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id="siteNotice"></div> </div> <div class="pre-content heading-holder"> <div class="page-heading"> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Climbing gourami</span></h1> <div class="tagline"></div> </div> <ul id="p-associated-pages" class="minerva__tab-container"> <li class="minerva__tab selected"> <a class="minerva__tab-text" href="/wiki/Climbing_gourami" rel="" data-event-name="tabs.subject">Article</a> </li> <li class="minerva__tab "> <a class="minerva__tab-text" href="/wiki/Talk:Climbing_gourami" rel="discussion" data-event-name="tabs.talk">Talk</a> </li> </ul> <nav class="page-actions-menu"> <ul id="p-views" class="page-actions-menu__list"> <li id="language-selector" class="page-actions-menu__list-item"> <a role="button" href="#p-lang" data-mw="interface" data-event-name="menu.languages" title="Language" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only 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class="minerva-icon minerva-icon--edit"></span> <span>Edit</span> </a> </li> </ul> </nav>  <div id="mw-content-subtitle"></div> </div> <div id="bodyContent" class="content"> <div id="mw-content-text" class="mw-body-content"><script>function mfTempOpenSection(id){var block=document.getElementById("mf-section-"+id);block.className+=" open-block";block.previousSibling.className+=" open-block";}</script><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><section class="mf-section-0" id="mf-section-0"> <p class="mw-empty-elt"> </p> <p>The <b>Anabantidae</b> are a <a href="/wiki/Family_(biology)" title="Family (biology)">family</a> of <a href="/wiki/Ray-finned_fish" class="mw-redirect" title="Ray-finned fish">ray-finned fish</a> within the <a href="/wiki/Order_(biology)" title="Order (biology)">order</a> <a href="/wiki/Anabantiformes" title="Anabantiformes">Anabantiformes</a> commonly called the <b>climbing gouramies</b> or <b>climbing perches</b>.<sup id="cite_ref-FBF_2-0" class="reference"><a href="#cite_note-FBF-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The family includes about 34 species. As <a href="/wiki/Anabantoidei" title="Anabantoidei">labyrinth fishes</a>, they possess a <a href="/wiki/Labyrinth_organ" class="mw-redirect" title="Labyrinth organ">labyrinth organ</a>, a structure in the fish's head which allows it to breathe atmospheric <a href="/wiki/Oxygen" title="Oxygen">oxygen</a>. Fish of this family are commonly seen gulping at air at the surface of the water. The air is held in a structure called the suprabranchial chamber, where oxygen diffuses into the bloodstream via the respiratory epithelium covering the labyrinth organ. This therefore allows the fish to move small distances across land. </p><table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Climbing gourami<br><div style="font-size: 85%;">Temporal range: <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">26–0 <a href="/wiki/Megaannum" class="mw-redirect" title="Megaannum">Ma</a></span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:211.2px; width:8.8px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:211.2px; width:8.8px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:212.2px; width:6.8px; background-color:#6c3;"></div> </div> </div></span> Late Oligocene to Recent</div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Anabas_testudineus_Day.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Anabas_testudineus_Day.png/220px-Anabas_testudineus_Day.png" decoding="async" width="220" height="101" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Anabas_testudineus_Day.png/330px-Anabas_testudineus_Day.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/da/Anabas_testudineus_Day.png/440px-Anabas_testudineus_Day.png 2x" data-file-width="1200" data-file-height="550"></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><i><a href="/wiki/Anabas_testudineus" title="Anabas testudineus">Anabas testudineus</a></i> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Anabantidae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20"></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Actinopterygii" title="Actinopterygii">Actinopterygii</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Anabantiformes" title="Anabantiformes">Anabantiformes</a> </td></tr> <tr> <td>Family: </td> <td><a class="mw-selflink selflink">Anabantidae</a><br><small><a href="/wiki/Charles_Lucien_Bonaparte" title="Charles Lucien Bonaparte">Bonaparte</a>, 1831<sup id="cite_ref-VDLEF_1-0" class="reference"><a href="#cite_note-VDLEF-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Genera </th></tr> <tr> <td colspan="2" style="text-align: left"> <p>see text </p> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:AnabasLyd.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/b/b7/AnabasLyd.jpg/220px-AnabasLyd.jpg" decoding="async" width="220" height="204" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/b7/AnabasLyd.jpg/330px-AnabasLyd.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/b7/AnabasLyd.jpg/440px-AnabasLyd.jpg 2x" data-file-width="681" data-file-height="633"></a><figcaption>Climbing perch (<i><a href="/wiki/Anabas_testudineus" title="Anabas testudineus">Anabas testudineus</a></i>) on land</figcaption></figure> </section><div class="mw-heading mw-heading2 section-heading" onclick="mfTempOpenSection(1)"><span class="indicator mf-icon mf-icon-expand mf-icon--small"></span><h2 id="Genera">Genera</h2><span class="mw-editsection"> <a role="button" href="/w/index.php?title=Climbing_gourami&action=edit&section=1" title="Edit section: Genera" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet "> <span class="minerva-icon minerva-icon--edit"></span> <span>edit</span> </a> </span> </div><section class="mf-section-1 collapsible-block" id="mf-section-1"> <p>There are four extant genera within the family Anabantidae:<sup id="cite_ref-FBF_2-1" class="reference"><a href="#cite_note-FBF-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <ul><li><i><a href="/wiki/Anabas" title="Anabas">Anabas</a></i> <small><a href="/wiki/Hippolyte_Cloquet" title="Hippolyte Cloquet">Cloquet</a>, 1816</small></li> <li><i><a href="/wiki/Ctenopoma" title="Ctenopoma">Ctenopoma</a></i> <small><a href="/wiki/Wilhelm_Peters" title="Wilhelm Peters">Peters</a>, 1844</small></li> <li><i><a href="/wiki/Microctenopoma" title="Microctenopoma">Microctenopoma</a></i> <small><a href="/w/index.php?title=Steven_Mark_Norris&action=edit&redlink=1" class="new" title="Steven Mark Norris (page does not exist)">Norris</a>, 1995</small></li> <li><i><a href="/wiki/Sandelia" title="Sandelia">Sandelia</a></i> <small>(<a href="/wiki/Francis_de_Laporte_de_Castelnau" class="mw-redirect" title="Francis de Laporte de Castelnau">Castelnau</a>, 1861)</small></li></ul> <p>There is also at least one extinct genus known:<sup id="cite_ref-Eoanabas_3-0" class="reference"><a href="#cite_note-Eoanabas-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <ul><li>†<i><a href="/wiki/Eoanabas" title="Eoanabas">Eoanabas</a></i> <small>Wu, Chang, Miao <i>et al</i>, 2016</small></li></ul> <p>Of the four genera, <i><a href="/wiki/Anabas" title="Anabas">Anabas</a></i> are the only climbing gouramies found in <a href="/wiki/South_Asia" title="South Asia">South Asia</a>. They are widely distributed and are found in countries including <a href="/wiki/Pakistan" title="Pakistan">Pakistan</a>, <a href="/wiki/India" title="India">India</a>, <a href="/wiki/Bangladesh" title="Bangladesh">Bangladesh</a>, <a href="/wiki/Sri_Lanka" title="Sri Lanka">Sri Lanka</a>, and <a href="/wiki/China" title="China">China</a>. Climbing gouramies have distinct names in numerous South Asian languages. In <a href="/wiki/Tamil_language" title="Tamil language">Tamil</a>, they are பனையேறி கெண்டை (<i>panaieri kendai</i>); in <a href="/wiki/Malayalam" title="Malayalam">Malayalam</a>, <i>chemballi</i>; in <a href="/wiki/Odia_language" title="Odia language">Odia</a>, they are କଉ ମାଛ (<i>kau</i>); in <a href="/wiki/Assamese_language" title="Assamese language">Assamese</a> they are কাৱৈ মাছ (<i>kawoi maas</i>); in <a href="/wiki/Bengali_language" title="Bengali language">Bengali</a>, they are কই মাছ (<i>koi mach</i>). </p><p>The remaining three genera are endemic to Africa: <i>Ctenopoma</i> and <i>Microctenopoma</i> are primarily found in the <a href="/wiki/Congo_Basin" title="Congo Basin">Congo Basin</a>, whereas <i>Sandelia</i> are endemic to the <a href="/wiki/Eastern_Cape" title="Eastern Cape">Eastern</a> and <a href="/wiki/Western_Cape" title="Western Cape">Western Capes</a> of South Africa. Anabantidae are primarily <a href="/wiki/Freshwater_fish" title="Freshwater fish">freshwater fish</a> and only very rarely are found in <a href="/wiki/Brackish" class="mw-redirect" title="Brackish">brackish</a> water. Parental care is varied; <i>Anabas</i> and <i>Ctenopoma</i> do not tend their <a href="/wiki/Egg_(biology)" class="mw-redirect" title="Egg (biology)">eggs</a>, <i>Microctenopoma</i> species produce <a href="/wiki/Bubble_nest" title="Bubble nest">bubble nests</a> like many other <a href="/wiki/Labyrinth_fish" class="mw-redirect" title="Labyrinth fish">labyrinth fish</a>, and <i>Sandelia</i> lays their eggs on the substrate. </p><p>Climbing gouramis are so named due to their ability to "climb" out of water and "walk" short distances. Even though it has not been reliably observed, some authors have mentioned about them having a tree climbing ability.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="What authors? (June 2018)">citation needed</span></a></i>]</sup> Their method of terrestrial locomotion uses the gill plates as supports, and the fish pushes itself using its fins and tail. </p> </section><div class="mw-heading mw-heading2 section-heading" onclick="mfTempOpenSection(2)"><span class="indicator mf-icon mf-icon-expand mf-icon--small"></span><h2 id="References">References</h2><span class="mw-editsection"> <a role="button" href="/w/index.php?title=Climbing_gourami&action=edit&section=2" title="Edit section: References" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet "> <span class="minerva-icon minerva-icon--edit"></span> <span>edit</span> </a> </span> </div><section class="mf-section-2 collapsible-block" id="mf-section-2"> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-VDLEF-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-VDLEF_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFRichard_van_der_LaanWilliam_N._EschmeyerRonald_Fricke2014" class="citation journal cs1">Richard van der Laan; William N. Eschmeyer & Ronald Fricke (2014). <a rel="nofollow" class="external text" href="https://doi.org/10.11646%2Fzootaxa.3882.1.1">"Family-group names of Recent fishes"</a>. <i>Zootaxa</i>. <b>3882</b> (2): 001–230. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.11646%2Fzootaxa.3882.1.1">10.11646/zootaxa.3882.1.1</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/25543675">25543675</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Zootaxa&rft.atitle=Family-group+names+of+Recent+fishes&rft.volume=3882&rft.issue=2&rft.pages=001-230&rft.date=2014&rft_id=info%3Adoi%2F10.11646%2Fzootaxa.3882.1.1&rft_id=info%3Apmid%2F25543675&rft.au=Richard+van+der+Laan&rft.au=William+N.+Eschmeyer&rft.au=Ronald+Fricke&rft_id=https%3A%2F%2Fdoi.org%2F10.11646%252Fzootaxa.3882.1.1&rfr_id=info%3Asid%2Fen.wikipedia.org%3AClimbing+gourami" class="Z3988"></span></span> </li> <li id="cite_note-FBF-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-FBF_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-FBF_2-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFroesePauly" class="citation web cs1"><a href="/wiki/Rainer_Froese" title="Rainer Froese">Froese, Rainer</a>; Pauly, Daniel (eds.). <a rel="nofollow" class="external text" href="http://www.fishbase.org/Summary/FamilySummary.php?family=Anabantidae">"Family Anabantidae"</a>. <i><a href="/wiki/FishBase" title="FishBase">FishBase</a></i>. August 2019 version.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=FishBase&rft.atitle=Family+Anabantidae&rft_id=http%3A%2F%2Fwww.fishbase.org%2FSummary%2FFamilySummary.php%3Ffamily%3DAnabantidae&rfr_id=info%3Asid%2Fen.wikipedia.org%3AClimbing+gourami" class="Z3988"></span></span> </li> <li id="cite_note-Eoanabas-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-Eoanabas_3-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFeixiang_WuDesui_MiaoMee-mann_ChangGongle_Shi2017" class="citation journal cs1">Feixiang Wu; Desui Miao; Mee-mann Chang; Gongle Shi & Ning Wang (2017). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5429824">"Fossil climbing perch and associated plant megafossils indicate a warm and wet central Tibet during the late Oligocene"</a>. <i>Scientific Reports</i>. <b>7</b> (878): 878. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fs41598-017-00928-9">10.1038/s41598-017-00928-9</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5429824">5429824</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/28408764">28408764</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Scientific+Reports&rft.atitle=Fossil+climbing+perch+and+associated+plant+megafossils+indicate+a+warm+and+wet+central+Tibet+during+the+late+Oligocene&rft.volume=7&rft.issue=878&rft.pages=878&rft.date=2017&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5429824%23id-name%3DPMC&rft_id=info%3Apmid%2F28408764&rft_id=info%3Adoi%2F10.1038%2Fs41598-017-00928-9&rft.au=Feixiang+Wu&rft.au=Desui+Miao&rft.au=Mee-mann+Chang&rft.au=Gongle+Shi&rft.au=Ning+Wang&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5429824&rfr_id=info%3Asid%2Fen.wikipedia.org%3AClimbing+gourami" class="Z3988"></span></span> </li> </ol></div></div> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=172583">"Anabantidae"</a>. <a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">Integrated Taxonomic Information System</a><span class="reference-accessdate">. Retrieved <span class="nowrap">April 28,</span> 2004</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Anabantidae&rft.pub=Integrated+Taxonomic+Information+System&rft_id=https%3A%2F%2Fwww.itis.gov%2Fservlet%2FSingleRpt%2FSingleRpt%3Fsearch_topic%3DTSN%26search_value%3D172583&rfr_id=info%3Asid%2Fen.wikipedia.org%3AClimbing+gourami" class="Z3988"></span></li></ul> </section><div class="mw-heading mw-heading2 section-heading" onclick="mfTempOpenSection(3)"><span class="indicator mf-icon mf-icon-expand mf-icon--small"></span><h2 id="External_links">External links</h2><span class="mw-editsection"> <a role="button" href="/w/index.php?title=Climbing_gourami&action=edit&section=3" title="Edit section: External links" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet "> <span class="minerva-icon minerva-icon--edit"></span> <span>edit</span> </a> </span> </div><section class="mf-section-3 collapsible-block" id="mf-section-3"> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation encyclopaedia cs1"><span class="cs1-ws-icon" title="s:The New International Encyclopædia/Climbing Fish"><a class="external text" href="https://en.wikisource.org/wiki/The_New_International_Encyclop%C3%A6dia/Climbing_Fish">"Climbing Fish" </a></span>. <i><a href="/wiki/New_International_Encyclopedia" class="mw-redirect" title="New International Encyclopedia">New International Encyclopedia</a></i>. 1905.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Climbing+Fish&rft.btitle=New+International+Encyclopedia&rft.date=1905&rfr_id=info%3Asid%2Fen.wikipedia.org%3AClimbing+gourami" class="Z3988"></span></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output 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href="https://en.wikipedia.org/w/index.php?title=Climbing_gourami&oldid=1254687721">https://en.wikipedia.org/w/index.php?title=Climbing_gourami&oldid=1254687721</a>"</div></div> </div> <div class="post-content" id="page-secondary-actions"> </div> </main> <footer class="mw-footer minerva-footer" role="contentinfo"> <a class="last-modified-bar" href="/w/index.php?title=Climbing_gourami&action=history"> <div class="post-content last-modified-bar__content"> <span class="minerva-icon minerva-icon-size-medium minerva-icon--modified-history"></span> <span class="last-modified-bar__text modified-enhancement" data-user-name="Naftali Honig" data-user-gender="unknown" data-timestamp="1730438139"> <span>Last edited on 1 November 2024, at 05:15</span> </span> <span class="minerva-icon minerva-icon-size-small minerva-icon--expand"></span> </div> </a> <div class="post-content footer-content"> <div id='mw-data-after-content'> <div class="read-more-container"></div> </div> <div id="p-lang"> <h4>Languages</h4> <section> <ul id="p-variants" class="minerva-languages"></ul> <ul class="minerva-languages"><li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Anabantidae" title="Anabantidae – Afrikaans" lang="af" hreflang="af" data-title="Anabantidae" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%A3%D8%B3%D9%85%D8%A7%D9%83_%D8%A7%D9%84%D8%AC%D9%88%D8%B1%D8%A7%D9%85%D9%8A_%D8%A7%D9%84%D9%85%D8%AA%D8%B3%D9%84%D9%82%D8%A9" title="أسماك الجورامي المتسلقة – Arabic" lang="ar" hreflang="ar" data-title="أسماك الجورامي المتسلقة" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%90%D0%BD%D0%B0%D0%B1%D0%B0%D1%81%D0%BE%D0%B2%D0%B8" title="Анабасови – Bulgarian" lang="bg" hreflang="bg" data-title="Анабасови" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Anab%C3%A0ntids" title="Anabàntids – Catalan" lang="ca" hreflang="ca" data-title="Anabàntids" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ceb mw-list-item"><a href="https://ceb.wikipedia.org/wiki/Anabantidae" title="Anabantidae – Cebuano" lang="ceb" hreflang="ceb" data-title="Anabantidae" data-language-autonym="Cebuano" data-language-local-name="Cebuano" class="interlanguage-link-target"><span>Cebuano</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Lezounovit%C3%AD" title="Lezounovití – Czech" lang="cs" hreflang="cs" data-title="Lezounovití" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Kletterfische_und_Buschfische" title="Kletterfische und Buschfische – German" lang="de" hreflang="de" data-title="Kletterfische und Buschfische" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Anabantidae" title="Anabantidae – Spanish" lang="es" hreflang="es" data-title="Anabantidae" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Anabantidae" title="Anabantidae – Basque" lang="eu" hreflang="eu" data-title="Anabantidae" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DA%AF%D9%88%D8%B1%D8%A7%D9%85%DB%8C_%D8%A8%D8%A7%D9%84%D8%A7%D8%B1%D9%88" title="گورامی بالارو – Persian" lang="fa" hreflang="fa" data-title="گورامی بالارو" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Anabantidae" title="Anabantidae – French" lang="fr" hreflang="fr" data-title="Anabantidae" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%93%B1%EB%AA%A9%EC%96%B4%EA%B3%BC" title="등목어과 – Korean" lang="ko" hreflang="ko" data-title="등목어과" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Anabantidae" title="Anabantidae – Indonesian" lang="id" hreflang="id" data-title="Anabantidae" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Anabantidae" title="Anabantidae – Italian" lang="it" hreflang="it" data-title="Anabantidae" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%90%D7%A0%D7%90%D7%91%D7%A1%D7%99%D7%99%D7%9D" title="אנאבסיים – Hebrew" lang="he" hreflang="he" data-title="אנאבסיים" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D2%9A%D1%83%D1%8B%D1%81%D1%82%D1%8B%D0%BB%D0%B0%D1%80" title="Қуыстылар – Kazakh" lang="kk" hreflang="kk" data-title="Қуыстылар" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/%C5%A0liau%C5%BEikin%C4%97s" title="Šliaužikinės – Lithuanian" lang="lt" hreflang="lt" data-title="Šliaužikinės" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Labirintkopolty%C3%BAs_halak" title="Labirintkopoltyús halak – Hungarian" lang="hu" hreflang="hu" data-title="Labirintkopoltyús halak" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%95%E0%B4%B0%E0%B4%BF%E0%B4%AA%E0%B5%8D%E0%B4%AA%E0%B4%BF%E0%B4%9F%E0%B4%BF" title="കരിപ്പിടി – Malayalam" lang="ml" hreflang="ml" data-title="കരിപ്പിടി" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%A7%D8%B3%D9%85%D8%A7%D9%83_%D8%A7%D9%84%D8%AC%D9%88%D8%B1%D8%A7%D9%85%D9%89_%D8%A7%D9%84%D9%85%D8%AA%D8%B3%D9%84%D9%82%D9%87" title="اسماك الجورامى المتسلقه – Egyptian Arabic" lang="arz" hreflang="arz" data-title="اسماك الجورامى المتسلقه" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Klimbaarzen" title="Klimbaarzen – Dutch" lang="nl" hreflang="nl" data-title="Klimbaarzen" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Klatreguramier" title="Klatreguramier – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Klatreguramier" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/B%C5%82%C4%99dnikowate" title="Błędnikowate – Polish" lang="pl" hreflang="pl" data-title="Błędnikowate" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Anabantidae" title="Anabantidae – Portuguese" lang="pt" hreflang="pt" data-title="Anabantidae" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9F%D0%BE%D0%BB%D0%B7%D1%83%D0%BD%D0%BE%D0%B2%D1%8B%D0%B5" title="Ползуновые – Russian" lang="ru" hreflang="ru" data-title="Ползуновые" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Kiipij%C3%A4kalat" title="Kiipijäkalat – Finnish" lang="fi" hreflang="fi" data-title="Kiipijäkalat" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Kl%C3%A4tterfiskar" title="Klätterfiskar – Swedish" lang="sv" hreflang="sv" data-title="Klätterfiskar" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B8%A7%E0%B8%87%E0%B8%A8%E0%B9%8C%E0%B8%9B%E0%B8%A5%E0%B8%B2%E0%B8%AB%E0%B8%A1%E0%B8%AD" title="วงศ์ปลาหมอ – Thai" lang="th" hreflang="th" data-title="วงศ์ปลาหมอ" data-language-autonym="ไทย" data-language-local-name="Thai" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Anabantidae" title="Anabantidae – Turkish" lang="tr" hreflang="tr" data-title="Anabantidae" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%90%D0%BD%D0%B0%D0%B1%D0%B0%D1%81%D0%BE%D0%B2%D1%96" title="Анабасові – Ukrainian" lang="uk" hreflang="uk" data-title="Анабасові" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/H%E1%BB%8D_C%C3%A1_r%C3%B4_%C4%91%E1%BB%93ng" title="Họ Cá rô đồng – Vietnamese" lang="vi" hreflang="vi" data-title="Họ Cá rô đồng" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Anabantidae" title="Anabantidae – Waray" lang="war" hreflang="war" data-title="Anabantidae" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E6%94%80%E9%B2%88%E7%A7%91" title="攀鲈科 – Chinese" lang="zh" hreflang="zh" data-title="攀鲈科" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li></ul> </section> </div> <div class="minerva-footer-logo"><img 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