). These units form covalent β-(1→4)-linkages (like the linkages between <a href="/wiki/Glucose" title="Glucose">glucose</a> units forming <a href="/wiki/Cellulose" title="Cellulose">cellulose</a>). Therefore, chitin may be described as <a href="/wiki/Cellulose" title="Cellulose">cellulose</a> with one <a href="/wiki/Hydroxyl" class="mw-redirect" title="Hydroxyl">hydroxyl</a> group on each <a href="/wiki/Monomer" title="Monomer">monomer</a> replaced with an <a href="/wiki/Acetyl" class="mw-redirect" title="Acetyl">acetyl</a> <a href="/wiki/Amine" title="Amine">amine</a> group. This allows for increased <a href="/wiki/Hydrogen_bonding" class="mw-redirect" title="Hydrogen bonding">hydrogen bonding</a> between adjacent <a href="/wiki/Polymers" class="mw-redirect" title="Polymers">polymers</a>, giving the chitin-polymer matrix increased strength. </p><p>In its pure, unmodified form, chitin is translucent, pliable, resilient, and quite tough. In most <a href="/wiki/Arthropod" title="Arthropod">arthropods</a>, however, it is often modified, occurring largely as a component of <a href="/wiki/Composite_material" title="Composite material">composite materials</a>, such as in <a href="/wiki/Sclerotin" title="Sclerotin">sclerotin</a>, a tanned <a href="/wiki/Protein" title="Protein">proteinaceous</a> matrix, which forms much of the <a href="/wiki/Exoskeleton" title="Exoskeleton">exoskeleton</a> of <a href="/wiki/Insect" title="Insect">insects</a>. Combined with <a href="/wiki/Calcium_carbonate" title="Calcium carbonate">calcium carbonate</a>, as in the shells of <a href="/wiki/Crustacean" title="Crustacean">crustaceans</a> and <a href="/wiki/Mollusc" class="mw-redirect" title="Mollusc">molluscs</a>, chitin produces a much stronger composite. This composite material is much harder and stiffer than pure chitin, and is tougher and less brittle than pure <a href="/wiki/Calcium_carbonate" title="Calcium carbonate">calcium carbonate</a>.<sup id="cite_ref-Campbell_10-0" class="reference"><a href="#cite_note-Campbell-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> Another difference between pure and composite forms can be seen by comparing the flexible body wall of a <a href="/wiki/Caterpillar" title="Caterpillar">caterpillar</a> (mainly chitin) to the stiff, light <a href="/wiki/Elytron" title="Elytron">elytron</a> of a <a href="/wiki/Beetle" title="Beetle">beetle</a> (containing a large proportion of <a href="/wiki/Sclerotin" title="Sclerotin">sclerotin</a>).<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p><p>In butterfly wing scales, chitin is organized into stacks of <a href="/wiki/Gyroid" title="Gyroid">gyroids</a> constructed of chitin <a href="/wiki/Photonic_crystal" title="Photonic crystal">photonic crystals</a> that produce various <a href="/wiki/Iridescent" class="mw-redirect" title="Iridescent">iridescent</a> colors serving <a href="/wiki/Phenotype" title="Phenotype">phenotypic</a> signaling and communication for mating and foraging.<sup id="cite_ref-wings_12-0" class="reference"><a href="#cite_note-wings-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> The elaborate chitin gyroid construction in butterfly wings creates a model of optical devices having potential for innovations in <a href="/wiki/Biomimicry" class="mw-redirect" title="Biomimicry">biomimicry</a>.<sup id="cite_ref-wings_12-1" class="reference"><a href="#cite_note-wings-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Scarab_beetle" class="mw-redirect" title="Scarab beetle">Scarab beetles</a> in the genus <i><a href="/wiki/Cyphochilus" title="Cyphochilus">Cyphochilus</a></i> also utilize chitin to form extremely thin <a href="/wiki/Scale_(anatomy)" class="mw-redirect" title="Scale (anatomy)">scales</a> (five to fifteen <a href="/wiki/Micrometre" title="Micrometre">micrometres</a> thick) that diffusely reflect white light. These scales are networks of randomly ordered filaments of chitin with diameters on the scale of hundreds of <a href="/wiki/Nanometre" title="Nanometre">nanometres</a>, which serve to scatter light. The <a href="/wiki/Scattering#Single_and_multiple_scattering" title="Scattering">multiple scattering</a> of light is thought to play a role in the unusual whiteness of the scales.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Burresi_14-0" class="reference"><a href="#cite_note-Burresi-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> In addition, some social wasps, such as <i><a href="/wiki/Protopolybia_chartergoides" title="Protopolybia chartergoides">Protopolybia chartergoides</a></i>, orally secrete material containing predominantly chitin to reinforce the outer nest envelopes, composed of paper.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p><p><a href="/wiki/Chitosan" title="Chitosan">Chitosan</a> is produced commercially by <a href="/wiki/Deacetylation" class="mw-redirect" title="Deacetylation">deacetylation</a> of chitin by treatment with <a href="/wiki/Sodium_hydroxide" title="Sodium hydroxide">sodium hydroxide</a>. Chitosan has a wide range of biomedical applications including wound healing, drug delivery and tissue engineering.<sup id="cite_ref-:0_2-2" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:1_3-3" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Due to its specific intermolecular hydrogen bonding network, dissolving chitin in water is very difficult.<sup id="cite_ref-Bedian2017rev_16-0" class="reference"><a href="#cite_note-Bedian2017rev-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> Chitosan (with a degree of deacetylation of more than ~28%), on the other hand, can be dissolved in dilute acidic aqueous solutions below a pH of 6.0 such as acetic, formic and lactic acids. Chitosan with a degree of deacetylation greater than ~49% is soluble in water<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Humans_and_other_mammals">Humans and other mammals</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=3" title="Edit section: Humans and other mammals"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Humans and other mammals have <a href="/wiki/Chitinase" title="Chitinase">chitinase</a> and <a href="/wiki/CHI3L1" title="CHI3L1">chitinase-like proteins</a> that can degrade chitin; they also possess several <a href="/wiki/Immune_receptor" title="Immune receptor">immune receptors</a> that can recognize chitin and its degradation products, initiating an <a href="/wiki/Immune_response" title="Immune response">immune response</a>.<sup id="cite_ref-Komi2017rev_19-0" class="reference"><a href="#cite_note-Komi2017rev-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> </p><p>Chitin is sensed mostly in the lungs or <a href="/wiki/Gastrointestinal_tract" title="Gastrointestinal tract">gastrointestinal tract</a> where it can activate the <a href="/wiki/Innate_immune_system" title="Innate immune system">innate immune system</a> through <a href="/wiki/Eosinophil" title="Eosinophil">eosinophils</a> or <a href="/wiki/Macrophage" title="Macrophage">macrophages</a>, as well as an <a href="/wiki/Adaptive_immune_response" class="mw-redirect" title="Adaptive immune response">adaptive immune response</a> through <a href="/wiki/T_helper" class="mw-redirect" title="T helper">T helper</a> cells.<sup id="cite_ref-Komi2017rev_19-1" class="reference"><a href="#cite_note-Komi2017rev-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Keratinocyte" title="Keratinocyte">Keratinocytes</a> in skin can also react to chitin or chitin fragments.<sup id="cite_ref-Komi2017rev_19-2" class="reference"><a href="#cite_note-Komi2017rev-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Plants">Plants</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=4" title="Edit section: Plants"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Plants also have receptors that can cause a response to chitin, namely chitin elicitor receptor kinase 1 and chitin elicitor-binding protein.<sup id="cite_ref-Komi2017rev_19-3" class="reference"><a href="#cite_note-Komi2017rev-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> The first chitin receptor was cloned in 2006.<sup id="cite_ref-Sanchez2015rev_20-0" class="reference"><a href="#cite_note-Sanchez2015rev-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> When the receptors are activated by chitin, genes related to plant defense are expressed, and <a href="/wiki/Jasmonate" title="Jasmonate">jasmonate</a> hormones are activated, which in turn activate systemic defenses.<sup id="cite_ref-Sharp2013rev_21-0" class="reference"><a href="#cite_note-Sharp2013rev-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Commensalism" title="Commensalism">Commensal</a> fungi have ways to interact with the host immune response that, as of 2016<sup class="plainlinks noexcerpt noprint asof-tag update" style="display:none;"><a class="external text" href="https://en.wikipedia.org/w/index.php?title=Chitin&action=edit">[update]</a></sup>, were not well understood.<sup id="cite_ref-Sanchez2015rev_20-1" class="reference"><a href="#cite_note-Sanchez2015rev-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> </p><p>Some pathogens produce chitin-binding proteins that mask the chitin they shed from these receptors.<sup id="cite_ref-Sharp2013rev_21-1" class="reference"><a href="#cite_note-Sharp2013rev-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> <i><a href="/wiki/Zymoseptoria_tritici" title="Zymoseptoria tritici">Zymoseptoria tritici</a></i> is an example of a fungal pathogen that has such blocking proteins; it is a major pest in <a href="/wiki/Wheat" title="Wheat">wheat</a> crops.<sup id="cite_ref-Kettles2016rev_23-0" class="reference"><a href="#cite_note-Kettles2016rev-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Fossil_record">Fossil record</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=5" title="Edit section: Fossil record"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">For more on the preservation potential of chitin and other biopolymers, see <a href="/wiki/Taphonomy" title="Taphonomy">taphonomy</a>.</div> <p>Chitin was probably present in the exoskeletons of <a href="/wiki/Cambrian" title="Cambrian">Cambrian</a> arthropods such as <a href="/wiki/Trilobite" title="Trilobite">trilobites</a>. The oldest preserved (intact) chitin samples thus far reported are dated to the <a href="/wiki/Oligocene" title="Oligocene">Oligocene</a>, about <span class="nourlexpansion plainlinks" style="white-space:nowrap;"><a rel="nofollow" class="external text" href="https://geoltime.github.io/?Ma=25">25</a> million years ago</span>, from specimens encased in <a href="/wiki/Amber" title="Amber">amber</a> where the chitin has not completely degraded.<sup id="cite_ref-Briggs1999_24-0" class="reference"><a href="#cite_note-Briggs1999-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Uses">Uses</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=6" title="Edit section: Uses"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Agriculture">Agriculture</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=7" title="Edit section: Agriculture"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Chitin is a good inducer of plant <a href="/wiki/Plant_defense_against_herbivory" title="Plant defense against herbivory">defense mechanisms</a> for controlling <a href="/wiki/Plant_disease" title="Plant disease">diseases</a>.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> It has potential for use as a soil <a href="/wiki/Fertilizer" title="Fertilizer">fertilizer</a> or <a href="/wiki/Soil_conditioner" title="Soil conditioner">conditioner</a> to improve fertility and plant resilience that may enhance crop yields.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Industrial">Industrial</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=8" title="Edit section: Industrial"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Chitin is used in many industrial processes. Examples of the potential uses of chemically modified chitin in <a href="/wiki/Food_processing" title="Food processing">food processing</a> include the formation of edible films and as an additive to thicken and stabilize foods and food emulsions.<sup id="cite_ref-28" class="reference"><a href="#cite_note-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Shahidi_29-0" class="reference"><a href="#cite_note-Shahidi-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> Processes to <a href="/wiki/Sizing" title="Sizing">size</a> and strengthen <a href="/wiki/Paper" title="Paper">paper</a> employ chitin and chitosan.<sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-31" class="reference"><a href="#cite_note-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Research">Research</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=9" title="Edit section: Research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>How chitin interacts with the <a href="/wiki/Immune_system" title="Immune system">immune system</a> of plants and animals has been an active area of research, including the identity of key <a href="/wiki/Receptor_(biochemistry)" title="Receptor (biochemistry)">receptors</a> with which chitin interacts, whether the size of chitin particles is relevant to the kind of immune response triggered, and mechanisms by which immune systems respond.<sup id="cite_ref-Gomez-Casado2016rev_32-0" class="reference"><a href="#cite_note-Gomez-Casado2016rev-32"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Kettles2016rev_23-1" class="reference"><a href="#cite_note-Kettles2016rev-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> Chitin is deacetylated chemically or enzymatically to produce <a href="/wiki/Chitosan" title="Chitosan">chitosan</a>, a highly <a href="/wiki/Biocompatibility" title="Biocompatibility">biocompatible</a> polymer which has found a wide range of applications in the biomedical industry.<sup id="cite_ref-:0_2-3" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-33" class="reference"><a href="#cite_note-33"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-34" class="reference"><a href="#cite_note-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup> Chitin and chitosan have been explored as a <a href="/wiki/Vaccine_adjuvant" class="mw-redirect" title="Vaccine adjuvant">vaccine adjuvant</a> due to its ability to stimulate an immune response.<sup id="cite_ref-:0_2-4" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Komi2017rev_19-4" class="reference"><a href="#cite_note-Komi2017rev-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> </p><p>Chitin and chitosan are under development as <a href="/wiki/Tissue_engineering#Scaffolds" title="Tissue engineering">scaffolds</a> in studies of how tissue grows and how <a href="/wiki/Wound_healing" title="Wound healing">wounds heal</a>, and in efforts to invent better <a href="/wiki/Bandages" class="mw-redirect" title="Bandages">bandages</a>, <a href="/wiki/Surgical_suture" title="Surgical suture">surgical thread</a>, and materials for <a href="/wiki/Allotransplantation" title="Allotransplantation">allotransplantation</a>.<sup id="cite_ref-:0_2-5" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Bedian2017rev_16-1" class="reference"><a href="#cite_note-Bedian2017rev-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-35" class="reference"><a href="#cite_note-35"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Surgical_suture" title="Surgical suture">Sutures</a> made of chitin have been experimentally developed, but their lack of elasticity and problems making thread have prevented commercial success so far.<sup id="cite_ref-36" class="reference"><a href="#cite_note-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </p><p><a href="/wiki/Chitosan" title="Chitosan">Chitosan</a> has been demonstrated and proposed to make a reproducible form of <a href="/wiki/Biodegradable" class="mw-redirect" title="Biodegradable">biodegradable</a> plastic.<sup id="cite_ref-37" class="reference"><a href="#cite_note-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup> Chitin <a href="/wiki/Nanofiber" title="Nanofiber">nanofibers</a> are extracted from crustacean waste and mushrooms for possible development of products in <a href="/wiki/Tissue_engineering" title="Tissue engineering">tissue engineering</a>, drug delivery and medicine.<sup id="cite_ref-:0_2-6" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-38" class="reference"><a href="#cite_note-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup> </p><p>Chitin has been proposed for use in building structures, tools, and other solid objects from a <a href="/wiki/Composite_material" title="Composite material">composite material</a>, combining chitin with <a href="/wiki/Martian_soil" class="mw-redirect" title="Martian soil">Martian regolith</a>.<sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">[</span>39<span class="cite-bracket">]</span></a></sup> To build this, the <a href="/wiki/Biopolymers" class="mw-redirect" title="Biopolymers">biopolymers</a> in the chitin are suggested as the <a href="/wiki/Binder_(material)" title="Binder (material)">binder</a> for the regolith <a href="/wiki/Aggregate_(composite)" title="Aggregate (composite)">aggregate</a> to form a <a href="/wiki/Concrete" title="Concrete">concrete</a>-like <a href="/wiki/Composite_material" title="Composite material">composite material</a>. The authors believe that waste materials from food production (e.g. scales from fish, exoskeletons from crustaceans and insects, etc.) could be put to use as feedstock for manufacturing processes. </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=10" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Chitobiose" title="Chitobiose">Chitobiose</a></li> <li><a href="/wiki/Lorica_(biology)" title="Lorica (biology)">Lorica</a></li> <li><a href="/wiki/Sporopollenin" title="Sporopollenin">Sporopollenin</a></li> <li><a href="/wiki/Tectin_(secretion)" title="Tectin (secretion)">Tectin</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=11" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width reflist-columns-2"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFNelson,_D.L.,_Cox,_M.M.2017" class="citation book cs1">Nelson, D.L., Cox, M.M. 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href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7494075">7494075</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/32936806">32936806</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=PLOS+ONE&rft.atitle=Martian+biolith%3A+A+bioinspired+regolith+composite+for+closed-loop+extraterrestrial+manufacturing&rft.volume=15&rft.issue=9&rft.pages=e0238606&rft.date=2020-09-16&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7494075%23id-name%3DPMC&rft_id=info%3Apmid%2F32936806&rft_id=info%3Adoi%2F10.1371%2Fjournal.pone.0238606&rft_id=info%3Abibcode%2F2020PLoSO..1538606S&rft.aulast=Shiwei&rft.aufirst=Ng&rft.au=Dritsas%2C+Stylianos&rft.au=Fernandez%2C+Javier+G.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7494075&rfr_id=info%3Asid%2Fen.wikipedia.org%3AChitin" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Chitin&action=edit&section=12" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output 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href="https://commons.wikimedia.org/wiki/Category:Chitin" class="extiw" title="commons:Category:Chitin">Chitin</a> at Wikimedia Commons</li> <li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Wiktionary-logo-en-v2.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/16px-Wiktionary-logo-en-v2.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/24px-Wiktionary-logo-en-v2.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/32px-Wiktionary-logo-en-v2.svg.png 2x" data-file-width="512" data-file-height="512" /></a></span> The dictionary definition of <a href="https://en.wiktionary.org/wiki/Special:Search/chitin" class="extiw" title="wiktionary:Special:Search/chitin"><i>chitin</i></a> at Wiktionary</li></ul> <div class="navbox-styles"><style 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abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Carbohydrates" title="Template:Carbohydrates"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Carbohydrates" title="Template talk:Carbohydrates"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Carbohydrates" title="Special:EditPage/Template:Carbohydrates"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Types_of_carbohydrates184" style="font-size:114%;margin:0 4em">Types of <a href="/wiki/Carbohydrate" title="Carbohydrate">carbohydrates</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aldose" title="Aldose">Aldose</a></li> <li><a href="/wiki/Ketose" title="Ketose">Ketose</a></li> <li><a href="/wiki/Furanose" title="Furanose">Furanose</a></li> <li><a href="/wiki/Pyranose" title="Pyranose">Pyranose</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Geometry</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anomer" title="Anomer">Anomer</a></li> <li><a href="/wiki/Cyclohexane_conformation" title="Cyclohexane conformation">Cyclohexane conformation</a></li> <li><a href="/wiki/Epimer" title="Epimer">Epimer</a></li> <li><a href="/wiki/Mutarotation" title="Mutarotation">Mutarotation</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Monosaccharide" title="Monosaccharide">Monosaccharides</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Diose" title="Diose">Dioses</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Aldodiose <ul><li><a href="/wiki/Glycolaldehyde" title="Glycolaldehyde">Glycolaldehyde</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Triose" title="Triose">Trioses</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Aldotriose <ul><li><a href="/wiki/Glyceraldehyde" title="Glyceraldehyde">Glyceraldehyde</a></li></ul></li> <li>Ketotriose <ul><li><a href="/wiki/Dihydroxyacetone" title="Dihydroxyacetone">Dihydroxyacetone</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tetrose" title="Tetrose">Tetroses</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Aldotetroses <ul><li><a href="/wiki/Erythrose" title="Erythrose">Erythrose</a></li> <li><a href="/wiki/Threose" title="Threose">Threose</a></li></ul></li> <li>Ketotetrose <ul><li><a href="/wiki/Erythrulose" title="Erythrulose">Erythrulose</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pentose" title="Pentose">Pentoses</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Aldopentoses <ul><li><a href="/wiki/Arabinose" title="Arabinose">Arabinose</a></li> <li><a href="/wiki/Lyxose" title="Lyxose">Lyxose</a></li> <li><a href="/wiki/Ribose" title="Ribose">Ribose</a></li> <li><a href="/wiki/Xylose" title="Xylose">Xylose</a></li></ul></li> <li>Ketopentoses <ul><li><a href="/wiki/Ribulose" title="Ribulose">Ribulose</a></li> <li><a href="/wiki/Xylulose" title="Xylulose">Xylulose</a></li></ul></li> <li><a href="/wiki/Deoxy_sugar" title="Deoxy sugar">Deoxy sugars</a> <ul><li><a href="/wiki/Deoxyribose" title="Deoxyribose">Deoxyribose</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Hexose" title="Hexose">Hexoses</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aldohexose" class="mw-redirect" title="Aldohexose">Aldohexoses</a> <ul><li><a href="/wiki/Allose" title="Allose">Allose</a></li> <li><a href="/wiki/Altrose" title="Altrose">Altrose</a></li> <li><a href="/wiki/Galactose" title="Galactose">Galactose</a></li> <li><a href="/wiki/Glucose" title="Glucose">Glucose</a></li> <li><a href="/wiki/Gulose" title="Gulose">Gulose</a></li> <li><a href="/wiki/Idose" title="Idose">Idose</a></li> <li><a href="/wiki/Mannose" title="Mannose">Mannose</a></li> <li><a href="/wiki/Talose" title="Talose">Talose</a></li></ul></li> <li><a href="/wiki/Ketohexose" class="mw-redirect" title="Ketohexose">Ketohexoses</a> <ul><li><a href="/wiki/Fructose" title="Fructose">Fructose</a></li> <li><a href="/wiki/Psicose" title="Psicose">Psicose</a></li> <li><a href="/wiki/Sorbose" title="Sorbose">Sorbose</a></li> <li><a href="/wiki/Tagatose" title="Tagatose">Tagatose</a></li></ul></li> <li>Deoxy sugars <ul><li><a href="/wiki/Fucose" title="Fucose">Fucose</a></li> <li><a href="/wiki/Fuculose" title="Fuculose">Fuculose</a></li> <li><a href="/wiki/Rhamnose" title="Rhamnose">Rhamnose</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Heptose" title="Heptose">Heptoses</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Ketoheptoses <ul><li><a href="/wiki/Mannoheptulose" title="Mannoheptulose">Mannoheptulose</a></li> <li><a href="/wiki/Sedoheptulose" title="Sedoheptulose">Sedoheptulose</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Above 7</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Octose" title="Octose">Octoses</a></li> <li><a href="/wiki/Nonose" title="Nonose">Nonoses</a> <ul><li><a href="/wiki/Neuraminic_acid" title="Neuraminic acid">Neuraminic acid</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Multiple</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Disaccharide" title="Disaccharide">Disaccharides</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cellobiose" title="Cellobiose">Cellobiose</a></li> <li><a href="/wiki/Isomaltose" title="Isomaltose">Isomaltose</a></li> <li><a href="/wiki/Isomaltulose" title="Isomaltulose">Isomaltulose</a></li> <li><a href="/wiki/Lactose" title="Lactose">Lactose</a></li> <li><a href="/wiki/Lactulose" title="Lactulose">Lactulose</a></li> <li><a href="/wiki/Maltose" title="Maltose">Maltose</a></li> <li><a href="/wiki/Sucrose" title="Sucrose">Sucrose</a></li> <li><a href="/wiki/Trehalose" title="Trehalose">Trehalose</a></li> <li><a href="/wiki/Turanose" title="Turanose">Turanose</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Trisaccharide" title="Trisaccharide">Trisaccharides</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Maltotriose" title="Maltotriose">Maltotriose</a></li> <li><a href="/wiki/Melezitose" title="Melezitose">Melezitose</a></li> <li><a href="/wiki/Raffinose" title="Raffinose">Raffinose</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tetrasaccharide" title="Tetrasaccharide">Tetrasaccharides</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Stachyose" title="Stachyose">Stachyose</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other<br /> <a href="/wiki/Oligosaccharide" title="Oligosaccharide">oligosaccharides</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acarbose" title="Acarbose">Acarbose</a></li> <li><a href="/wiki/Fructooligosaccharide" title="Fructooligosaccharide">Fructooligosaccharide</a> (FOS)</li> <li><a href="/wiki/Galactooligosaccharide" title="Galactooligosaccharide">Galactooligosaccharide</a> (GOS)</li> <li><a href="/wiki/Isomaltooligosaccharide" title="Isomaltooligosaccharide">Isomaltooligosaccharide</a> (IMO)</li> <li><a href="/wiki/Maltodextrin" title="Maltodextrin">Maltodextrin</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Polysaccharide" title="Polysaccharide">Polysaccharides</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Beta-glucan" title="Beta-glucan">Beta-glucan</a> <ul><li><a href="/wiki/Oat_beta-glucan" title="Oat beta-glucan">Oat beta-glucan</a></li> <li><a href="/wiki/Lentinan" title="Lentinan">Lentinan</a></li> <li><a href="/wiki/Schizophyllan" title="Schizophyllan">Sizofiran</a></li> <li><a href="/wiki/Zymosan" title="Zymosan">Zymosan</a></li> <li><a href="/wiki/Cellulose" title="Cellulose">Cellulose</a></li> <li><a class="mw-selflink selflink">Chitin</a></li></ul></li> <li><a href="/wiki/Chitosan" title="Chitosan">Chitosan</a></li> <li><a href="/wiki/Dextrin" title="Dextrin">Dextrin</a> / <a href="/wiki/Dextran" title="Dextran">Dextran</a></li> <li><i><a href="/wiki/Fructose" title="Fructose">Fructose</a> / <a href="/wiki/Fructan" title="Fructan">Fructan</a></i> <ul><li><a href="/wiki/Inulin" title="Inulin">Inulin</a></li></ul></li> <li><i><a href="/wiki/Galactose" title="Galactose">Galactose</a> / <a href="/wiki/Galactose" title="Galactose">Galactan</a></i></li> <li><i><a href="/wiki/Glucose" title="Glucose">Glucose</a> / <a href="/wiki/Glucan" title="Glucan">Glucan</a></i> <ul><li><a href="/wiki/Glycogen" title="Glycogen">Glycogen</a></li></ul></li> <li><a href="/wiki/Hemicellulose" title="Hemicellulose">Hemicellulose</a></li> <li><a href="/wiki/Levan_polysaccharide" title="Levan polysaccharide">Levan beta 2→6</a></li> <li><a href="/wiki/Lignin" title="Lignin">Lignin</a></li> <li><a href="/wiki/Mannans" title="Mannans">Mannan</a></li> <li><a href="/wiki/Pectin" title="Pectin">Pectin</a></li> <li><a href="/wiki/Starch" title="Starch">Starch</a> <ul><li><a href="/wiki/Amylopectin" title="Amylopectin">Amylopectin</a></li> <li><a href="/wiki/Amylose" title="Amylose">Amylose</a></li></ul></li> <li><a href="/wiki/Xanthan_gum" title="Xanthan gum">Xanthan gum</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Carbohydrates" title="Category:Carbohydrates">Category</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /></div><div role="navigation" class="navbox" aria-labelledby="Human_interactions_with_insects236" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231" /><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Insects_in_culture" title="Template:Insects in culture"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Insects_in_culture" title="Template talk:Insects in culture"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Insects_in_culture" title="Special:EditPage/Template:Insects in culture"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Human_interactions_with_insects236" style="font-size:114%;margin:0 4em"><a href="/wiki/Human_interactions_with_insects" title="Human interactions with insects">Human interactions with insects</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Aspects<br />of <a href="/wiki/Insect" title="Insect">insects</a><br />in culture</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">In the arts</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Insects_in_art" title="Insects in art">Insects in art</a> <ul><li><a href="/wiki/Beetlewing" title="Beetlewing">Beetlewing</a></li> <li><i><a href="/wiki/Musca_depicta" title="Musca depicta">Musca depicta</a></i></li></ul></li> <li><a href="/wiki/Arthropods_in_film" title="Arthropods in film">Arthropods in film</a></li> <li><a href="/wiki/Insects_in_literature" title="Insects in literature">Insects in literature</a></li> <li><a href="/wiki/Insects_in_music" title="Insects in music">Insects in music</a> <ul><li><a href="/wiki/List_of_insect-inspired_songs" title="List of insect-inspired songs">List of insect-inspired songs</a></li></ul></li> <li><a href="/wiki/Insects_on_stamps" title="Insects on stamps">Insects on stamps</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">In fishing</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Fishing_bait" title="Fishing bait">Fishing bait</a></li> <li><a href="/wiki/Fly_fishing" title="Fly fishing">Fly fishing</a></li> <li><a href="/wiki/Artificial_fly" title="Artificial fly">Artificial fly</a></li> <li><a href="/wiki/Fly_tying" title="Fly tying">Fly tying</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Insects_in_medicine" title="Insects in medicine">In medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Apitherapy" title="Apitherapy">Apitherapy</a> <ul><li><a href="/wiki/Apitoxin" title="Apitoxin">Apitoxin</a></li> <li><a href="/wiki/Melittin" title="Melittin">Melittin</a></li></ul></li> <li><a href="/wiki/Lytta_vesicatoria" title="Lytta vesicatoria">Spanish fly</a> <ul><li><a href="/wiki/Cantharidin" title="Cantharidin">Cantharidin</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Insects_in_mythology" title="Insects in mythology">In mythology</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bees_in_mythology" title="Bees in mythology">Bees in mythology</a></li> <li><a href="/wiki/Cicadas_in_mythology" title="Cicadas in mythology">Cicadas in mythology</a></li> <li><a href="/wiki/Scarab_(artifact)" title="Scarab (artifact)">Scarab (artifact)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other aspects</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biomimetics" title="Biomimetics">Biomimicry</a></li> <li><a href="/wiki/Cockroach_racing" title="Cockroach racing">Cockroach racing</a></li> <li><a href="/wiki/Cockroaches_in_popular_culture" title="Cockroaches in popular culture">Cockroaches in popular culture</a></li> <li><a href="/wiki/Cricket_fighting" title="Cricket fighting">Cricket fighting</a></li> <li><a href="/wiki/Entomological_warfare" title="Entomological warfare">Entomological warfare</a></li> <li><a href="/wiki/Entomophagy_in_humans" title="Entomophagy in humans">Entomophagy</a></li> <li><a href="/wiki/Insect_farming" title="Insect farming">Insect farming</a></li> <li><a href="/wiki/Flea_circus" title="Flea circus">Flea circus</a></li> <li><a href="/wiki/Insects_in_ethics" title="Insects in ethics">Insects in ethics</a></li> <li><a href="/wiki/Insects_in_Japanese_culture" title="Insects in Japanese culture">Insects in Japan</a></li> <li><a href="/wiki/Insects_in_religion" title="Insects in religion">Insects in religion</a></li> <li><a href="/wiki/Jingzhe" title="Jingzhe">Jingzhe</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Economic_entomology" title="Economic entomology">Economic<br />entomology</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Beneficial_insect" title="Beneficial insect">Beneficial insect</a></li> <li><a href="/wiki/Biological_pest_control" title="Biological pest control">Biological pest control</a></li> <li><a href="/wiki/Beekeeping" title="Beekeeping">Beekeeping</a> <ul><li><a href="/wiki/Bee_pollen" title="Bee pollen">Bee pollen</a></li> <li><a href="/wiki/List_of_crop_plants_pollinated_by_bees" title="List of crop plants pollinated by bees">List of crop plants pollinated by bees</a></li> <li><a href="/wiki/Beeswax" title="Beeswax">Beeswax</a></li> <li><a href="/wiki/Honey" title="Honey">Honey</a></li> <li><a href="/wiki/Propolis" title="Propolis">Propolis</a></li> <li><a href="/wiki/Royal_jelly" title="Royal jelly">Royal jelly</a></li></ul></li> <li><a href="/wiki/Carmine" title="Carmine">Carmine/Cochineal</a> <ul><li><a href="/wiki/Polish_cochineal" title="Polish cochineal">Polish</a></li></ul></li> <li><a class="mw-selflink selflink">Chitin</a></li> <li><a href="/wiki/Kermes_(dye)" title="Kermes (dye)">Kermes</a></li> <li><a href="/wiki/Sericulture" title="Sericulture">Sericulture</a> <ul><li><a href="/wiki/Silk" title="Silk">Silk</a></li></ul></li> <li><a href="/wiki/Shellac" title="Shellac">Shellac</a></li> <li><a href="/wiki/Model_organism" title="Model organism">Model organism</a> <ul><li><i><a href="/wiki/Drosophila_melanogaster" title="Drosophila melanogaster">Drosophila melanogaster</a></i></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pest_(organism)#Insects" title="Pest (organism)">Harmful<br />insects</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Insect_bites_and_stings" class="mw-redirect" title="Insect bites and stings">Insect bites and stings</a></li> <li><a href="/wiki/Insect_sting_allergy" title="Insect sting allergy">Insect sting allergy</a></li> <li><a href="/wiki/Bed_bug" title="Bed bug">Bed bug</a></li> <li><a href="/wiki/Woodworm" title="Woodworm">Woodworm</a></li> <li><a href="/wiki/Home-stored_product_entomology" title="Home-stored product entomology">Home-stored product entomology</a></li> <li><a href="/wiki/Clothes_moth" title="Clothes moth">Clothes moth</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Pioneers</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Jan_Swammerdam" title="Jan Swammerdam">Jan Swammerdam</a></li> <li><a href="/wiki/Alfred_Russel_Wallace" title="Alfred Russel Wallace">Alfred Russel Wallace</a></li> <li><a href="/wiki/Jean-Henri_Fabre" title="Jean-Henri Fabre">Jean-Henri Fabre</a></li> <li><a href="/wiki/Hans_Zinsser" title="Hans Zinsser">Hans Zinsser</a> (<i><a href="/wiki/Rats,_Lice_and_History" title="Rats, Lice and History">Rats, Lice and History</a></i>)</li> <li><a href="/wiki/Lafcadio_Hearn" title="Lafcadio Hearn">Lafcadio Hearn</a> (<i>Insect Literature</i>)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Concerns</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bees_and_toxic_chemicals" title="Bees and toxic chemicals">Bees and toxic chemicals</a></li> <li><a href="/wiki/Colony_collapse_disorder" title="Colony collapse disorder">Colony collapse disorder</a></li> <li><a href="/wiki/Decline_in_insect_populations" title="Decline in insect populations">Decline in insect populations</a></li> <li><a href="/wiki/Habitat_destruction" title="Habitat destruction">Habitat destruction</a></li> <li><a href="/wiki/List_of_endangered_insects" title="List of endangered insects">List of endangered insects</a></li> <li><a href="/wiki/Pesticide" title="Pesticide">Pesticide</a> <ul><li><a href="/wiki/Insecticide" title="Insecticide">Insecticide</a></li> <li><a href="/wiki/Neonicotinoid" title="Neonicotinoid">Neonicotinoid</a></li> <li><a href="/wiki/Pesticide_toxicity_to_bees" title="Pesticide toxicity to bees">Pesticide toxicity to bees</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Categories,<br />templates</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Category:Insects_and_humans" title="Category:Insects and humans">Insects and humans</a></li> <li><a href="/wiki/Template:Insecticides" title="Template:Insecticides">Insecticides</a></li> <li><a href="/wiki/Template:Pesticides" title="Template:Pesticides">Pesticides</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="font-weight:bold;"><div><span class="nowrap"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Beetle_icon.svg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Beetle_icon.svg/17px-Beetle_icon.svg.png" decoding="async" width="17" height="19" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Beetle_icon.svg/26px-Beetle_icon.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Beetle_icon.svg/34px-Beetle_icon.svg.png 2x" data-file-width="143" data-file-height="159" /></a></span> </span><a href="/wiki/Portal:Insects" title="Portal:Insects">Insects portal</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /><style data-mw-deduplicate="TemplateStyles:r1038841319">.mw-parser-output .tooltip-dotted{border-bottom:1px dotted;cursor:help}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1038841319" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1038841319" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1038841319" /></div><div role="navigation" class="navbox authority-control" aria-label="Navbox1570" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a>: National <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q161219#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/4147737-6">Germany</a></span></li><li><span class="uid"><span class="rt-commentedText tooltip tooltip-dotted" title="Chitin"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/sh85024494">United States</a></span></span></li><li><span class="uid"><span class="rt-commentedText tooltip tooltip-dotted" title="Chitine"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb11977844t">France</a></span></span></li><li><span class="uid"><span class="rt-commentedText tooltip tooltip-dotted" title="Chitine"><a rel="nofollow" class="external 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